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A crinoid fauna and a new species of Pycnocrinus from the Martinsburg Formation (Upper Ordovician), lower Hudson Valley, New York

Published online by Cambridge University Press:  13 May 2024

James C. Brower
Affiliation:
Earth Science Department, Heroy Geology Laboratory, 141 Crouse Drive, Syracuse University, Syracuse, New York 13244, USA.
Carlton E. Brett
Affiliation:
Department of Geosciences, 500 Geology-Physics Building, Clifton Court, University of Cincinnati, Cincinnati, Ohio 45221-0013, USA.
Howard R. Feldman*
Affiliation:
Biology Department, 227 West 60th Street, Lander College for Women, a division of Touro University, New York, NY 10023, USA.
*
*Corresponding author.

Abstract

A new crinoid fauna has been discovered in the Upper Ordovician (Katian) Martinsburg Formation at a small shale quarry, locally known as the ‘Shale Bank,’ on the Shawangunk Ridge in Ulster County, NY. The assemblage, which is from a relatively low energy, offshore mud-bottom environment, includes four identified species, including a new species of glyptocrinid camerate, Pycnocrinus mohonkensis n. sp., described herein. Crinoid taxa in order of increasing branch density in the assemblage include (1) the dicyclic inadunate Merocrinus curtus with irregularly isotomous and heterotomous, non-pinnulate arms and a stout cylindrical column exceeding 700 mm; (2) the disparids Cincinnaticrinus varibrachialus, with heterotomous non pinnulate arms, and Ectenocrinus simplex, with extensively branched ramulate arms and meric columns of 460–500 mm; and (3) the camerate Pycnocrinus mohonkensis n. sp., with uniserial pinnulate arms and a somewhat shorter column. Some cylindrical stems with nodose and holomeric columnals are thought to belong to unknown camerate crinoids with pinnulate arms. Filtration theory is used to model food capture in the Martinsburg crinoids. Surprisingly, even densely pinnulate camerates were able to survive in this setting, suggesting that ambient currents attained velocities exceeding 25 cm/sec even in this offshore setting. Similar assemblages were widespread in eastern Laurentia during the Late Ordovician.

UUID: http://zoobank.org/23ca31e8-f572-4520-ba1d-891e3abb950d

Type
Articles
Copyright
Copyright © Touro University, University of Cincinnati, and The Estate of Jim Brower, 2024. Published by Cambridge University Press on behalf of Paleontological Society

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Footnotes

Deceased.

References

Ausich, W.I., 1980, A model for differentiation in lower Mississippian crinoid communities: Journal of Paleontology, v. 54, p. 273288.Google Scholar
Ausich, W.I., 1998, Phylogeny of Arenig to Caradoc crinoids (Phylum Echinodermata) and suprageneric classification of the Crinoidea: The University of Kansas Paleontological Contributions Papers, New Series, no. 9, 36 p.Google Scholar
Ausich, W.I., 1999, Upper Ordovician of the Cincinnati, Ohio, area, USA, in Hess, H., Ausich, W.I., Brett, C.E., and Simms, M.J., eds., Fossil Crinoids: Cambridge University Press, Cambridge, United Kingdom, p. 7580.CrossRefGoogle Scholar
Ausich, W.I., Kammer, T.W., and Baumiller, T.K., 1994, Demise of the middle Paleozoic crinoid fauna: a single extinction event or rapid faunal turnover?: Paleobiology v. 20, p. 345361.CrossRefGoogle Scholar
