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Late Ordovician to Early Silurian conodont faunas from the Kolyma Terrane, Omulev Mountains, northeast Russia, and their paleobiogeographic affinity

Published online by Cambridge University Press:  14 July 2015

Shunxin Zhang
Affiliation:
School of Earth and Ocean Sciences, University of Victoria, P.O. Box 1700 STN CSC, Victoria, British Columbia, Canada V8W 2Y2, ,
Christopher R. Barnes
Affiliation:
School of Earth and Ocean Sciences, University of Victoria, P.O. Box 1700 STN CSC, Victoria, British Columbia, Canada V8W 2Y2, ,

Abstract

The Late Ordovician and Early Silurian conodont faunas of northeast Russia are described, based on 39 conodont-bearing samples collected from Kanyon, Padun, Tirekhtyakh, Maut, and lower Sandugan Formations at Mirny Creek and Ina River sections, Omulev Mountains. The faunas are represented by a total of 776 identifiable conodont specimens assigned to 35 species representing 25 genera, among which a new species, Periodon mirnyensis, is established. Based on graptolite zones at the same locality and other conodont faunas worldwide, the faunas are assigned to the Caradocian, early Ashgillian, late Ashigillian, early Llandovery, and late Llandovery. The Ordovician faunas belong to outer platform to upper slope facies with a North Atlantic Realm affinity; the Silurian faunas are from slope and basin facies. The conodont faunas are the first to be described from the Kolyma Terrane, which has an enigmatic paleogeographic position during the Early Paleozoic.

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Research Article
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Copyright © The Paleontological Society 

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References

Aldridge, R. J., and Jeppsson, L. 1984. Ecological specialists among Silurian conodonts. Palaeontology, 32:141149.Google Scholar
Aldridge, R. J., and Mabillard, J. E. 1981. Local variations in the distribution of Silurian conodonts: An example from the amorphognathoides interval of the Welsh Basin, p. 1017. In Neale, J. W. and Brasier, M. D. (eds.), Microfossils from Recent and Fossil Shelf Seas. Ellis Horwood, Chichester.Google Scholar
Armstrong, H. A. 1990. Conodonts from the Upper Ordovician-Lower Silurian carbonate platform of North Greenland. Bulletin Grønlands Geologiske Undersogelse, 159, 151 p.Google Scholar
Bergstróm, S. M. 1964. Remarks on some Ordovician conodont faunas from Wales. Acta Universitatis Lundensis, Section II, No. 3, 66 p.Google Scholar
Bergström, S. M. 1971. Conodont biostratigraphy of the Middle and Upper Ordovician of Europe and Eastern North America. Geological Society of America Memoir, 127:83162.CrossRefGoogle Scholar
Bergström, S. M. 1973. Ordovician conodonts, p. 4758. In Hallam, A. (ed.), Atlas of Palaeobiogeography. Elsevier Scientific Publishing Company, Amsterdam.Google Scholar
Bergström, S. M., and Sweet, W. C. 1966. Conodonts from the Lexington Limestone (Middle Ordovician) of Kentucky and its lateral equivalents in Ohio and Indiana. Bulletin of American Paleontology, 50(229):271441.Google Scholar
Bischoff, G. C. O. 1986. Early and Middle Silurian conodonts from midwestern New South Wales. Courier Forschungsinstitut Senckenberg, 89, 337 p.Google Scholar
Bischoff, G. C. O., and Sanneman, D., 1958. Unterdevonische Conodonten aus dem Franken-wald. Notizblatt des hessischen Landes-amtes für Bodenforschung zu Weisbaden, 86:87110.Google Scholar
Blodgett, R. B. 1998. Emsian (late Early Devonian) fossils indicate a Siberian origin for the Farewell Terrane, p. 5361. In Clough, J. G. and Larson, F. (eds.), Short Notes on Alaska Geology 1997. Alaska Division of Geological and Geophysical Surveys Professional Report 118.CrossRefGoogle Scholar
