The nuculanid bivalve Costinuculana magharensis new genus new species is described from the middle to upper Bathonian Kehailia Formation of Gebel Maghara, North Sinai, Egypt. Costinuculana differs from other genera of the family Nuculanidae by the presence of opisthocline ribs along the rostrum. These ribs are variable in shape and size, straight to folded posteriorly, bifurcate ventrally and occasionally postero-dorsally, and cover an area ~45% of the total valve length from the posterior end. The life position of C. magharensis n. gen. n. sp. is reconstructed on the basis of a functional interpretation of its morphology and by comparison with closely related Recent forms. The asymmetrical commarginal ribs facilitated the burrowing process. The posterior oblique ribs are asymmetrical in cross-section with a steeply concave side in the burrowing direction and slightly convex side in the opposite direction. They probably kept the bivalve in a stable position once the desired depth had been reached. The thick oblique ribs probably also increased the strength of the rostrum and offered resistance against durophagous predators, being presumably partly exposed above the sediment-water interface. Based on the associated fauna, Costinuculana n. gen. lived in a low-energy environment characterized by a fine-grained, soft substrate.
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