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Biostratigraphy, paleoecology, and taxonomy of Devonian (Emsian and Famennian) crinoids from southeastern Morocco

Published online by Cambridge University Press:  20 May 2016

G. D. Webster ,
R. Thomas Becker  and
Christopher G. Maples
G. D. Webster
Affiliation:
Department of Geology, Washington State University, Pullman, Washington 99163-2812 USA,
R. Thomas Becker
Affiliation:
Geologisch-Paläontologisches Institut, Westfälische Wilhelms-Universität, Corrensstr. 24, D-48149 Münster, Germany,
Christopher G. Maples
Affiliation:
Desert Research Institute, 2215 Raggio Parkway, Reno, Nevada 89512,
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Abstract

An early late Emsian codiacrinid and Frasnian specimens of an indeterminate species of Parapisocrinus, an undesignated new genus of a catillocrinid, and two amabilicrinids are described from the eastern part of the Anti-Atlas Mountains of Morocco. These are the first articulated crinoid specimens reported from the Famennian of Morocco. Specimens occur within ammonoid-rich dysoxic shales, marls, and thin limestones at Ouidane Chebbi, Jebel Mrakib, Lambidia, and Rich Bou Kourazia. Parapisocrinus, the codiacrinid, and the catillocrinid are loose cups that are associated with other megafossils and may have been transported within the pelagic depositional settings. The amabilicrinids occur in three small unitaxial lenses, two of which have associated logs. The problems with and possibility of a pseudoplanktic lifestyle for stemmed crinoids are discussed.

These new collections expand the known stratigraphic and geographic distribution of the pisocrinid and amabilicrinids and the early evolutionary history of the catillocrinids. New taxa introduced are the amabilicrinids Mrakibocrinus bockwinkeli, Moroccocrinus ebbighauseni, and the codiacrinid Elicrinus? weyeri.

Type
Research Article
Information
Journal of Paleontology , Volume 79 , Issue 6 , November 2005 , pp. 1052 - 1071
Copyright
Copyright © The Paleontological Society 