Blodgett, R. B., Rohr, D. M., and Boucot, A. J. 2002. Paleozoic links among some Alaskan accreted terranes and Siberia based on megafossils. Geological Society of America Special Paper, 360:273290.Google Scholar
Branson, E. B., and Branson, C. C. 1947. Lower Silurian conodonts from Kentucky. Journal of Paleontology, 21:549556.Google Scholar
Branson, E. B., and Mehl, M. G. 1933. Conodonts from the Maquoketa-Thebes (Upper Ordovician) of Missouri. The University of Missouri Studies, 8:121131.Google Scholar
Branson, E. B., Mehl, M. G., and Branson, C. C. 1951. Richmond conodonts of Kentucky and Indiana. Journal of Paleontology, 25:117.Google Scholar
Cooper, B. J. 1975. Multielement conodonts from the Brassfield Limestone (Silurian) of southern Ohio. Journal of Paleontology, 49:9841008.Google Scholar
Craig, W. W., Ethington, R. L., and Repetski, J. E. 1986. Guidebook to the conodont paleontology of uppermost Lower Ordovician through Silurian strata, northeastern Arkansas. Pander Society Field Trip No. 5, Annual Meeting of the South-Central and Southeastern Sections of the Geological Society of America, Memphis, 57 p.Google Scholar
Drygant, D. M. 1974. Prostye konodonty silura i nizov devona Volyno-Podol'ia (Simple conodonts from the Silurian and lowermost Devonian of the Volyno-Podolia). Paleontologicheskiy sbornik, 10:6470.Google Scholar
Dumoulin, J. A., Harris, A. G., Gagiev, M., Bradley, D. C., and Repetski, J. E. 2002. Lithostratigraphic, conodont, and other faunal links between Lower Paleozoic strata in north and central Alaska and northeastern Russia. Geological Society of America Special Paper, 360:291312.Google Scholar
Dzik, J. 1976. Remarks on the evolution of Ordovician conodonts. Acta Palaeontologica Polonica, 21:395455.Google Scholar
Dzik, J. 1994. Conodonts from the Mójcza Limestone. Palaeontologia Polonica, 53:43128.Google Scholar
Dzik, J. 1999. Evolution of the Late Ordovician high-latitude conodonts and dating of Gondwana glaciations, p. 237253. In Serpagli, E. and Corradini, C. (eds.), Studies on Conodonts—Proceedings of the Seventh European Conodont Symposium. Bollettino della Società Paleontologica Italiana, 37 (2–3).Google Scholar
Ethington, R. L. 1959. Conodonts of the Ordovician Galena Formation. Journal of Paleontology, 33:257292.Google Scholar
Ferretti, A., and Barnes, C. R. 1997. Upper Ordovician conodonts from the Kalkbank Limestone of Thuringia, Germany. Palaeontology, 40:1542.Google Scholar
Ferretti, A., and Serpagli, E. 1999. Late Ordovician conodont faunas from southern Sardinia, Italy: biostratigraphic and paleogeographic implications, p. 215236. In Serpagli, E. and Corradini, C. (eds.), Studies on Conodonts—Proceedings of the Seventh European Conodont Symposium. Bollettino della Società Paleontologica Italiana, 37 (2–3).Google Scholar
Fortey, R. A., and Barnes, C. R. 1977. Early Ordovician conodont and trilobite communities of Spitsbergen: Influence on biogeography. Alcheringa, 1:297309.CrossRefGoogle Scholar
Hadding, Assor. 1913. Undre dicellograptusskiffern I Skäne jämte några därmed ekvivalenta bildningar. Lunds Universitets Årsskrift N. F. Avd. 2, 9(15):190.Google Scholar
Harris, A. G., Dumoulin, J. A., Repetski, J. E., and Carter, C. 1995. Correlation of Ordovician rocks of Northern Alaska, p. 2126. In Cooper, J. D., Droser, M. L., and Finney, S. C. (eds.), Ordovician Odyssey: Short papers for the Seventh International Symposium on the Ordovician System. Society of Economic Paleontologists and Mineralogists, Las Vegas.Google Scholar
Henningsmoen, G. 1948. The Tretaspis Series of the Killatorp Core. Bulletin of Geological Institutions of the University of Uppsala, 32:374432.Google Scholar