Ausich, W.I., Brett, C.E., and Hess, H., 1999, Taphonomy, in Hess, H., Ausich, W.I., Brett, C.E., and Simms, M.J., eds., Fossil Crinoids: Cambridge University Press, Cambridge, United Kingdom, p. 5059.CrossRefGoogle Scholar
Ausich, W.I., Wright, D.F., Cole, S.R., and Koniecki, J.M., 2018, Disparid and hybocrinid crinoids (Echinodermata) from the Upper Ordovician (lower Katian) Brechin Lagerstätte of Ontario: Journal of Paleontology, v. 92, p. 850871.CrossRefGoogle Scholar
Baird, G.C., and Brett, C.E., 2002, Indian Castle Shale: late synorogenic siliciclastic succession in an evolving Middle to Late Ordovician foreland basin, eastern New York State: Physics and Chemistry of the Earth, v. 27, p. 203230.CrossRefGoogle Scholar
Baumiller, T.K., 1993, Survivorship analysis of Paleozoic Crinoidea: effect of filter morphology on evolutionary rates: Paleobiology, v. 19, p. 304321.CrossRefGoogle Scholar
Billings, E., 1857, Report for the Year 1856: Canada Geological Survey Report of Progress for the Years 1853–54–55–56, p. 247345.Google Scholar
Billings, E., 1859, On the Crinoideae of the Lower Silurian rocks of Canada, figures and descriptions of Canadian organic remains, decade IV: Canadian Geological Survey, 72 p.CrossRefGoogle Scholar
Bretsky, P.W. Jr., 1970, Upper Ordovician ecology of the central Appalachians: Peabody Museum of Natural History Bulletin, v. 34, p. 1150.Google Scholar
Brett, C.E., 1984, Autecology of Silurian pelmatozoan echinoderms, in Bassett, M.G., and Lawson, J.D., eds., Autecology of Silurian Organisms: Special Papers in Palaeontology, v. 32, p. 87120.Google Scholar
Brett, C.E., 1999, Chapter 6, Middle Ordovician Trenton Group of New York, in Hess, H., Ausich, W.I., Brett, C.E., and Simms, M.J., eds., Fossil Crinoids: Cambridge University Press, Cambridge, United Kingdom, p. 6367.CrossRefGoogle Scholar
Brett, C.E., and Eckert, J.D., 1982, Paleoecology of a well-preserved crinoid colony from the Silurian Rochester Shale in Ontario: Royal Ontario Museum Contributions to Life Sciences, v. 131, p. 120.Google Scholar
Brett, C.E., Whiteley, T.E., Allison, P.A., and Yochelson, E.L., 1999, The Walcott–Rust Quarry: Middle Ordovician trilobite Konservat-Lagerstätten: Journal of Paleontology, v. 73, p. 288305.Google Scholar
Brett, C.E., Deline, B.L., and McLaughlin, P.I., 2008, Attachment, facies distribution, and life history strategies in crinoids from the Upper Ordovician of Kentucky, in Ausich, W.I. and Webster, G.D., eds., Echinoderm Paleobiology: Indiana University Press, Bloomington and Indianapolis, Indiana, p. 2252.Google Scholar
Brett, C.E., Aucoin, C.D., Dattilo, B.F, Freeman, R.L, Hartshorn, K.R., McLaughlin, P.I., and Schwalbach, C.E., 2020. Revised sequence stratigraphy of the upper Katian Stage (Cincinnatian) strata in the Cincinnati Arch reference area: geological and paleontological implications: Palaeogeography, Palaeoclimatology, Palaeoecology, v. 540, 109483, https://doi.org/10.1016/j.palaeo.2019.109483.Google Scholar
Brower, J.C., 1974, Upper Ordovician xenocrinids (Crinoidea, Camerata) from Scotland: University of Kansas Paleontological Contributions, Paper 67, p. 125.Google Scholar
Brower, J.C., 1992, Hybocrinid and disparid crinoids from the Middle Ordovician (Galena Group, Dunleith Formation) of northern Iowa and southern Minnesota: Journal of Paleontology, v. 66, p. 973993.CrossRefGoogle Scholar
