Hostname: page-component-848d4c4894-tn8tq Total loading time: 0 Render date: 2024-06-14T01:10:48.209Z Has data issue: false hasContentIssue false
  • English
  • Français

Ontogeny and systematics of Toernquistiidae (Trilobita, Proetida) from the Ordovician of the Argentine Precordillera

Published online by Cambridge University Press:  20 May 2016

Brian D. E. Chatterton ,
Gregory D. Edgecombe ,
Beatriz G. Waisfeld  and
Norberto E. Vaccari
Brian D. E. Chatterton
Affiliation:
Department of Earth and Atmospheric Sciences, University of Alberta, Edmonton, T6G 2E3, Canada
Gregory D. Edgecombe
Affiliation:
Division of Earth and Environmental Sciences, Australian Museum, 6 College Street, Sydney South, NSW 2000, Australia
Beatriz G. Waisfeld
Affiliation:
División Paleozoología de Invertebrados, Museo Argentino de Ciencias Naturales “Bernardino Rivadavia”, Avenida ángel Gallardo 470, 1405 Buenos Aires, Argentina
Norberto E. Vaccari
Affiliation:
CONICET, Cátedra de Estratigrafía y Geología Histórica, Universidad Nacional de Córdoba, Avenida Vélez Sársfield 299, 5000 Córdoba, Argentina
Get access Rights & Permissions [Opens in a new window]

Abstract

The following new Toernquistiidae from Whiterockian (Llanvirn to lower Caradoc) strata of the Precordillera, western Argentina, are described: Chomatopyge canasi new species, Lasarchopyge benedettoi new genus and species, Lasarchopyge correae new genus and species, Lasarchopyge new species A, and Paratoernquistia sanchezae new genus and species. Phylogenetic analysis suggests that the Toernquistiidae Hupé, 1953 (including Chomatopyge, Lasarchopyge, Mesotaphraspis, Paratoernquistia, and Toernquistia) is closely related to but distinct from the Dimeropygidae Hupé, 1953 (including Ischyrotoma, Ischyrophyma and Dimeropyge). It is difficult to find unique synapomorphies that unite toernquistiids and dimeropygids into a monophyletic group that excludes other proetides (e.g., hystricurids), although details of their ontogenies are similar. Several species of Toernquistiidae are known from the Argentine Precordillera (one species previously described), and other species of this family have been described from Australia, Baltica, China, Kazakhstan and Laurentia. Members of the Dimeropygidae are known only from Baltica and Laurentia.

Type
Research Article
Information
Journal of Paleontology , Volume 72 , Issue 2 , March 1998 , pp. 273 - 303
Copyright
Copyright © The Paleontological Society 

Access options

Get access to the full version of this content by using one of the access options below. (Log in options will check for institutional or personal access. Content may require purchase if you do not have access.)