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References

Aboussalam, Z. S., and Becker, R. T. 2002. The Base of the hermanni Zone as the Base of an Upper Givetian Substage. Document of the International Subcommission on Devonian Stratigraphy, Annual Meeting, Toulouse 2002, 10 p.Google Scholar
Angelin, N. P. 1878. Iconographia Crinoideorum. In Stratis Sueciae Siluricis Fossilium. Samson and Wallin, Holmiae, 62 p., 29 pls.Google Scholar
Anstedt, D. T. 1838. On a new genus of fossil multilocular shells, found in the slate-rocks of Cornwall. Transactions of the Cambridge Philosophical Society, 6:415–422.Google Scholar
Arendt, Yu. A., and Hecker, R. T. 1964. Klass Crinoidea. Morski lilii. Sistematitcheckaia tchast [Class Crinoidea. Crinoids. Systematic part]. In Orlov, Yu. A. (ed.), Osnovi Paleontologii, Iglokozhi, Gemikhordovye, Pogonofory, i Shchetinkochelyustnye [Fundamentals of paleontology, Echinodermata, Hemichordata, Pogonophora, and Chaetognatha]. Moscow, Izdatelstvo Nedra, p. 76–105, 214–231, pls. 8–16.Google Scholar
Ausich, W. I. 1977. The functional morphology and evolution of Pisocrinus (Crinoidea: Silurian). Journal of Paleontology, 51:672–686, 2 pls.Google Scholar
Ausich, W. I. 1980. A model for niche differentiation in Lower Mississippian crinoid communities. Journal of Paleontology, 54:273–288.Google Scholar
Barrande, J. 1865. Systême Silurien du centre de la Bohême. I. Volume II. Classe des Mollusques, orde des Céphalopodes. Praha, Paris, 107 pls.Google Scholar
Barrande, J. 1881. Système Silurien du centre de la Bohéme, première partie, recherches paléontologique. Volume 6. Classe des mollusques, ordre des acèphalès. 342 p., 48 pls.Google Scholar
Bartels, C., Briggs, D. E. G., and Brassel, G. 1998. The Fossils of the Hunsrück Slate: Marine Life in the Devonian. Cambridge University Press, Cambridge, 309 p.Google Scholar
Bather, F. A. 1890. The classification of the Inadunata Fistulata. Annals and Magazine of Natural History, ser. 6, 5:310–334, 373–388, 485–486, pls. 14, 15.Google Scholar
Bather, F. A. 1893. The Crinoidea of Gotland, Pt. 1, The Crinoidea Inadunata. Kongliga Svenska Vetenskaps-Akademiens Handlingar, 25(2):200 p., 10 pls.Google Scholar
Bather, F. A. 1899. A phylogenetic classification of the Pelmatozoa. British Association for the Advancement of Science (1898):916–923.Google Scholar
Becker, R. T. 1992a. Zur Kenntnis von Hemberg-Stufe und Annulata-Schiefer im Nordsauerland (Oberdevon, Rheinisches Schiefergebirge, GK 4611 Hohenlimburg). Berliner geowissenschaftliche Abhandlungen, E3:3–41.Google Scholar
Becker, R. T. 1992b. Regional developments of the global hypoxic Annulata Event (middle Famennian). Fifth International Conference on Global Bioevents, Göttingen, 16–19 February 1992, Abstract, p. 13–14.Google Scholar
Becker, R. T. 1993. Stratigraphische Gliederung und Ammonoideen-Faunen im Nehdenium (Oberdevon) von Europa und Nord-Afrika. Courier Forschungsinstitut Senckenberg, 155, 405 p.Google Scholar
Becker, R. T. 1995. Taxonomy and Evolution of Late Famennian Tornocerataceae (Ammonoidea). Berliner geowissenschaftliche Abhandlungen, (Gundolf-Ernst-Festschrift), E16:607–643.Google Scholar
Becker, R. T. 2000. Palaeobiogeographic relationships and diversity of Upper Devonian ammonoids from western Australia, p. 385–401. In Mawson, R., Talent, J., and Long, J. A. (eds.), Mid-Palaeozoic Biota and Biogeography. Records of the Western Australian Museum supplement 58, 420 p.Google Scholar
Becker, R. T., and House, M. R. 1994. International Devonian goniatite zonation, Emsian to Givetian, with new records from Morocco. Courier Forschungsinstitut Senckenberg, Willi Ziegler-Gestschrift II, 169:79–135.Google Scholar
Becker, R. T., and House, M. R. 2000a. Devonian ammonoid zones and their correlation with established series and stage boundaries. Courier Forschungsinstitut Senckenberg, 220:113–151.Google Scholar
Becker, R. T., and House, M. R. 2000b. The Famennian ammonoid succession at Bou Tchrafine (Anti-Atlas, Southern Morocco). Notes et Mémoires du Service géologique Maroc, 399:37–42.Google Scholar
Becker, R. T., Bockwinkel, J., Ebbighausen, V., and House, M. R. 1999. Jebel Mrakib, Anti-Atlas (Morocco), a potential Upper Famennian substage boundary stratotype section, p. 91–107. In El Hassani, A. and Tahiri, A. (eds.), SDS—IGCP 421 Morocco Meeting, 23 April–1 May 1999, Excursion Guidebook, Pt. 1, Tafilalt and Maider (eastern Anti-Atlas), Rabat.Google Scholar