Brower, J.C., 1994, Camerate crinoids from the Middle Ordovician (Galena Group, Dunleith Formation) of northern Iowa and southern Minnesota: Journal of Paleontology, v. 68, p. 570599.CrossRefGoogle Scholar
Brower, J.C., 1997, Homocrinid crinoids from the Upper Ordovician of northern Iowa and southern Minnesota: Journal of Paleontology, v. 71, p. 442458.CrossRefGoogle Scholar
Brower, J.C., 2005, The paleobiology and ontogeny of Cincinnaticrinus varibrachialus Warn and Strimple, 1977 from the Middle Ordovician (Shermanian) Walcott–Rust Quarry of New York: Journal of Paleontology, v. 79, p. 152174.2.0.CO;2>CrossRefGoogle Scholar
Brower, J.C., 2006, Ontogeny of the food-gathering system in Ordovician crinoids: Journal of Paleontology, v. 80, p. 430446.CrossRefGoogle Scholar
Brower, J.C., 2007, The application of filtration theory to food gathering in Ordovician crinoids: Journal of Paleontology, v. 81, p. 12841300.CrossRefGoogle Scholar
Brower, J.C., 2008, Some disparid crinoids from the Upper Ordovician (Shermanian) Walcott–Rust Quarry of New York: Journal of Paleontology, v. 82, p. 5777.CrossRefGoogle Scholar
Brower, J.C., 2010, Camerate and cladid crinoids from the Upper Ordovician (Katian, Shermanian) Walcott–Rust Quarry of New York: Journal of Paleontology, v. 84, p. 626645.CrossRefGoogle Scholar
Brower, J.C., 2011, Paleoecology of suspension-feeding echinoderm assemblages from the Upper Ordovician (Katian, Shermanian) Walcott–Rust Quarry of New York: Journal of Paleontology, v. 85, p. 369391.CrossRefGoogle Scholar
Brower, J.C., 2013, Paleoecology of echinoderm assemblages from the Upper Ordovician (Katian) Dunleith Formation of northern Iowa and southern Minnesota: Journal of Paleontology v. 87, p. 1643.CrossRefGoogle Scholar
Brower, J.C., and Kile, K.M., 1994, Paleoautecology and ontogeny of Cupulocrinus levorsoni Kolata, a Middle Ordovician crinoid from the Guttenberg Formation of Wisconsin, in Landing, E., ed., Studies in Stratigraphy and Paleontology in Honor of Donald W. Fisher: New York State Museum Bulletin, v. 481, p. 2544.Google Scholar
Brower, J.C. and Veinus, J, 1978, Middle Ordovician crinoids from the Twin Cities area of Minnesota: Bulletins of American Paleontology, v. 74, p. 373506.Google Scholar
Cole, S.R., 2017, Phylogeny and morphologic evolution of the Ordovician Camerata (Class Crinoidea, Phylum Echinodermata): Journal of Paleontology, v. 91, p. 815828.CrossRefGoogle Scholar
Cole, S.R., Ausich, W.I., Wright, D.F., and Koniecki, J.M., 2018, An echinoderm Lagerstätte from the Upper Ordovician (Katian), Ontario: taxonomic re-evaluation and description of new dicyclic camerate crinoids: Journal of Paleontology, v. 92, p. 488505.CrossRefGoogle Scholar
Cole, S.R., Wright, D.F., and Ausich, W.I., 2019, Phylogenetic community paleoecology of one of the earliest complex crinoid faunas (Brechin Lagerstätte, Ordovician): Palaeogeography, Palaeoclimatology, Palaeoecology, v. 521, p. 8298.CrossRefGoogle Scholar
Cole, S.R., Wright, D.F., Ausich, W.I., and Koniecki, J.M., 2020, Paleocommunity composition, relative abundance, and new camerate crinoids from the Brechin Lagerstätte (Upper Ordovician): Palaeogeography, Palaeoclimatology, Palaeoecology, v. 94, p. 11031123.Google Scholar
Conrad, T.A., 1838, Report of the Palaeontological Department of the Survey: New York State Geological Survey, Annual Report, v. 2, p. 107119.Google Scholar