References

Adrain, J. M., and Chatterton, B. D. E. 1994. The aulacopleurid trilobite Otarion, with new species from the Silurian of northwestern Canada. Journal of Paleontology, 68:305–323.Google Scholar
Adrain, J. M., and Chatterton, B. D. E. 1995a. The otarionine trilobites Harpidella and Maurotarion, with species from northwestern Canada, the United States, and Australia. Journal of Paleontology, 69:307–326.Google Scholar
Adrain, J. M., and Chatterton, B. D. E. 1995b. Aulacopleurine trilobites from the Llandovery of northwestern Canada. Journal of Paleontology, 69:326–340.Google Scholar
Albanesi, G. L., and Astini, R. A. 1994. Conodontofauna de los niveles cuspidales de la Formación San Juan (Llanvirniano) en el perfil de Las Chacritas, Provincia de San Juan. VI Congreso Argentino de Paleontología y Bioestratigrafía. Resumenes, p. 48–49. Trelew.Google Scholar
Astini, R. A. 1994. Significado estratigráfico del Miembro Superior de la Formacion San Juan, cordón de Las Chacritas, Ordovícico medio de la Precordillera de San Juan. Revista de la Asociación Geológica Argentina, 49:365–367.Google Scholar
Astini, R. A. 1995. Sedimentología de la Formación Las Aguaditas (talud carbonático) y evolución de la cuenca precordillera durante el Ordovícico medio. Revista de la Asociación Geológica Argentina, 50:143–164.Google Scholar
Baldis, B. A. J., and Blasco, G. 1973. Trilobites ordovícicos de Ponon Trehue, Sierra Pintada de San Rafael, Provincia de Mendoza. Ameghiniana, 10:72–88.Google Scholar
Brussa, E. D. 1994. Las graptofaunas ordovícicas del sector central de la Precordillera Occidental sanjuanina. Unpubl. , Universidad Nacional de Córdoba, 323 p.Google Scholar
Brussa, E. D. 1996. Las graptofaunas ordovícicas de la Formación Las Aguaditas, Precordillera de San Juan, Argentina. Parte I: Familias Thamnograptidae, Dichograptidae, Abrograptidae y Glossograptidae. Ameghiniana, 33:421–434.Google Scholar
Bremer, K. 1994. Branch support and tree stability. Cladistics, 10:295–304.Google Scholar
Bruton, D. L. 1983. The morphology of Celmus (Trilobita) and its classification. Special Papers in Palaeontology, 30:213–219.Google Scholar
Chang, W.-T., and Fan, C.-S. 1960. Class Trilobita of the Ordovician and Silurian periods of the Chilian Mountains. Geological Gazetteer of the Chilian Mountains, 4:83–148. (In Chinese.).Google Scholar
Chatterton, B. D. E. 1971. Taxonomy and ontogeny of Siluro-Devonian trilobites from near Yass, New South Wales. Palaeontographica A, 137, 108 p.Google Scholar
Chatterton, B. D. E. 1980. Ontogenetic studies of Middle Ordovician trilobites from the Esbataottine Formation, Mackenzie Mountains, Canada. Palaeontographica A, 171, 74 p.Google Scholar
Chatterton, B. D. E. 1994. Ordovician proetide trilobite Dimeropyge, with a new species from northwestern Canada. Journal of Paleontology, 68:541–556.CrossRefGoogle Scholar
Chatterton, B. D. E., and Campbell, M. 1993. Enrolling in trilobites: a review and some new characters. Memoirs of the Association of Australasian Palaeontologists 15:103–123.Google Scholar
Chatterton, B. D. E., Edgecombe, G. D., Vaccari, N. E., and Waisfeld, B. G. 1997. Ontogeny and relationships of the Ordovician odontopleurid trilobite Ceratocara, with new species from Argentina and New York. Journal of Paleontology, 71:108–125.Google Scholar
Chatterton, B. D. E., and Ludvigsen, R. 1976. Silicified Middle Ordovician trilobites from the South Nahanni River area, District of Mackenzie, Canada. Palaeontographica A, 154, 106 p.Google Scholar
