Hostname: page-component-848d4c4894-ndmmz Total loading time: 0 Render date: 2024-05-31T03:11:27.278Z Has data issue: false hasContentIssue false

Trilobites of the suborder Illaenina from the Silurian of north Queensland, Australia

Published online by Cambridge University Press:  01 July 2016

David J. Holloway
Affiliation:
Invertebrate Palaeontology, Museum Victoria, GPO Box 666, Melbourne, Victoria 3001, Australia 〈dhollow@museum.vic.gov.au〉
Philip D. Lane
Affiliation:
School of Earth Sciences, William Smith Building, University of Keele, Staffordshire ST5 5BG, United Kingdom 〈p.d.lane@keele.ac.uk〉

Abstract

The trilobite fauna of the middle Silurian (Telychian to possibly earliest Sheinwoodian) Tomcat Creek limestone in the Broken River Province of north Queensland is dominated by the suborder Illaenina, including illaenimorphs (Illaeninae and Bumastinae) and members of the Scutelluidae. Scutelluidae are most diverse, with eight genera, of which Dolabrapex, Iotoryx, Perizostra, and Quintonia are new. Perizostra is the first scutelluid with a cephalon that may be described as of phacomorph appearance. Illaenimorphs are represented by three genera, including Opsypharus, which is regarded as a senior synonym of Paracybantyx but distinct from Failleana with which it has been placed in synonymy by some authors. Thirteen species are new: Cybantyx? ergodes, Opsypharus pandanensis, Australoscutellum talenti, Dolabrapex acomus, Illaenoscutellum psephos, Iotoryx clarksoni, Japonoscutellum mawsonae, J. drakton, J. fractum, Kosovopeltis avita, Perizostra campbelli, Quintonia arata, and Q. pavo. A species of Stenoparia is placed in open nomenclature. The species of Australoscutellum, Illaenoscutellum, and possibly Kosovopeltis are the oldest known representatives of those genera. These genera and Japonoscutellum are also common in faunas from limestones of Wenlock to Ludlow age in central western New South Wales, reflecting the similarity in lithofacies. The monotypic Late Ordovician genus Craigheadia, which has been regarded as a scutelluid, belongs to the Lichidae and is probably a junior synonym of Leiolichas.

Type
Articles
Copyright
Copyright © 2016, The Paleontological Society 

Access options

Get access to the full version of this content by using one of the access options below. (Log in options will check for institutional or personal access. Content may require purchase if you do not have access.)

References

Adrain, J.M., 2013, A synopsis of Ordovician trilobite distribution and diversity, in Harper, D.A.T., and Servais, T., eds., Early Palaeozoic Biogeography and Palaeogeography: Geological Society of London Memoir 38, p. 297–336.Google Scholar
Adrain, J.M., Chatterton, B.D.E., and Blodgett, R.B., 1995, Silurian trilobites from southwestern Alaska: Journal of Paleontology, v. 69, p. 723736.CrossRefGoogle Scholar
Alberti, G.K.B., 1970, Trilobiten des jüngeren Siluriums sowie des Unter- und Mitteldevons. 2: Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, v. 525, p. 1233.Google Scholar
Angelin, N.P., 1854, Palaeontologica Scandinavica. Pars II, Holmiae, Academiae Regiae Scientarium Suecanae, p. i–ix, 21–92.Google Scholar
Apollonov, M.K., 1980, Klass Trilobita. Trilobity, in Apollonov, M.K., Bandaletov, S.M., and Nikitin, I.F., eds., Granitsa Ordovika i Silura v Kazakhstane, Alma-Ata, Nauka, p. 86118.Google Scholar
Balashova, E.A., 1959, Sredne- i verkhneordovikskie i nizhnesiluriiskie trilobity Vostochnogo Taimyra i ikh stratigraficheskoe znachenie (Part 2): Sbornik statei po paleontologii i biostratigrafii, v. 15, p. 2755.Google Scholar