DeKay, J.E., 1824, Observations on the structure of trilobites, and descriptions of an apparently new genus, with notes on the geology of Trenton Falls by J. Renwick: Annals of the Lyceum of Natural History, New York, v. 1, p. 174189.Google Scholar
Eckert, J. D., 1987, Pycnocrinus altilis, a new Late Ordovician channel-dwelling crinoid from southern Ontario: Canadian Journal of Earth Sciences, v. 24, p. 851859.CrossRefGoogle Scholar
Emmons, E., 1842, Natural History of New York, Geology 2: Albany, NY, Carroll and Cooke Printers to the Assembly.Google Scholar
English, A.E., Landing, E., and Baird, G.C., 2006, Snake Hill—reconstructing eastern Taconic foreland basin litho- and biofacies from a giant mélange block in eastern New York, USA: Palaeogeography, Palaeoclimatology, Palaeoecology, v. 242, p. 201213.CrossRefGoogle Scholar
Epstein, J.B., and Lyttle, P.T., 1987, Structure and stratigraphy above, below and within the Taconic unconformity, southeastern New York, in Waines, R.H., ed., Fieldtrip Guidebook, New York State Geological Association, 59th Annual Meeting, Kingston, New York. November 6–8, 1987. New Paltz, New York, State University of New York, College at New Paltz, p. C1C8.Google Scholar
Epstein, J.B., and Lyttle, P.T., 2012, A journey along the Taconic unconformity: interpretations, perplexities, and wonderments, northeastern Pennsylvania, northern New Jersey, and southeasternmost New York: 77th Annual Field Conference of Pennsylvania Geologists. Shawnee-on-Delaware, Pennsylvania, v. 77, p. 136.Google Scholar
Ettensohn, F.R., 2008, Chapter 4: The Appalachian foreland basin in the eastern United States, in Miall, A., ed., Sedimentary Basins of the World: The Sedimentary Basins of the United States and Canada: Amsterdam, Elsevier, v. 5, p. 105179.CrossRefGoogle Scholar
Feldman, H.R., 1980, Level-bottom brachiopod communities in the Middle Devonian of New York: Lethaia, v. 13, p. 2746.CrossRefGoogle Scholar
Feldman, H.R., Smoliga, J., and Feldman, B.A., 2012, Notes on the Geology of the Shawangunk Ridge on the Mohonk Preserve and environs: Northeast Natural History Conference 2011: Selected Papers 2012: Northeastern Naturalist, v. 19, p. 312.CrossRefGoogle Scholar
Frest, T.J., Brett, C.E., and Witzke, B.J., 1999, Caradocian to Gedinnian echinoderm associations of central and eastern North America, in Boucot, A.J., and Lawson, J.D., eds., Paleocommunities: A Case Study from the Silurian and Lower Devonian: Cambridge University Press, Cambridge, UK, p. 638783.Google Scholar
Gorzelak, P., Dorota, K., Salamon, M.A., Magdalena, Ł., Ausich, W.I., and Baumiller, T.K., 2020, Bringing planktonic crinoids back to the bottom: reassessment of the functional role of scyphocrinoid loboliths: Paleobiology, v. 46, p. 104122.CrossRefGoogle Scholar
Hall, J., 1847, Palaeontology of New York, v. 1, containing descriptions of the organic remains of the lower division of the New-York system (equivalent of the Lower Silurian rocks of Europe). Natural History of New York: Albany, State of New York, v. 6, 338 p.Google Scholar
Hall, J., 1860, New species of fossils from the Hudson River Group of Ohio, and other western states: Annual Report of the Regents of the University of the State of New York, on the Condition of the State Cabinet of Natural History, v. 13, p. 119121.Google Scholar