Chatterton, B. D. E., Siveter, D. J., Edgecombe, G. D., and Hunt, A. S. 1990. Larvae and relationships of the Calymenina (Trilobita). Journal of Paleontology, 64:255–277.Google Scholar
Chatterton, B. D. E., and Speyer, S. E. 1997. Trilobite ontogeny, p. 171–247. In Whittington, H. B. (ed.), Treatise on Invertebrate Paleontology, Part O, Trilobita, Revised Edition. Geological Society of America and University of Kansas Press, Lawrence, Kansas.Google Scholar
Churkin, M. Jr. 1963. Ordovician trilobites from graptolitic shale in central Idaho. Journal of Paleontology, 37:421–428.Google Scholar
Dean, W. T. 1974. The trilobites of the Chair of Kildare Limestone (Upper Ordovician) of eastern Ireland. Part 2. Palaeontographical Society Monographs, 128:61–98.Google Scholar
Edgecombe, G. D., Chatterton, B. D. E., Vaccari, N. E., and Waisfeld, B. G. 1997. Ontogeny of the proetoid trilobite Stenoblepharum and relationships of a new species from the Upper Ordovician of Argentina. Journal of Paleontology, 71:419–433.Google Scholar
Eriksson, T., and Wikström, N. 1995. AutoDecay version 3.0. Program distributed by authors, University of Stockholm.Google Scholar
Espisua, E. 1968. El Paleozóico inferior del Rio de Las Chacritas, Departamento Jáchal, Provincia de San Juan. Con especial referencia al Silúrico. Revista de la Asociación Geológica Argentina, 23:297–311.Google Scholar
Fortey, R. A. 1979. Early Ordovician trilobites from the Catoche Formation (St. George Group), western Newfoundland. Geological Survey of Canada, Bulletin, 321:61–114.Google Scholar
Fortey, R. A. 1980. The Ordovician trilobites of Spitsbergen III. Remaining trilobites of the Valhallfonna Formation. Norsk Polarinstitut Skrifter, 171, 163 p.Google Scholar
Fortey, R. A. 1983. Cambrian-Ordovician trilobites from the boundary beds in western Newfoundland and their phylogenetic significance. Special Papers in Palaeontology, 30:179–211.Google Scholar
Fortey, R. A., and Peel, J. S. 1989. Stratigraphy and hystricurid trilobites of the Christian Elv Formation (Lower Ordovician) of western northern Greenland. Rapport Grønlands geologiske Undersokning, 144:5–15.CrossRefGoogle Scholar
Fortey, R. A., and Owens, R. M. 1975. Proetida—a new order of trilobites. Fossils and Strata, 4:227–239.Google Scholar
Furque, G. 1975. Estratigrafía de la Serrania Mogotes Azules, SO de Jáchal, San Juan. I Congreso Argentino de Paleontología y Bioestratigrafía, Tucumán, 1:275–282.Google Scholar
Hintze, L. F. 1952. Lower Ordovician trilobites from western Utah and eastern Nevada. Utah Geological and Mineralogical Survey, Bulletin, 48, 249 p.Google Scholar
Hupé, P. 1953. Trilobita, p. 44–246 In Piveteau, J. (ed.), Traité de Paléontologie, Volume 3. Masson et Compagnie, Paris.Google Scholar
Ingham, J. K., and Tripp, R. P. 1991. The trilobite fauna of the Middle Ordovician Doularg Formation of the Girvan District, Scotland, and its paleoenvironmental significance. Transactions of the Royal Society of Edinburgh, Earth Sciences, 82:27–54.Google Scholar
Ingham, J. K., Curry, G. B., and Williams, A. 1985. Early Ordovician Dounans Limestone fauna, Highland Border Complex, Scotland. Transactions of the Royal Society of Edinburgh, Earth Sciences, 76:481–513.CrossRefGoogle Scholar
Jaanusson, V. 1956. On the trilobite genus Celmus Angelin, 1854. Bulletin of the Geological Institute of the University of Uppsala, 36:35–49.Google Scholar
Jell, P. A., and Stait, B. 1985. Tremadoc trilobites from the Florentine Valley Formation, Tim Shea area, Tasmania. Memoirs of the Museum of Victoria, 46:1–34.Google Scholar