Jeppsson, L. 1983. Silurian conodont faunas from Gotland. Fossils and Strata, 15:121144.CrossRefGoogle Scholar
Jeppsson, L. 1989. Latest Silurian conodonts from Klonk, Czechoslovakia. Geologica et Palaeontologica, 23:2137.Google Scholar
Johnston, D. I., and Barnes, C. R. 1999. Early and Middle Ordovician (Arenig) conodonts from St. Paul's Inlet and Martin Point, Cow Head Group, western Newfoundland, Canada. 1. Biostratigraphy and paleoecology. Geologica et Palaeontologica, 33:2170.Google Scholar
Johnston, D. I., and Barnes, C. R. 2000. Early and Middle Ordovician (Arenig) conodonts from St. Paul's Inlet and Martin Point, Cow Head Group, western Newfoundland, Canada. 2. Systematic paleontology. Geologica et Palaeontologica, 34:1187.Google Scholar
Kennedy, D. J., Barnes, C. R., and Uyeno, T. T. 1979. A Middle Ordovician conodont faunule from the Tetagouche Group, Camel Back Mountain, New Brunswick. Canadian Journal of Earth Sciences, 16:540551.CrossRefGoogle Scholar
Khaisnikova, K. B., and Latypov, J. J. 1979. Ordovician-Silurian boundary in South Verkhoyan Range based on corals, p. 9799. In Oradovskaya, M. M. and Sobolevskaya, R. F. (eds.), Guidebook to Field Excursion to the Omulev Mountains. USSR Pacific Science Congress XIV.Google Scholar
Klapper, G., and Murphy, M. A. 1975. Silurian-Lower Devonian conodont sequence in the Roberts Mountains Formation of Central Nevada. University of California Publications in Geological Sciences, 111, 62 p.Google Scholar
Koren, T. N. 1991. Evolutionary crisis of the Ashgill graptolites, p. 157164. In Barnes, C. R. and Williams, S. H. (eds.), Advances in Ordovician Geology. Geological Survey of Canada Paper 90-9.Google Scholar
Koren, T. N., and Sobolevskaya, R. F. 1979. A graptolite zonation of the Ordovician-Silurian boundary deposits of the Omulev Mountains, p. 9192. In Oradovskaya, M. M. and Sobolevskaya, R. F. (eds.), Guidebook to Field Excursion to the Omulev Mountains. USSR Pacific Science Congress XIV.Google Scholar
Koren, T. N., Oradovskaya, M. M., and Sobolevskaya, R. F. 1988. The Ordovician-Silurian boundary beds of the north-east USSR, p. 133138. In Cocks, L. R. M. and Rickards, R. B. (eds.), A Global Analysis of the Ordovician-Silurian Boundary. Bulletin of the British Museum of Natural History (Geology), 43.Google Scholar
Koren, T. N., Oradovskaya, M. M., Sobolevskaya, R. F., and Chungaeva, M. N. (eds.). 1979. Supplement to a guidebook of the field excursion to the Omulev Mountains—Atlas of the paleontological plates. USSR Pacific Science Congress XIV.Google Scholar
Koren, T. N., Oradovskaya, M. M., Pylma, L. J., Sobolevskaya, R. F., and Chungaeva, M. N. 1983. The Ordovician and Silurian boundary in the northeast of the USSR. Academy of Sciences of the USSR, Ministry of Geology of the USSR, Interdepartmental Stratigraphic Committee of the USSR, Transections, Volume 11, 205 p. (In Russian)Google Scholar
Kulkov, N. P., and Vladimirskaya, E. V. 1979. Ordovician-Silurian boundary in Tuva based on brachiopods, p. 93. In Oradovskaya, M. M. and Sobolevskaya, R. F. (eds.), Guidebook to Field Excursion to the Omulev Mountains. USSR Pacific Science Congress XIV.Google Scholar
Lenz, A. C., and McCracken, A. D. 1982. The Ordovician-Silurian boundary, northern Canadian Cordillera: Graptolite and conodont correlation. Canadian Journal of Earth Sciences, 19:13081322.CrossRefGoogle Scholar
Lindström, M. 1971. Lower Ordovician conodonts of Europe. Geological Society of America Memoir, 127:2161.CrossRefGoogle Scholar
Löfgren, A. 1978. Arenigian and Llanvirnian conodonts from Jämtland, northern Sweden. Fossils and Strata, 13, 129 p.Google Scholar