Holland, S.M., Miller, A.I., Meyer, D.L., and Datillo, B.F., 2001, The detection and importance of subtle biofacies within a single lithofacies: the Upper Ordovician Kope Formation of the Cincinnati, Ohio region: Palaios, v. 16, p. 205217.2.0.CO;2>CrossRefGoogle Scholar
Holterhoff, P.F., 1997, Paleocommunity and evolutionary ecology of Paleozoic crinoids, in Waters J.A. and Maples, C.G., eds, Geobiology of Echinoderms: Paleontological Society Papers, v. 3, p. 69106.CrossRefGoogle Scholar
Jaekel, O., 1918, Phylogenie und System der Pelmatozoen: Paläontologische Zeitschrift, v. 3, p. 1128.CrossRefGoogle Scholar
Kallmeyer, J.W., and Ausich, W.L., 2016. Deepwater occurrence of a new Glyptocrinus (Crinoidea, Camerata) from the Late Ordovician of southwestern Ohio and northern Kentucky: revision of crinoid community composition: Journal of Paleontology, v. 89, p. 10681075.CrossRefGoogle Scholar
Kidd, W.S.F., Plesch, A., and Vollmer, F.W., 1995, Lithofacies and structure of the Taconic Flysch, mélange, and allochthon, in the New York Capital District, in Garver, J.I., Smith, J.A., eds., Field Trips for the 67th Annual Meeting of the New York State Geological Association: Schenectady, NY, Union College, v. 67, p. 5780.Google Scholar
Kirk, E., 1914, Notes on the fossil crinoid genus Homocrinus Hall: United States National Museum Proceedings, v. 46, p. 473483.CrossRefGoogle Scholar
Kolata, D.R., 1976, Crinoids from the Upper Ordovician Bighorn Formation of Wyoming: Journal of Paleontology, v. 50, p. 445453.Google Scholar
Kolata, D.R., 1982, Camerates, in Sprinkle, J., ed., Echinoderm Faunas from the Bromide Formation (Middle Ordovician) of Oklahoma: University of Kansas Paleontological Contributions, Monograph 1, p. 170205.Google Scholar
Kolata, D.R., 1986, Crinoids of the Champlainian (Middle Ordovician) Guttenberg Formation—upper Mississippi Valley region: Journal of Paleontology, v. 60, p. 711718.CrossRefGoogle Scholar
Landing, E., 1988, Depositional tectonics and biostratigraphy of the western portion of the Taconic allochthon, eastern New York State, in Landing, E., ed., The Canadian Paleontology and Biostratigraphy Seminar: New York State Museum Bulletin 462, p. 96110.Google Scholar
Landing, E., Pe-Piper, G., Kidd, W.S.F., and Azmy, K., 2003, Tectonic setting of outer trench slope volcanism: pillow basalt and limestone in the Ordovician Taconian orogen of eastern New York: Canadian Journal of Earth Sciences, v. 40, p. 11731187.CrossRefGoogle Scholar
Lehman, D., and Pope, J.K., 1989, Upper Ordovician tempestites from Swatara Gap, Pennsylvania: depositional processes affecting the sediments and paleoecology of the fossil faunas: Palaios, v. 89, p. 553564.CrossRefGoogle Scholar
Liberty, B.A., 1969, Palaeozoic geology of the Lake Simcoe area, Ontario: Geological Survey of Canada, Memoir 355, 201 p.Google Scholar
Meek, F.B., 1871, On some new Silurian (Ordovician) crinoids and shells: American Journal of Science, ser. 3, v. 1, p. 295299.CrossRefGoogle Scholar
Meek, F.B., 1872, Descriptions of new western Palaeozoic fossils mainly from the Cincinnati Group of the Lower Silurian series of Ohio: Proceedings of the Academy of Natural Sciences of Philadelphia, v. 23, p. 308337.Google Scholar
Meek, F.B., 1873a, Fossils of the Cincinnati Group: Geological Survey of Ohio, v. 1, pt. 2 (palaeontology), 175 p.Google Scholar
Meek, F.B., 1873b, Descriptions of invertebrate fossils of the Silurian and Devonian Systems: Report of the Geological Survey of Ohio, v. 1, p. 1243.Google Scholar
Messing, C.G., 1997, Living comatulids, in Waters, J.A., and Maples, C.G., eds., Geobiology of Echinoderms: Paleontological Society Papers, v. 3, p. 330.Google Scholar
Meyer, D.L., 1982, Food and feeding mechanisms: Crinozoa, in Jangoux, M., and Lawrence, J.M., eds., Echinoderm Nutrition: Rotterdam, The Netherlands, A.A. Balkema, p. 2542.Google Scholar
Meyer, D.L., Miller, A.I., Holland, S.I., and Datillo, B.F., 2002, Crinoid distribution and feeding morphology through a depositional sequence: Kope and Fairview formations, Upper Ordovician, Cincinnati Arch region: Journal of Paleontology, v. 76, p. 725732.Google Scholar
Miller, J.S., 1821, A Natural History of the Crinoidea, or lily-shaped animals with observations on the genera, Asteria, Euryale, Comatula and Marsupites: Bristol, England, Bryan & Co., 150 p.Google Scholar
Miller, S.A., 1875, Glyptocrinus shafferi: Cincinnati Quarterly Journal of Science, v. 2, p. 277279.Google Scholar
Miller, S.A., 1880, Description of four new species and a new variety of Silurian fossils and remarks upon others: Journal of the Cincinnati Society of Natural History, v. 3, p. 232236.Google Scholar
Miller, S.A., 1883, Glyptocrinus redefined and restricted, Gaurocrinus, Pycnocrinus and Compsocrinus established, and two new species described: Journal of the Cincinnati Society of Natural History, v. 6, p. 217234.Google Scholar
Miller, S.A., 1889, North American Geology and Palaeontology: Cincinnati, Ohio, Western Methodist Book Concern, 664 p.Google Scholar
Miller, S.A., 1890, The structure, classification, and arrangement of American Palaeozoic crinoids into families: American Geologist, v. 6, p. 275286, 340–357.Google Scholar
Moore, R.C., 1952, Evolution rates among crinoids: Journal of Paleontology, v. 26, p. 338352.Google Scholar
Moore, R.C., and Laudon, L.R., 1943, Evolution and classification of Paleozoic crinoids: Geological Society of America Special Paper, v. 46, p. 1167.CrossRefGoogle Scholar
Paton, T.R., Brett, C.E., and Kampouris, G.E., 2019, Genesis, modification, and preservation of complex Upper Ordovician hardgrounds: implications for sequence stratigraphy and the Great Ordovician Biodiversification Event: Palaeogeography, Palaeoclimatology, Palaeoecology, v. 526, p. 5371.CrossRefGoogle Scholar
Rowley, D.B., and Kidd, W.S.F., 1981, Stratigraphic relationships and detrital composition of the medial Ordovician flysch of western New England: implications for the tectonic evolution of the Taconic orogeny: Journal of Geology v. 89, p. 199218.CrossRefGoogle Scholar
Ruedemann, R., 1926, Utica and Lorraine formations of New York. Part 2, systematic paleontology: New York State Museum Bulletin, v. 272, 168 p.Google Scholar
Sowerby, G.B., 1839, A Conchological Manual: London, George Odell, Printer, 130 p.CrossRefGoogle Scholar
Titus, R., 1989, Clinal variation in the evolution of Ectenocrinus simplex: Journal of Paleontology, v. 63, p. 8191.CrossRefGoogle Scholar
Turpaeva, E.P., 1957, Food interrelationships of dominant species in marine benthic Biocoenoses, in Nikitkin, B.N., ed., Transactions, Institute Oceanography, Marine Biology USSR Academy of Science Press v. 20, p.137148. [published in the U.S. by the American Institute of Biological Sciences, Washington, D.C.]Google Scholar
Ubaghs, G., 1978, Skeletal morphology of fossil crinoids, in Moore, R.C., and Teichert, C., eds., Treatise on Invertebrate Paleontology, Part T, Echinodermata 2: Lawrence, Kansas, The Geological Society of America and University of Kansas Press, p. T58T216.Google Scholar
Ulrich, E.O., 1878, Descriptions of new species of fossils from the Cincinnati Group: Journal of the Cincinnati Natural History Society, v. 1(3), p. 92100.Google Scholar
Ulrich, E.O., 1879, Descriptions of new genera and species of fossils from the Lower Silurian about Cincinnati: Cincinnati Society of Natural History Journal, v. 2, p. 830.Google Scholar
Ulrich, E.O., 1882, Descriptions of two new species of crinoids: Cincinnati Society of Natural History Journal, v. 5, p. 175177.Google Scholar
Vollmer, F.W., and Bosworth, W., 1984, Formation of mélange in a foreland basin overthrust setting: example from the Taconic Orogen, in Raymond, L.A., ed., Mélanges: Their Nature, Origin, and Significance: Geological Society of America Special Paper, v. 198, p. 5370.Google Scholar
Wachsmuth, C., and Springer, F., 1885, Revision of the Paleocrinoidea, Part 3, Section 1. Discussion of the classification and relations of the brachiate crinoids, and conclusion of the generic descriptions: Academy of Natural Sciences, Philadelphia, Proceedings for 1885, p. 223364 (1–138).Google Scholar
Wachsmuth, C., and Springer, F., 1897, The North American Crinoidea Camerata: Harvard University Museum of Comparative Zoology, Memoir 20, v. 21, p. 1897.Google Scholar
Walcott, C.D., 1883, Descriptions of new species of fossils from the Trenton Group of New York: Thirty-fifth Annual Report of the New York State Museum of Natural History, p. 207214. [advanced print, 15 October 1883, p. 1–8.]Google Scholar
Walcott, C.D., 1884, Descriptions of new species of fossils from the Trenton Group of New York: Thirty-fifth Annual Report of the New York State Museum of Natural History, p. 207214.Google Scholar
Walker, K.R., 1972, Trophic analysis: a method for studying the function of ancient communities: Journal of Paleontology, v. 46, p. 8293.Google Scholar
Warn, J.M., and Strimple, H.L., 1977, The disparid inadunate superfamilies Homocrinacea and Cincinnaticrinacea (Echinodermata: Crinoidea), Ordovician–Silurian, North America: Bulletins of American Paleontology, v. 72, p. 1138.Google Scholar
Wilson, A.E., 1946, Echinodermata of the Ottawa Formation of the Ottawa–St. Lawrence Lowland: Canada Geological Survey Bulletin, v. 4, p. 161.Google Scholar
Woodley, J.D., 1980, The biomechanics ophiuroid tube-feet, in Jangoux, M., ed., Echinoderms: Present and Past. Proceedings of the European Colloquium on Echinoderms, 3–8 September, 1979: Rotterdam, The Netherlands, A. A. Balkema, p. 293299.Google Scholar
Wright, D.F., Ausich, W.I., Cole, S.R., Rhenberg, E.C., and Peter, M.E., 2017, Phylogenetic taxonomy and classification of the Crinoidea (Echinodermata): Journal of Paleontology, v. 91, p. 829846.CrossRefGoogle Scholar
Wright, D.F., Cole, S.R., and Ausich, W.I., 2019, Biodiversity, systematics, and new taxa of cladid crinoids from the Ordovician Brechin Lagerstätte: Journal of Paleontology, v. 94, p. 334357.CrossRefGoogle Scholar
Zittel, K.A. von., 1876–1880, Handbuch der Palaeontologie, Band 1, Palaeozoologie, Abt. 1: München und Leipzig, Germany, R. Oldenbourg, p. 1765.Google Scholar