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    Anderson, Tavis K. and Sukhdeo, Michael V. K. 2016. A Century of Parasitology.

    Ash, Anirban de Chambrier, Alain Shimazu, Takeshi Ermolenko, Alexey and Scholz, Tomáš 2015. An annotated list of the species of Gangesia Woodland, 1924 (Cestoda: Proteocephalidea), parasites of catfishes in Asia, with new synonyms and a key to their identification. Systematic Parasitology, Vol. 91, Issue. 1, p. 13.

    de Chambrier, Alain Waeschenbach, Andrea Fisseha, Makda Scholz, Tomas and Mariaux, Jean 2015. A large 28S rDNA-based phylogeny confirms the limitations of established morphological characters for classification of proteocephalidean tapeworms (Platyhelminthes, Cestoda). ZooKeys, Vol. 500, p. 25.

    Khalil, L. F. 1960. On a New Genus, Sandonella, for Proteocephalus sandoni Lynsdale, 1960, (Proteocephalidae) and the Erection of a New Subfamily, Sandonellinae. Journal of Helminthology, Vol. 34, Issue. 1-2, p. 47.

    Lynsdale, J. A. 1959. On a New Species of Proteocephalus from Brazil. Journal of Helminthology, Vol. 33, Issue. 2-3, p. 145.

    Meggitt, F. J. 1930. Report on a Collection of Cestoda, mainly from Egypt. Parasitology, Vol. 22, Issue. 03, p. 338.

    Meggitt, F. J. 1928. Report on a Collection of Cestoda, mainly from Egypt. Part III. Cyclophyllidea (Conclusion): Tetraphyllidea. Parasitology, Vol. 20, Issue. 03, p. 315.

    Southwell, T. 1928. Cestodes of the OrderPseudophyllideaRecorded from India and Ceylon. Annals of Tropical Medicine & Parasitology, Vol. 22, Issue. 4, p. 419.

    Meggitt, F. J. 1927. Remarks on the Cestode FamiliesMonticellidaeandIchthyotaeniidae. Annals of Tropical Medicine & Parasitology, Vol. 21, Issue. 1, p. 69.

    Southwell, T. 1925. On the GenusTetracampos, Wedl, 1861. Annals of Tropical Medicine & Parasitology, Vol. 19, Issue. 1, p. 71.

    Woodland, W. N. F. 1925. TetracamposWedl1861 as a Genus of the Bothriocephalidae. Annals of Tropical Medicine & Parasitology, Vol. 19, Issue. 2, p. 185.

    Woodland, W. N. F. 1925. On Three New Proteocephalids (Cestoda) and a Revision of the Genera of the Family. Parasitology, Vol. 17, Issue. 04, p. 370.


On a New Bothriocephalus and a New Genus of Proteocephalidae from Indian Fresh-water Fishes

  • W. N. F. Woodland (a1)
  • DOI:
  • Published online: 01 April 2009

1. A new species of Bothriocephalus—B. pycnomerus—is described from the intestine of Ophiocephalus marulius Ham. Buch., from the Ganges and Jumna at Allahabad, India.

2. B. pycnomerus closely resembles B. histiophorus (Shipley 1901; from the marine sword-fish Histiophorus) in structure but differs chiefly in the possession of an armed scolex and its crowded indistinct segmentation.

3. Lühe's definition of the genus Bothriocephalus must be amended to include forms with armed scolices and forms in which the ventral uterine apertures are a-median and irregularly alternate.

4. Two new species of Proteocephalidae are described which possess armed muscular rostella, and for these a new genus Gangesia is created and defined. The definitions of the Order Tetraphyllidea (Lühe 1910) and of the Family Proteocephalidae (La Rue 1914) must be amended to include forms possessing muscular rostella, armed or unarmed.

5. These two new species—Gangesia wallago and G. macrones—were found in the intestines of Wallago attu Bleek and Macrones seenghala Sykes (both Siluroids) respectively, from the Ganges and Jumna at Allahabad.

6. Southwell's “Ophryocotyle bengalensis,” from Ophiocephalus striatus, Labeo rohita and Wallago attu, is probably identical with Gangesia wallago.

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  • ISSN: 0031-1820
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