Mashkova, T. V. 1977. New conodonts of the amorphognathoides Zone for the Lower Silurian of Podolia. Paleontologicheskiy Zhurnal, 1977(4):127131. [In Russian; English translation, Scripta Publishing Co., 11(4):513–517]Google Scholar
Männik, P. 1998. Evolution and taxonomy of the Silurian conodont Pterospathodus. Palaeontology, 41:10011050.Google Scholar
McCracken, A. D. 1987. Description and correlation of late Ordovician conodonts from the D. ornatus and P. pacificus graptolite zones, Road River Group, northern Yukon Territory. Canadian Journal of Earth Science, 24:14501464.CrossRefGoogle Scholar
McCracken, A. D. 1989. Protopanderodus (Conodontata) from the Ordovician Road River Group, Northern Yukon Territory, and the evolution of the genus. Geological Survey of Canada Bulletin, 388, 39 p.Google Scholar
McCracken, A. D. 1991a. Taxonomy and biostratigraphy of Llandovery (Silurian) conodonts in the Canadian Cordillera, northern Yukon Territory. Geological Survey of Canada Bulletin, 417:6595.Google Scholar
McCracken, A. D. 1991b. Silurian conodont biostratigraphy of the Canadian Cordillera with a description of new Llandovery species. Geological Survey of Canada Bulletin, 417:97127.Google Scholar
McCracken, A. D., and Barnes, C. R. 1981. Conodont biostratigraphy and paleoecology of the Ellis Bay Formation, Anticosti Island, Quebec, with special reference to Late Ordovician—Early Silurian chronostratigraphy and the system boundary. Geological Survey of Canada Bulletin, 329:51134.Google Scholar
McCracken, A. D., and Nowlan, G. S. 1989. Conodont paleontology and biostratigraphy of Ordovician carbonates and petroliferous carbonates on Southampton, Baffin, and Akpatok Islands in the eastern Canadian Arctic. Canadian Journal of Earth Sciences, 26:18801903.CrossRefGoogle Scholar
McCracken, A. D., Nowlan, G. S., and Barnes, C. R. 1980. Gamachignathus, a new multielement conodont genus from the latest Ordovician, Anticosti Island, Quebec. Current Research Part C, Geological Survey of Canada Paper, 80–01C:103112.Google Scholar
Miller, S. A. 1889. North American geology and palaeontology for the use of amateurs, students, and scientists. Western Methodist Book Concern, Cincinnati, Ohio, 718 p.CrossRefGoogle Scholar
Moskalenko, T. A. 1972. Ordovician conodonts of the Siberian platform and their bearing on multielement taxonomy. Geologica et Palaeontologica, SB 1:4756.Google Scholar
Nicoll, R. S., and Rexroad, C. B. 1969. Stratigraphy and conodont paleontology of the Salamonie Dolomite and Lee Creek Member of the Brassfield Limestone (Silurian) in southeastern Indiana and adjacent Kentucky. Indiana Geological Survey Bulletin, 40, 73 p.Google Scholar
Nikolaev, A. A., Oradovskaya, M. M., and Sapelnikov, V. P. 1977. Biostratigraphical review of the Ordovician and Silurian pentamerids of the north-east USSR. Sverdlovsk, Akademiya Nauk SSSR, Uralskii Nauchnii Tsentr, Trudy Instituta Geologii i Geokhimii, 126:3267. (In Russian)Google Scholar
Nowlan, G. S. 1983. Biostratigraphic, paleogeographic, and tectonic implications of Late Ordovician conodonts from the Grog Brook Group, northwestern New Brunswick. Canadian Journal of Earth Sciences, 20:651671.CrossRefGoogle Scholar
Nowlan, G. S., and Barnes, C. R. 1981. Late Ordovician conodonts from the Vauréal Formation, Anticosti Island, Quebec. Geological Survey of Canada Bulletin, 329:150.Google Scholar
Nowlan, G. S., McCracken, A. D., and Chatterton, B. D. E. 1988. Conodonts from Ordovician-Silurian boundary strata, Whittaker Formation, Mackenzie Mountains, Northwest Territories. Geological Survey of Canada Bulletin, 373, 99 p.Google Scholar
Nowlan, G. S., McCracken, A. D., and McLeod, M. J. 1997. Tectonic and paleogeographic significance of late Ordovician conodonts in the Canadian Appalachians. Canadian Journal of Earth Sciences, 34:15211537.CrossRefGoogle Scholar
Obut, A. M., and Sennikov, N. V. 1979. Graptolite assemblages from the Ordovician-Silurian boundary deposits of Pacific palaeobiogeographic province in Siberia and neighbouring regions, p. 94. In Oradovskaya, M. M. and Sobolevskaya, R. F. (eds.), Guidebook to Field Excursion to the Omulev Mountains. USSR Pacific Science Congress XIV.Google Scholar
Obut, A. M., Sobolevskaya, R. F., and Nikolaev, A. A. 1967. Graptolites and the stratigraphy of the lower Silurian of marginal uplifts of the Kolyma Massif, the north-east USSR Moscow, 162 p. (In Russian)Google Scholar
Oradovskaya, M. M. 1963. The Ordovician deposits of the Chersky Ridge. In Materialy po geologii i poleznym iskopaemym Severo-Vostoka SSSR, 16:140162. (In Russian)Google Scholar
Oradovskaya, M. M. 1988a. The Ordovician System in the north-east and far east of the USSR, p. 85115. In Ross, R. J. Jr. and Talent, J. (eds.), The Ordovician System in most of Russian Asia. International Union of Geological Sciences Publication, No. 26.Google Scholar
Oradovskaya, M. M. 1988b. The Ordovician-Silurian biostratigraphy and facies of North-Eastern part of the USSR. Publication of Geological Ministry of the USSR North-Eastern Geological Department. Moscow“Nedra”.Google Scholar
Oradovskaya, M. M., and Sobolevskaya, R. F. (eds.). 1979. Guidebook to Field Excursion to the Omulev Mountains. USSR Pacific Science Congress XIV, 103 p.Google Scholar
Orchard, M. J. 1980. Upper Ordovician conodonts from England and Wales. Geologica et Palaeontologica, 14:944.Google Scholar
Pander, C. H. 1856. Monographie der fossilen Fische des Silurischen Systems der Russisch-Baltischen Gouvernements. Kaiserlichen Akademie der Wissenschaften St. Petersburg, 91 p.Google Scholar
Pohler, S. M. L. 1994. Conodont biofacies of Lower to lower Middle Ordovician megaconglomerates, Cow Head Group, western Newfoundland. Geological Survey of Canada Bulletin, 459, 71 p.Google Scholar
Pollock, C. A., Rexroad, C. B., and Nicoll, R. S. 1970. Lower Silurian conodonts from Northern Michigan and Ontario. Journal of Paleontology, 44:743764.Google Scholar
Preobrazhensky, B. V. 1966. Biostratigraphic substantiation of the Ordovician-Silurian in the north-east USSR based on tabulate corals. Avtoreferat kandidata dissertatsiya, Novosibirsk, 20 p. (In Russian)Google Scholar
Rasmussen, J. A. 1998. A reinterpretation of the conodont Atlantic realm in the late Early Ordovician (early Llanvirn). Acta Palaeontologica Polonica, 58:6777.Google Scholar
Rhodes, F. H. T. 1953. Some British Lower Paleozoic conodont faunas. Philosophical Transactions of the Royal Society of London, Series B, 647:261334.Google Scholar
Rhodes, F. H. T. 1955. The conodont fauna of the Keisley Limestone. Quarterly Journal of the Geological Society of London, 111:117142.CrossRefGoogle Scholar
Rexroad, C. B. 1967. Stratigraphy and conodont paleontology of the Brassfield (Silurian) in the Cincinnati Arch area. Indiana Geological Survey, Bulletin 36, 64 p.Google Scholar
Sennikov, N. V. 1998. Manifestations of global Ordovician-Silurian biotic crisis in central Siberia graptolite communities. Geologiya i geofizika, 39:557567. (In Russian)Google Scholar
Serpagli, T. J. M. 1967. I conodonti del'Ordoviciano Superiore (Ashgilliano) delle Alpi Carniche. Bollettino della Società Paleontologica Italiana, 6:30111.Google Scholar
Sheshegova, L. I. 1979. Upper Ordovician and Lower Silurian acritarchs of the Siberian platform and their stratigraphic significance, p. 100. In Oradovskaya, M. M. and Sobolevskaya, R. F. (eds.), Guidebook to Field Excursion to the Omulev Mountains. USSR Pacific Science Congress XIV.Google Scholar