Lee, D.-C., and Chatterton, B. D. E. 1996. Terminology of glabellar lobes in trilobite larvae based on homology. Journal of Paleontology, 70:439–442.CrossRefGoogle Scholar
Lee, D.-C., and Chatterton, B. D. E. 1997. Hystricurid trilobite larvae from the Garden City Formation (Lower Ordovician), Idaho and their phylogenetic implications. Journal of Paleontology, 71:862–877.Google Scholar
Lu, Y.-H. 1975. Ordovician trilobite faunas of central and southwestern China. Palaeontologia Sinica New Series B, 11, 463 p.Google Scholar
Lu, Y.-H., Chang, W.-T., Chu, C.-L., Chen, Y.-Y., and Hsiang, L.-W. 1965. Trilobites of China. Science Press, Beijing. (In Chinese).Google Scholar
Lu, Y.-H., Chu, C.-L., Chien, Y.-Y., Zhou, Z.-Y., Chen, J.-Y., Liu, C.-W., Yu, W., Chen, X., and Xu, H.-K. 1976. Ordovician biostratigraphy and paleozoogeography of China. Nanjing Institute of Geology and Paleontology, Academia Sinica, Memoir 7:1–83. (In Chinese).Google Scholar
Moore, R. C. (ed.). 1959. Treatise on Invertebrate Paleontology: (Part O) Arthropoda I. Geological Society of America and University of Kansas Press, Lawrence, Kansas, 560 p.Google Scholar
Morris, S. F. 1988. A review of British trilobites including a synoptic revision of Salter's monograph. Palaeontographical Society Monographs, 316 p.Google Scholar
Morris, S. F. and Tripp, R. P. 1986. Lectotype selections for Ordovician trilobites from the Girvan District, Strathclyde. Bulletin of the British Museum of natural History (Geology), 40:161–176.Google Scholar
Nixon, K. C., and Carpenter, J. M. 1993. On outgroups. Cladistics, 9:413–426.Google Scholar
Owen, A. W. 1981. The Ashgill trilobites of the Oslo region, Norway. Palaeontographica A, 175, 88p.Google Scholar
Owen, A. W., and Bruton, D. L. 1980. Late Caradoc-early Ashgill trilobites of the central Oslo Region, Norway. Paleontological Contributions from the University of Oslo, 245, 63p.Google Scholar
Owen, A. W., Tripp, R. P., and Morris, S. F. 1986. The trilobite fauna of the Raheen Formation (upper Caradoc), Co. Waterford, Ireland. Bulletin of the British Museum of Natural History, Geology, 40:91–122.Google Scholar
Owens, R. M. 1978. The trilobite genera Panarchaegonus Öpik, Isbergia Warburg and Cyamops gen. nov. from the Ordovician of Balto-Scandia and the British Isles. Norsk Geologisk Tiddskrift, 58:199–219.Google Scholar
Perry, D. G., and Chatterton, B. D. E. 1979. Wenlock trilobites and brachiopods from the Mackenzie Mountains, north-western Canada. Palaeontology, 22:569–607.Google Scholar
Raymond, P. E. 1925. Some trilobites of the lower Middle Ordovician of eastern North America. Bulletin of the Museum of Comparative Zoology at Harvard College, 17, 180 p.Google Scholar
Reed, F. R. C. 1896. The fauna of the Keisley Limestone, Part 1. Quarterly Journal of the Geological Society of London, 52:407–437.Google Scholar
Reed, F. R. C. 1904 (1903-1906). The Lower Palaeozoic trilobites of the Girvan District, Ayrshire. Palaeontographical Society Monographs, 186 p.Google Scholar
Reed, F. R. C. 1931. The Lower Palaeozoic trilobites of Girvan. Supplement number 2. Palaeontographical Society Monographs, 30 p.Google Scholar
Reed, F. R. C. 1935. The Lower Palaeozoic trilobites of Girvan. Supplement No. 3. Palaeontographical Society Monograph, 64 p.Google Scholar
Ross, R. J. Jr. 1951. Stratigraphy of the Garden City Formation in northeastern Utah, and its trilobite faunas. Peabody Museum of Natural History, Yale University, Bulletin 6, 161 p.Google Scholar
Shaw, F. C. 1968. Early Middle Ordovician (Chazy) trilobites of New York. New York State Museum and Science Service Memoir, 17, 163 p.Google Scholar
Stitt, J. H. 1983. Trilobites, biostratigraphy, and lithostratigraphy of the McKenzie Hill Limestone (Lower Ordovician), Wichita and Arbuckle Mountains, Oklahoma. Oklahoma Geological Survey, Bulletin, 134:1–54.Google Scholar
Swofford, D. L. 1993. PAUP. Phylogenetic analysis using parsimony. Version 3.1. Computer program distributed by the Illinois Natural History Survey, Champaign, Illinois.Google Scholar
Thorslund, P. 1940. On the Chasmops Series of Jemtland and Södermanland (Tvären). Sveriges Geologiska Undersökning. Avhhandlingar och Uppsatser (C), 436, 191 p.Google Scholar
Törnquist, S. L. 1884. Undersökningar öfver Siljanssimrådets Trilobitfauna. Sveriges Geologiska Udersökning, 66, 101 p.Google Scholar
Tripp, R. P. 1965. Trilobites from the Albany division (Ordovician) of the Girvan District, Ayrshire. Palaeontology, 8:577–603.Google Scholar
Tripp, R. P. 1967. Trilobites from the Upper Stinchar Limestone (Ordovician) of the Girvan District, Ayrshire. Transactions of the Royal Society of Edinburgh, 67:43–93.Google Scholar
Tripp, R. P. 1976. Trilobites of the basal superstes Mudstones (Ordovician) at Aldons Quarry, near Girvan, Ayrshire. Transactions of the Royal Society of Edinburgh, 69:369–423.Google Scholar
Tripp, R. P. 1979. Trilobites from the Ordovician Auchensoul and Stinchar limestones of the Girvan District, Strathclyde. Palaeontology, 22:339–361.Google Scholar
Tripp, R. P. 1980. Trilobites from the Ordovician Balclatchie and lower Ardwell groups of the Girvan district, Scotland. Transactions of the Royal Society of Edinburgh, Earth Sciences, 71:123–145.Google Scholar
Tripp, R. P., and Evitt, W. R. 1983. Silicified trilobites of the genus Dimeropyge from the Middle Ordovician of Virginia. Special Papers in Palaeontology, 30:229–240.Google Scholar
Warburg, E. 1925. The trilobites of the Leptaena Limestone in Dalarne with a discussion of the zoological position and the classification of the trilobites. Geological Institute of Uppsala, 17, 445 p.Google Scholar
Webby, B. D. 1974. Upper Ordovician trilobites from central New South Wales. Palaeontology, 17:203–252.Google Scholar
Weber, V. N. 1948. Trilobity silurijskich otlozhenij SSSR. 1. Nizhnesiluriskije trilobity. Monografii po paleontologii SSSR., 69:1–111.Google Scholar
Westrop, S. R., Knox, L. A., and Landing, E. 1993. Lower Ordovician (Ibexian) trilobites from the Tribes Hill Formation, central Mohawk Valley, New York State. Canadian Journal of Earth Sciences, 30:1618–1633.Google Scholar
Whittington, H. B. 1950. Sixteen Ordovician genotype trilobites. Journal of Paleontology, 24:531–565.Google Scholar
Whittington, H. B. 1956. Silicified Middle Ordovician trilobites: The Odontopleuridae. Bulletin of the Museum of Comparative Zoology at Harvard College, 114:155–288.Google Scholar
Whittington, H. B. 1963. Middle Ordovician trilobites from Lower Head, western Newfoundland. Bulletin of the Museum of Comparative Zoology at Harvard College, 129, 118 p.Google Scholar
Whittington, H. B. and Evitt, W. R. II. 1954. Silicified Middle Ordovician trilobites. Geological Society of America, Memoir, 59, 137 p.Google Scholar
Young, G. E. 1973. An Ordovician (Arenigian) trilobite faunule of great diversity in the Ibex area, western Utah. Brigham Young University Geology Studies, 20:91–115.Google Scholar
Zhou, Z.-Q., Li, J.-S., and Qu, X.-G. 1982. Class Trilobita, pp. 215–294. In Xian Institute of Geology and Mineral Resources (eds.). Paleontological Atlas of northwest China. Shuanxi-Gansu-Ninxia Volume. Part I. Precambrian and Early Paleozoic. Geological Publishing House, Beijing. (In Chinese).Google Scholar