Balashova, E.A., 1960, Trilobity Srednego i Verkhnego Ordovika i Nizhnego Silura Vostochnogo Taimyra, Leningrad, Izdatel'stvo Leningradskogo Universiteta, 111 p.Google Scholar
Basse, M., 2009, Fossilium Catalogus I. Animalia. Catalogus Typorum Trilobitorum Germaniae: I. Pars 147. Trilobites Cambrici, Ordovicici et Silurici—Saxa Erratica Inclusa. II. Trilobites Devonici et Infracarbonici, Leiden, Backhuys, 380 p.Google Scholar
Billings, E., 1859, Descriptions of some new species of trilobites from the lower and middle Silurian rocks of Canada: Canadian Naturalist and Geologist, v. 4, p. 367383.Google Scholar
Bolton, T.E., 1981, Ordovician and Silurian biostratigraphy, Anticosti Island, Québec, in Lespérance, P.J., ed., IUGS Subcommission on Silurian Stratigraphy, Ordovician–Silurian Boundary Working Group, Field Meeting, Anticosti–Gaspé, Québec, 1981. Volume 2, Stratigraphy and Paleontology. Montréal, Département de Géologie, Université de Montréal, p. 41–59.Google Scholar
Chatterton, B.D.E., and Campbell, K.S.W., 1980, Silurian trilobites from near Canberra and some related forms from the Yass Basin: Palaeontographica, Abteilung A, v. 167, p. 77119, pls. 1–16.Google Scholar
Chatterton, B.D.E., and Ludvigsen, R., 1976, Silicified Middle Ordovician trilobites from the South Nahanni River area, District of Mackenzie, Canada: Palaeontographica, Abteilung A, v. 154, p. 1106, pls. 1–22.Google Scholar
Chatterton, B.D.E., and Ludvigsen, R., 2004, Early Silurian trilobites of Anticosti Island, Québec, Canada: Palaeontographica Canadiana, v. 22, p. 1264.Google Scholar
Cockle, P., 1999, Conodont data in relation to time, space and environmental relationships in the Silurian (late Llandovery–Ludlow) succession at Boree Creek (New South Wales, Australia): Abhandlungen der geologischen Bundesanstalt, v. 54, p. 107133.Google Scholar
Curtis, N.J., and Lane, P.D., 1997, The Llandovery trilobites of England and Wales. Part 1: Monograph of the Palaeontographical Society, v. 151, no. 605, p. 150, pls. 1–3.Google Scholar
Edgecombe, G.D., and Sherwin, L., 2001, Early Silurian (Llandovery) trilobites from the Cotton Formation, near Forbes, New South Wales, Australia: Alcheringa, v. 25, p. 87105.Google Scholar
Edgecombe, G.D., and Webby, B.D., 2007, Ordovician trilobites with eastern Gondwanan affinites from central-west New South Wales and Tasmania: Memoirs of the Association of Australasian Palaeontologists, v. 34, p. 255281.Google Scholar
Edgecombe, G.D., Banks, M.R., and Banks, D.M., 2004, Late Ordovician trilobites from Tasmania: Styginidae, Asaphidae and Lichidae: Memoirs of the Association of Australasian Palaeontologists, v. 30, p. 5799.Google Scholar
Foerste, A.F., 1885, The Clinton Group of Ohio: Bulletin of the Scientific Laboratories of Denison University, v. 1, p. 63120, pls. 13–14.Google Scholar
Fortey, R.A., and Owens, R.M., 1990, Trilobites, in McNamara, K.J., ed., Evolutionary Trends, London, Belhaven Press, p. 121142.Google Scholar
Fortey, R.A., and Owens, R.M., 1997, Evolutionary history, in Kaesler, R.L., ed., Treatise on Invertebrate Paleontology. Part O. Arthropoda 1. Trilobita, revised. Volume 1: Introduction, Order Agnostida, Order Redlichiida, Boulder, Colorado, and Lawrence, Kansas, Geological Society of America and University of Kansas Press, p. 249287.Google Scholar
Gaertner, H.R. von, 1930, Silurische und tiefunterdevonische Trilobiten und Brachiopoden aus den Zentralkarnischen Alpen: Jahrbuch der Preussischen Geologischen Landesanstalt und Bergakademie, v. 51, p. 188252, pls. 24–26.Google Scholar
Gümbel, C.W., 1879, Geognostische Beschreibung des Königreichs Bayern. 3. Geognostische Beschreibung des Fichtelgebirges mit dem Frankenwalde und dem westlichen Vorlande, Gotha, Justus Perthes, 697 p.Google Scholar
Hawle, I., and Corda, A.J.C., 1847, Prodrom einer Monographie der böhmischen Trilobiten, Prague, J. G. Calve, 176 p.Google Scholar
Helbert, G.J., Lane, P.D., Owens, R.M., Siveter, D.J., and Thomas, A.T., 1982, Lower Silurian trilobites from the Oslo Region, in Worsley, D., ed., International Union of Geological Sciences Subcommission on Silurian Stratigraphy. Field meeting, Oslo Region, 1982. Palaeontological Contributions from the University of Oslo No. 278, Oslo, Paleontologisk Museum, p. 129148.Google Scholar
Henderson, R.A., and Withnall, I.W., 2013, Broken River Province, in Jell, P.A., ed., Geology of Queensland, Brisbane, Geological Survey of Queensland, p. 250279.Google Scholar
Holloway, D.J., 1994, Early Silurian trilobites from the Broken River area, north Queensland: Memoirs of the Museum of Victoria, v. 54, p. 243269.Google Scholar
Holloway, D.J., and Lane, P.D., 1998, Effaced styginid trilobites from the Silurian of New South Wales: Palaeontology, v. 41, p. 853896.Google Scholar
Holloway, D.J., and Lane, P.D., 2012, Scutelluid trilobites from the Silurian of New South Wales: Palaeontology, v. 55, p. 413490.Google Scholar
Holloway, D.J., and Sandford, A.C., 1993, An early Silurian trilobite fauna from Tasmania: Memoirs of the Association of Australasian Palaeontologists, v. 15, p. 85102.Google Scholar
Holm, G., 1883, De Svenska arterna af trilobitslägtet Illaenus Dalman: Bihang till Kungliga Svenska Vetenskaps-Akademiens Handlingar, v. 7, no. 3, p. 1148.Google Scholar
Holm, G., 1886, Revision der ostbaltischen silurischen Trilobiten, von Fr. Schmidt. Abtheilung 3. Illaeniden: Mémoires de l'Académie Impériale des Sciences de St-Pétersbourg, v. 33, no. 8, p. i–ii, 1–173, pls. 1–12. [=Zapiski Imperatorskoi Akademii Nauk.]Google Scholar
Howells, Y., 1982, Scottish Silurian trilobites: Monograph of the Palaeontographical Society, v. 135, no. 561, p. 176, pls. 1–15.Google Scholar
Hughes, H.E., and Thomas, A.T., 2011, Trilobite associations, taphonomy, lithofacies and environments of the Silurian reefs of North Greenland: Palaeogeography, Palaeoclimatology, Palaeoecology, v. 302, p. 142155.Google Scholar
Hughes, H.E., and Thomas, A.T., 2014, Trilobites from Silurian reefs in North Greenland: Special Papers in Palaeontology, v. 92, p. 1102.Google Scholar
Hupé, P., 1955, Classification des trilobites (Part 2): Annales de Paléontologie, v. 41, p. 91325.Google Scholar
Jaanusson, V., 1959, Family Illaenidae Hawle & Corda, 1847, in Moore, R.C., ed., Treatise on Invertebrate Paleontology. Part O. Arthropoda 1, Boulder, Colorado, and Lawrence, Kansas, Geological Society of America and University of Kansas Press, p. 372376.Google Scholar
Jell, P.A., and Adrain, J.M., 2003, Available generic names for trilobites: Memoirs of the Queensland Museum, v. 48, p. 331551.Google Scholar
Kácha, P., and Šarič, R., 1991, Ontogeny of the trilobite Kosovopeltis svobodai Šnajdr from the Bohemian Silurian: Věstník Ústředního ústavu geologického, v. 66, p. 257273, pls. 1–6.Google Scholar
Kiaer, J., 1908, Das Obersilur im Kristianiagebiete. Eine stratigraphisch-faunistische Untersuchung: Skrifter udgivne af Videnskabs-Selskabet i Christiania. 1. Matematisk-Naturvidenskabelig Klasse, v. 1906(2), p. ixvi, 1–596 [volume for 1906].Google Scholar
Kobayashi, T., and Hamada, T., 1965, An occurrence of a new Scutellum in the Silurian of Shikoku Island: Transactions and Proceedings of the Palaeontological Society of Japan, v. 58, p. 7481.Google Scholar
Kobayashi, T., and Hamada, T., 1974, Silurian trilobites of Japan in comparison with Asian, Pacific and other faunas: Special Papers of the Palaeontological Society of Japan, v. 18, p. iviii, 1–155, pls. 1–12.Google Scholar