Becker, R. T., Bockwinkel, J., Ebbighausen, V., and House, M. R. 2000a. Jebel Mrakib, Anti-Atlas (Morocco), a potential Upper Famennian substage boundary stratotype section. Notes et Mémoires du Service géologique Maroc, 399:75–86.Google Scholar
Becker, R. T., House, M. R., Menner, V. V., and Ovnatanova, N. S. 2000b. Revision of ammonoid biostratigraphy in the Frasnian (Upper Devonian) of the southern Timan (Northeast Russian Platform). Acta Geologica Polonica, 50(1):67–97.Google Scholar
Becker, R. T., House, M. R., Bockwinkel, J., Ebbighausen, V., and Aboussalam, Z. S. 2002. Famennian ammonoid zones of the eastern Anti-Atlas (southern Morocco). Münstersche Forschungen zur Geologie und Paläontologie, 93:159–205.Google Scholar
Belka, Z., Kaufmann, B., and Bultynck, P. 1997. Conodont-based quantitative biostratigraphy for the Eifelian of the eastern Anti-Atlas, Morocco. Geological Society of America Bulletin, 109:643–651.Google Scholar
Belka, Z., Klug, C., Kaufmann, B., Korn, D., Döring, S., Feist, R., and Wendt, J. 1999. Devonian conodont and ammonoid succession of the eastern Tafilalt (Ouidane Chebbi section), Anti-Atlas, Morocco. Acta Geologica Polonica, 49:1–23.Google Scholar
Bogoslovskij, B. I. 1975. A new clymeniid family. Paleontological Journal, 9:304–310.Google Scholar
Bouska, J. 1956. Pisocrinidae Angelin ceského Siluru a Devonu (Crinoidea) [Pisocrinidae Angelin from the Silurian and Devonian of Bohemia (Crinoidea)]. Rozpravy Ustredního Ustavu Geologického, 20:1–138, 6 pls. (In Czech; Russian summary, p. 55–96; English summary p. 97–138) Google Scholar
Brett, C. E. 1981. Terminology and functional morphology of attachment structures in pelmatozoan echinoderms. Lethaia, 14:343–370.CrossRefGoogle Scholar
Brower, J. C. 1973. Crinoids from the Girardeau Limestone (Ordovician). Palaeontographia Americana, 7(46):263–499, pls. 59–79.Google Scholar
Brower, J. C. 1974. Ontogeny of camerate crinoids. University of Kansas Paleontological Contributions Paper, 72, 53 p.Google Scholar
Bultynck, P., and Walliser, O. H. 2000. Emsian to middle Frasnian sections in the northern Tafilalt. Notes et Mémoirs du Service géologique Maroc, 399:11–20.Google Scholar
Carpenter, P. H., and Etheridge, R. Jr. 1881. Contributions to the study of the British Paleozoic crinoids. No. 1. On Allagecrinus, the representative of the Carboniferous limestone series. Annals and Magazine of Natural History, ser. 5, 7:281–298, pls. 15, 16.Google Scholar
Czarnocki, J. 1989. Klimenie Gór Swietokrzyskich. Prace Panstwowego Instytutu Geologicznego, 127, 91 p.Google Scholar
Davies, R. A., Mapes, R. H., and Klofak, S. M. 1999. Epizoa on externally shelled cephalopods, p. 32–51. In Rozanov, A. Y. and Shevyrev, A. A. (eds.), Fossil Cephalopods: Recent Advances in Their Study. Russian Academy of Science, Paleontological Institute, Moscow.Google Scholar
de Koninck, L. G. 1841–1844. Description des animaux fossiles qui se trouvent dans le terrain carbonifère de Belgique. H. Dessain, Leige, 650 p., 55 pls.Google Scholar
de Koninck, L. G. 1858. Sur quelques Crinoides paleozoiques nouveaux de l'Angleterre et de l'Ecosse. Bulletin de la Academie Royal des Sciences, des Lettres et des Beaux-Arts de Belgique, ser. 2, 4:93–108, pl. 2.Google Scholar
Destombes, J., and Hollard, H. (eds.). 1986. Carte géologique du Maroc. Tafilalt–Taouz. Echeele 1/200,000. Ministère de L'Energie et des Mines, Direction de la Géologie, Rabat.Google Scholar
Destombes, J., and Hollard, H. (eds.). 1988. Carte géologique du Maroc. Todrha-Mader. Echelle 1/200,000. Ministère de LEnergie et des Mines, Direction de la Géologie, Rabat.Google Scholar
Ebbighausen, V., Becker, R. T., and Bockwinkel, J. 2002. Morpho-metric analyses and taxonomy of oxyconic Goniatites (Paratornoceratinae n. subfam.) from the early Famennian of the Tafilalt (Anti-Atlas, Morocco). Abhandlungen der Geologisches Bundesanstalt, 57:167–180.Google Scholar
El Hassani, A., and Tahiri, A. (eds.). 2000. Moroccan meeting of the Subcommission on Devonian Stratigraphy, (SDS)-IGCP 421, 24 April–1 May 1999. Notes et Mémoires du Service Géologique, 399, 128 p.Google Scholar
Frech, F. 1891. Die devonischen Aviculiden Deutschlands, ein Beitrag zur Systematik und Stammesgeschichte der Zweischaler. Abhandlungen zur Geologische Spezialkarte von Preussen, 9, 253 p., 18 pls.Google Scholar
Frech, F. 1902. Über devonische Ammoneen. Beiträge zur Paläontologie Österreich-Ungarns und des Orients, 14:27–112.Google Scholar
Goldring, W. 1923. The Devonian crinoids of the state of New York. New York State Museum Memoir, 16, 670 p., 60 pls.Google Scholar
Gürich, G. 1900. Nachträge zum Palaeozoikum des Polnischen Mittel-gebirges. