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    Huang, Jiandong Sinitshenkova, N. D. and Ren, Dong 2011. New mayfly nymphs (Insecta: Ephemeroptera) from Yixian Formation, China. Paleontological Journal, Vol. 45, Issue. 2, p. 167.

    Zhang, HaiChun Wang, Bo and Fang, Yan 2010. Evolution of insect diversity in the Jehol Biota. Science China Earth Sciences, Vol. 53, Issue. 12, p. 1908.

    Zhang, Jun-Feng and Kluge, N. J. 2007. Jurassic larvae of mayflies (Ephemeroptera) from the Daohugou formation in Inner Mongolia, China. Oriental Insects, Vol. 41, Issue. 1, p. 351.

    Zhang, J. -f. 2006. New mayfly nymphs from the Jurassic of Northern and Northeastern China (Insecta: Ephemeroptera). Paleontological Journal, Vol. 40, Issue. 5, p. 553.


Description of Caenoephemera shangyuanensis, gen.nov., sp.nov. (Ephemeroptera), from the Yixian Formation

  • Qi-Bin Lin (a1) and Di-Ying Huang (a1)
  • DOI:
  • Published online: 01 May 2012

A fossil mayfly nymph Caenoephemera shangyuanensis, gen.nov., sp.nov., discovered in the lower part of the Upper Jurassic – Lower Cretaceous Yixian Formation of Jianshangou, Beipiao district, West Liaoning, China, is described. It has several archaic features typical of Paleozoic mayfly nymphs, such as 5 tarsal segments and hind wing pads not covered by fore wing pads, but also exhibits specialized modern features, such as a short metathorax, abdominal tracheal gills, and cerci. It appears to be intermediate in structure between Paleozoic and extant mayfly nymphs.


On trouvera ici la description de Caenoephemera shangyuanensis, gen.nov., sp.nov., à partir d’une larve d’éphémère fossile découverte à la base de la formation Yixian (Jurassique supérieur – Crétacé inférieur, Jianshangou, district de Beipiao, Ouest Liaoning, Chine). Ce fossile présente non seulement plusieurs caractères archaïques typiques des éphémères paléozoïques comme des tarses à 5 articles et des fourreaux d’ailes postérieures non couverts par ceux des ailes antérieures, mais également des structures nettement "modernes" comme un métathorax court, des branchies abdominales et des cerques. Ce fossile semble être "intermédiaire" entre les éphémères paléozoïques et modernes.

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P-J Chen , Z-M Dong , S-N Zhen . 1998. An exceptionally well-preserved theropod dinosaur from the Yixian Formation of China. Nature (London) 391(1): 147–52

J. Kukalova-Peck 1985. Ephemeroid wing venation based upon new gigantic carboniferous mayflies and basic morphology, phylogeny, and metamorphosis of pterygote insects (Insecta, Ephemerida). Canadian Journal of Zoology 63: 933–55

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The Canadian Entomologist
  • ISSN: 0008-347X
  • EISSN: 1918-3240
  • URL: /core/journals/canadian-entomologist
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