Koroleva, M.N., 1984, Ordovikskie trilobity semeistva Scutellidae v Kazakhstane: Paleontologicheskii Zhurnal, v. 1984, no. 4, p. 8087, pl. 8. [English translation published 1985 in Paleontological Journal, v. 18 no. 4, p. 79–87.]Google Scholar
Lane, P.D., 1984, Silurian trilobites from Hall Land and Nyeboe Land, western North Greenland: Rapport Grønlands Geologiske Undersøgelse, v. 121, p. 5375.CrossRefGoogle Scholar
Lane, P.D., and Siveter, D.J., 1991, A Silurian trilobite fauna dominated by Calymene from Kap Tyson, Hall Land, western North Greenland: Rapport Grønlands Geologiske Undersøgelse, v. 150, p. 514.CrossRefGoogle Scholar
Lane, P.D., and Thomas, A.T., 1978a, Silurian trilobites from NE Queensland and the classification of effaced trilobites: Geological Magazine, v. 115, p. 351358.Google Scholar
Lane, P.D., and Thomas, A.T., 1978b, Family Scutelluidae Richter & Richter, 1955, in Thomas, A.T., British Wenlock trilobites. Part 1: Monograph of the Palaeontographical Society, v. 132, no. 552, p. 829, pls. 1–7.Google Scholar
Lane, P.D., and Thomas, A.T., 1980, A replacement name for Rhax Lane & Thomas, 1978 (Trilobita) non Hermann, 1804: Geological Magazine, v. 117, p. 191.CrossRefGoogle Scholar
Lane, P.D., and Thomas, A.T., 1983, A review of the trilobite Suborder Scutelluina: Special Papers in Palaeontology, v. 30, p. 141160.Google Scholar
Lane, P.D., and Wu, H.-J., 2002, Trilobites, in Holland, C.H., and Bassett, M.G., eds., Telychian Rocks of the British Isles and China (Silurian, Llandovery Series): An Experiment to Test Precision in Stratigraphy. National Museum of Wales Geological Series No. 21, Cardiff, National Museum of Wales, p. 142146.Google Scholar
Lerosey-Aubril, R., and McNamara, K.J., 2008, The cephalic median organ of trilobites, in Rábano, I., Gozalo, R., and García-Bellido, D., eds., Advances in Trilobite Research. Cuadernos del Museo Geominero no. 9, Madrid, Instituto Geológico y Minero de España, p. 229235.Google Scholar
Li, S.-J., 1978, Trilobita Walch, 1771, in Southwest Geological Research Institute, ed., Palaeontological Atlas of Southwest China. Sichuan Volume. 1. Sinian to Devonian: Beijing, Geological Publishing House, p. 179–284 [in Chinese].Google Scholar
Lin, T.-R., 1987, Two new Thysanopeltidae trilobite genera from lower Silurian of western Hunan: Acta Palaeontologica Sinica, v. 26, p. 746752, pls. 1–2 [in Chinese].Google Scholar
Lindström, G., 1885, Förteckning på Gotlands Siluriska crustacéer: Öfversigt af Kungliga Svenska Vetenskapsakademiens Förhandlingar, v. 42, no. 6, p. 3799, pls. 12–16.Google Scholar
Liu, Y.-R., 1982, Trilobita, in Geological Bureau of Hunan, ed., Palaeontological Atlas of Hunan. Geological Memoirs of the Ministry of Geology and Mineral Resources, People's Republic of China, Series 2, Number 1: Beijing, Geological Publishing House, p. 290–347 [in Chinese].Google Scholar
Ludvigsen, R., and Chatterton, B.D.E., 1980, The ontogeny of Failleana and the origin of the Bumastinae (Trilobita): Geological Magazine, v. 117, p. 471478, pl. 1.Google Scholar
Ludvigsen, R., and Tripp, R.P., 1990, Silurian trilobites from the northern Yukon Territory: Life Sciences Contributions, Royal Ontario Museum, v. 153, p. iiv, 1–59.Google Scholar
Mikulic, D.G., 1999, Silurian trilobite associations in North America, in Boucot, A.J., and Lawson, J.D., eds., Paleocommunities: A Case Study from the Silurian and Lower Devonian, Cambridge, Cambridge University Press, p. 793798.Google Scholar
Mitchell, J., 1887, On some new trilobites from Bowning, NSW: Proceedings of the Linnean Society of New South Wales, v. 2, p. 435440, pl. 16.Google Scholar