Simpson, A. J., and Talent, J. A. 1995. Silurian conodonts from the headwaters of the Indi (upper Murray) and Buchan rivers, southeastern Australia, and their implications, p. 79215. In Mawson, R. and Talent, J. A. (eds.), Contribution to the First Australian Conodont Symposium (AUSCOS 1), Courier Forschungsinstitut Senckenberg, 182.Google Scholar
Sobolevskaya, R. F. 1970. Middle and Upper Ordovician biostratigraphy of the Kolyma uplifts based on graptolites. Avtoreferat kandidata dissertatsiya, Moskva, 23 p. (In Russian)Google Scholar
Sobolevskaya, R. F. 1974. New Ashgillian graptolites from the middle Kolyma River basin, p. 6371. In Obut, A. M. (ed.), Graptolity SSSR, Trudy I Vses. Kollokviuma, Novosibirsk. (In Russian)Google Scholar
Stauffer, C. R. 1930. Conodonts from the Decorah Shale. Journal of Paleontology, 4:121128.Google Scholar
Stauffer, C. R. 1935. The conodont fauna of the Decorah Shale (Ordovician). Journal of Paleontology, 9:596620.Google Scholar
Stauffer, C. R. 1940. Conodonts from the Devonian and associated clays of Minnesota. Journal of Paleontology, 14:417435.Google Scholar
Sweet, W. C. 1979. Late Ordovician conodonts and biostratigraphy of the western Midcontinent Province, p. 4585. In Sandberg, C. A. and Clark, D. L. (eds.), Conodont Biostratigraphy of the Great Basin and Rocky Mountains. Brigham Young University Geology Studies, 26 (3).Google Scholar
Sweet, W. C. 1988. The Conodonta: Morphology, taxonomy, paleoecology, and evolutionary history of a long-exinct animal phylum. Oxford Monographs on Geology and Geophysics No. 10, 212 p.Google Scholar
Sweet, W. C., and Schönlaub, H. P. 1975. Conodonts of the Genus Oulodus Branson and Mehl, 1933. Geologica et Palaeontologica, 9:4159.Google Scholar
Uyeno, T. T. 1981. Stratigraphy and conodonts of Upper Silurian and Lower Devonian rocks in the environs of the Boothia Uplift, Canadian Arctic Archipelago, Pt. II, Systematic study of conodonts. Geological Survey of Canada Bulletin, 292:3975.Google Scholar
Uyeno, T. T., and Barnes, C. R. 1983. Conodonts of the Jupiter and Chicotte Formations (Lower Silurian), Anticosti Island, Québec. Geological Survey of Canada, Bulletin 355, 49 p.CrossRefGoogle Scholar
Uyeno, T. T. 1990. Biostratigraphy and conodont faunas of Upper Ordovician through Middle Devonian rocks, eastern Arctic Archipelago. Geological Survey of Canada Bulletin, 401, 211 p.Google Scholar
Uyeno, T. T., and Barnes, C. R. 1983. Conodonts of the Jupiter and Chicotte Formations (Lower Silurian), Anticosti Island, Québec. Geological Survey of Canada, Bulletin 355, 49 p.CrossRefGoogle Scholar
Viira, V. 1974. Ordovician conodonts of the Baltic Region. Tallin, USSR, 142 p. (In Russian)Google Scholar
Walliser, O. H. 1957. Conodonten aus dem oberen Gotlandium Deutschlands und der Karnischen Alpen. Notizblatt des Hessischen Landesamtes für Bodenforschung zu Wiesbaden, 85:2852.Google Scholar
Walliser, O. H. 1964. Conodonten des Silurs. Abhandlungen des Hessische Landesamtes für Bodenforschung, 41, 106 p.Google Scholar
Zaslavskaya, N. M. 1979. Upper Ordovician and Early Silurian chitinozoan assemblages from the Pacific paleobiogeographic province in Siberia and neighbouring regions, p. 90. In Oradovskaya, M. M. and Sobolevskaya, R. F. (eds.), Guidebook to Field Excursion to the Omulev Mountains. USSR Pacific Science Congress XIV.Google Scholar
Zhang, S., and Barnes, C. R. 2002. A new Llandovery (Early Silurian) conodont biozonation and conodonts from the Becscie, Merrimack and Gun River Formations, Anticosti Island, Québec. Paleontological Society Memoir, 57 (Journal of Paleontology, Volume 76, no. 2, supp.), 46p.Google Scholar