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  • James E. O’Hara (a1)

Eight species of the siphonine genus Actia Robineau-Desvoidy are recognized in the Nearctic Region, including four that are new: A. dasymyia sp.nov. (type-locality 21 mi. east Tuktoyaktuk, North West Territories, Canada), A. dimorpha sp.nov. (type-locality Sapelo Island, Georgia, USA), A. radialis sp.nov. (type-locality Gatineau Park, Quebec, Canada), and A. sternalis sp.nov. (type-locality mi. 51 Dempster Highway, Yukon, Canada). Adults of the eight species are keyed, described, illustrated (heads, wings, and male terminalia), and their distributions mapped. The species are arranged into three species groups: the autumnalis-group with Nearctic members A. autumnalis (Townsend), A. rufescens (Greene), A. diffidens Curran, and A. dimorpha sp.nov.; the lamia-group with Nearctic members A. interrupta Curran, A. dasymyia sp.nov., and A. radialis sp.nov.; and the crassicornis-group with the single Nearctic member A. sternalis sp.nov. Nearctic Actia are inferred to be of relatively recent origin because only the A. autumnalis/A. rufescens species pair seems to have speciated within the Nearctic Region. Actia autumnalis/A. rufescens, A. diffidens, and A. dimorpha are each hypothesized to have their closest phylogenetic affinities with Neotropical species, and A. sternalis, A. interrupta, A. dasymyia, and questionably A. radialis are each hypothesized to have their closest affinities with Palearctic species.

Huit espèces siphonines du genre Actia Robineau-Desvoidy sont reconnues de la région néarctique, y compris quatre espèces nouvelles : A. dasymyia sp.nov. (site-type 21 mi. à l’est de Tuktoyaktuk, Territoires-du-nord-ouest, Canada), A. dimorpha sp.nov. (site-type Sapelo Island, Georgia, USA), A. radialis sp.nov. (site-type Parc Gatineau, Québec, Canada), et A. sternalis sp.nov. (site-type 51e mille route Dempster, Yukon, Canada). Les individus adultes ont été clavetés, décrits, illustrés (têtes, ailles et terminaux mâles) et leurs dispersions ont été entrées sur une carte. Les espèces ont été rangées dans trois groupes d’espèces : le groupe autumnalis avec des membres néarctiques A. autumnalis (Townsend), A. rufescens (Greene), A. diffidens Curran et A. dimorpha sp.nov.; le groupe lamia avec des membres néarctiques A. interrupta Curran, A. dasymyia sp.nov. et A. radialis sp.nov.; et le groupe crassicornis avec le membre néarctique unique A. sternalis sp.nov. Il a été conclu que les Actia néarctiques sont d’origine relativement récente, parce que la paire-espèce A. autumnalis/A. rufescens seule semble d’être devenue une espèce en dedans de la région néarctique. Il est proposé que Actia autumnalis/A. rufescens, A. diffidens et A. dimorpha ont leurs affinités les plus proches avec les espèces néarctiques, et que A. sternalis, A. interrupta, A. dasymyia et peut-être A. radialis ont leurs affinités les plus proches avec les espèces paléarctiques.

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The Canadian Entomologist
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