Morris, S.F., 1988, A review of British trilobites, including a synoptic revision of Salter's monograph: Monograph of the Palaeontographical Society, v. 140, no. 574, p. 1316.Google Scholar
Morris, S.F., and Tripp, R.P., 1986, Lectotype selections for Ordovician trilobites from the Girvan district, Strathclyde: Bulletin of the British Museum (Natural History), Geology Series, v. 40, p. 161176.Google Scholar
Munson, T.J., and Jell, J.S., 1999, Llandovery rugose corals from the Quinton Formation, Broken River Province, northeast Queensland: Memoirs of the Association of Australasian Palaeontologists, v. 21, p. 165.Google Scholar
Nicholson, H.A., and Etheridge, R., 1879, A monograph of the Silurian fossils of the Girvan district in Ayrshire with special reference to those contained in the ‘Gray Collection.’ Volume 1. Fasciculus 2, Edinburgh, William Blackwood and Sons, p. 137207.Google Scholar
Norford, B.S., 1994, Biostratigraphy and trilobite fauna of the lower Silurian Tegart Formation, southeastern British Columbia: Bulletin of the Geological Survey of Canada, v. 479, p. 1347.Google Scholar
Pickett, J.W., et al, 2000, Silurian palaeobiogeography of Australia: Memoirs of the Association of Australasian Palaeontologists, v. 23, p. 127165.Google Scholar
Poulsen, C., 1934, The Silurian faunas of North Greenland. I. The fauna of the Cape Schuchert Formation: Meddelelser om Grønland, v. 72, p. 146.Google Scholar
Prantl, F., and Přibyl, A., 1947, Classification and division of the genus Scutellum Pusch, 1833, from the Barrandian (central Bohemia): Bulletin international de l'Académie tchèque des Sciences, v. 47, no. 9, p. 4778, pls. 1–7.Google Scholar
Přibyl, A., and Vaněk, J., 1971, Studie über die Familie Scutelluidae Richter et Richter (Trilobita) und ihre phylogenetische Entwicklung: Acta Universitatis Carolinae, Geologica, v. 1971, no. 4, p. 361394, pls. 1–10.Google Scholar
Raymond, P.E., 1910, Notes on Ordovician trilobites. 4. New and old species from the Chazy: Annals of the Carnegie Museum, v. 7, p. 6080, pls. 17–19.Google Scholar
Raymond, P.E., 1916, New and old Silurian trilobites from southeastern Wisconsin, with notes on the genera of the Illaenidae: Bulletin of the Museum of Comparative Zoology, Harvard University, v. 60, p. 141, pls. 1–4.Google Scholar
Reed, F.R.C., 1904, The lower Palaeozoic trilobites of the Girvan district, Ayrshire. Part 2: Monograph of the Palaeontographical Society, v. 58, no. 276, p. 4996, pls. 7–13.Google Scholar
Reed, F.R.C., 1906, The lower Palaeozoic trilobites of the Girvan district, Ayrshire. Part 3: Monograph of the Palaeontographical Society, v. 60, no. 286, p. 97186, pls. 14–20.Google Scholar
Reed, F.R.C., 1928, Notes on the Bronteidae [= Goldiidae]: Annals and Magazine of Natural History, Series 10, v. 1, p. 4978.Google Scholar
Richter, R., and Richter, E., 1955, Scutelluidae n.n. (Tril.) durch 'kleine Änderung' eines Familien-Namens wegen Homonymie: Senckenbergiana lethaea, v. 36, p. 291293.Google Scholar
Richter, R., and Richter, E., 1956a, Ergänzendes zu den Scutelluidae mit Ausblick auf den Sprossungs-Gürtel der Trilobiten: Senckenbergiana lethaea, v. 37, p. 467485.Google Scholar
Richter, R., and Richter, E., 1956b, Grundlagen für die Beurteilung und Einteilung der Scutelluidae (Tril.): Senckenbergiana lethaea, v. 37, p. 79124.Google Scholar
Richter, R., and Richter, E., 1959, Family Thysanopeltidae Hawle & Corda, 1847, in Moore, R.C., ed., Treatise on Invertebrate Paleontology. Part O. Arthropoda 1, Boulder, Colorado, and Lawrence, Kansas, Geological Society of America and University of Kansas Press, p. 367372.Google Scholar
Rickards, R.B., and Jell, J.S., 2002, New graptolite faunas from the Llandovery, lower Silurian of the Graveyard Creek Subprovince, Broken River region, Queensland, Australia: Proceedings of the Geologists’ Association, v. 113, p. 111120.Google Scholar
Salter, J.W., 1867, A monograph of the British trilobites from the Cambrian, Silurian and Devonian formations. Part 4: Monograph of the Palaeontographical Society, v. 20, no. 86, p. 177214, pls. 25*–30.Google Scholar
Sandford, A.C., and Holloway, D.J., 2006, Early Silurian phacopide trilobites from central Victoria, Australia: Memoirs of Museum Victoria, v. 63, p. 215255.Google Scholar
Santel, W., 2001, Trilobiten aus dem Silur der Karnischen Alpen/Österreich. Teil 1: Palaeontographica, Abteilung A, v. 262, p. 87191, pls. 1–9.Google Scholar
Schmidt, F., 1885, Revision der ostbaltischen silurischen Trilobiten. Abtheilung 2. Acidaspiden und Lichiden: Mémoires de l'Academie Impériale des Sciences de St-Pétersbourg, Series 7, v. 33, no. 1, p. 1–127.Google Scholar
Shaw, F.C., 1968, Early Middle Ordovician Chazy trilobites of New York: New York State Museum Memoir, v. 17, p. ivii, 1–163.Google Scholar
Shaw, F.C., and Bolton, T.E., 2011, Ordovician trilobites from the Romaine and Mingan formations (Ibexian–late Whiterockian), Mingan Islands, Quebec: Journal of Paleontology, v. 85, p. 406441.Google Scholar
Simpson, A., 1999, Early Silurian conodonts from the Quinton Formation of the Broken River region (north-eastern Australia): Abhandlungen der geologischen Bundesanstalt, v. 54, p. 181199.Google Scholar
Sinclair, G.W., 1949, The Ordovician trilobite Eobronteus : Journal of Paleontology, v. 23, p. 4556.Google Scholar
Sloan, T.R., et al, 1995, Conodont data from Silurian–Middle Devonian carbonate fans, debris flows, allochthonous blocks and adjacent autochthonous platform margins: Broken River and Camel Creek areas, north Queensland, Australia: Courier Forschungsinstitut Senckenberg, v. 182, p. 177.Google Scholar
Šnajdr, M., 1957, Klasifikace čeledě Illaenidae (Hawle a Corda) v českém starším paleozoiku: Sborník Ústředního ústavu geologického, Oddíl paleontologický, v. 23, p. 125284, pls. 1–12.Google Scholar
Šnajdr, M., 1958, Několik nových rodů trilobitů z čeledě Scutelluidae: Věstník Ústředního ústavu geologického, v. 33, p. 177184, pls. 1–2.Google Scholar
Šnajdr, M., 1960, Studie o čeledi Scutelluidae (Trilobitae): Rozpravy Ústředního ústavu geologického, v. 26, p. 1264, pls. 1–36.Google Scholar
Suzuki, Y., and Bergström, J., 1999, Trilobite taphonomy and ecology in Upper Ordovician carbonate buildups in Dalarna, Sweden: Lethaia, v. 32, p. 159172.Google Scholar
Talent, J.A., Mawson, R., Simpson, A., and Brock, G.A., 2002, Palaeozoics of NE Queensland: Broken River region. Ordovician–Carboniferous of the Townsville Hinterland: Broken River and Camel Creek Regions, Burdekin and Clarke River Basins. IPC2002 Field Excursion Guidebook, Sydney, Macquarie University Centre for Ecostratigraphy and Palaeobiology, 82 p.Google Scholar
Thomas, A.T., and Holloway, D.J., 1988, Classification and phylogeny of the trilobite order Lichida: Philosophical Transactions of the Royal Society of London, Series B, Biological Sciences, v. 321, no. 1205, p. 179262.Google Scholar
Thomas, A.T., and Lane, P.D., 1999, Trilobite assemblages of the North Atlantic region, in Boucot, A.J., and Lawson, J.D., eds., Paleocommunities: A Case Study from the Silurian and Lower Devonian, Cambridge, Cambridge University Press, p. 444457.Google Scholar
Tremblay, J.V., and Westrop, S.R., 1991, Middle Ordovician (Whiterockian) trilobites from the Sunblood Formation, District of Mackenzie, Canada: Journal of Paleontology, v. 65, p. 801824.Google Scholar
Umeda, M., 1998, The Siluro-Devonian Yokokurayama Group in the Yokokurayama area, Kôchi, southwest Japan: Journal of the Geological Society of Japan, v. 104, p. 365376 [in Japanese].Google Scholar
Warburg, E., 1925, The trilobites of the Leptaena Limestone in Dalarne: Bulletin of the Geological Institutions of the University of Uppsala, v. 17, p. ivi, 1–446, pls. 1–11.Google Scholar
Webby, B.D., 1974, Upper Ordovician trilobites from central New South Wales: Palaeontology, v. 17, p. 203252.Google Scholar
Weber, V.N., 1945, O rode Thysanopeltis Corda i nekotorykh ural'skikh Bronteidae: Materialy Vsesoyuznyi Nauchno-Issledovatel'skii Geologicheskii Institut, Paleontologiya i Stratigrafiya, Sbornik, v. 4, p. 2445, pls. 3–5.Google Scholar
Weber, V.N., 1948, Trilobity siluriiskikh otlozhenii SSSR. Vypusk 1. Nizhnesiluriiskie trilobity: Monografii po Paleontologii SSSR, v. 69, no. 4, p. 1–114, pls. 1–11.Google Scholar
Weber, V.N., 1951, Verkhnesiluriiskie trilobity SSSR: Trudy Vsesoyuznogo Nauchno-Issledovatel'skogo Geologicheskogo Instituta, v. 2, p. 172.Google Scholar
White, D.A., 1965, The geology of the Georgetown/Clarke River area, Queensland: Bulletin of the Bureau of Mineral Resources, Geology and Geophysics, v. 71, p. ivi, 1–165, pls. 1–8.Google Scholar
Whitfield, R.P., 1880, Descriptions of new species of fossils from the Paleozoic formations of Wisconsin: Annual Report of the Wisconsin Geological Survey, v. for 1897, p. 4471.Google Scholar
Whittard, W.F., 1938, The upper Valentian trilobite fauna of Shropshire: Annals and Magazine of Natural History, Series 10, v. 1, p. 85140.Google Scholar
Whittard, W.F., 1939, The Silurian illaenids of the Oslo Region: Norsk Geologisk Tidsskrift, v. 19, p. 275295, pls. 1–4.Google Scholar
Whittington, H.B., 1992, Trilobites, Woodbridge, Suffolk, Boydell Press, 145 p.Google Scholar
Whittington, H.B., 1997, Illaenidae (Trilobita): Morphology of thorax, classification, and mode of life: Journal of Paleontology, v. 71, p. 878896.Google Scholar
Whittington, H.B., 1999, Siluro-Devonian Scutelluinae (Trilobita) from the Czech Republic: Morphology and classification: Journal of Paleontology, v. 73, p. 414430.Google Scholar
Whittington, H.B., 2000, Stygina, Eobronteus (Ordovician Styginidae, Trilobita): Morphology, classification, and affinities of Illaenidae: Journal of Paleontology, v. 74, p. 879889.Google Scholar
Whittington, H.B., and Kelly, S.R.A., 1997, Morphological terms applied to Trilobita, in Kaesler, R.L., ed., Treatise on Invertebrate Paleontology. Part O. Arthropoda 1. Trilobita, revised. Volume 1: Introduction, Order Agnostida, Order Redlichiida, Boulder, Colorado, and Lawrence, Kansas, Geological Society of America and University of Kansas Press, p. 313329.Google Scholar
Withnall, I.W., 1993, Quinton Formation, in Withnall, I.W., and Lang, S.C., eds., Geology of the Broken River Province, North Queensland. Queensland Geology 4, Brisbane, Queensland Department of Minerals and Energy, p. 5662.Google Scholar
Wu, H.-J., 1977, Comments on new genera and species of Silurian–Devonian trilobites in southwest China and their significance: Acta Palaeontologica Sinica, v. 16, p. 95119, pls. 1–3 [in Chinese].Google Scholar
Zhang, W.-T., 1974, [Silurian] Trilobites, in Nanjing Insitute of Geology and Palaeontology, Academia Sinica, ed., A handbook of the Stratigraphy and Palaeontology in Southwest China: Beijing, Science Press, p. 173–187 [in Chinese].Google Scholar
Zhou, Z.-Y., Yin, G.-Z., and Zhou, Z.-Q., 2014, Ordovician (Darriwilian–early Katian) trilobite faunas of northwestern Tarim, Xinjiang, China: Memoirs of the Association of Australasian Palaeontologists, v. 46, p. 1142.Google Scholar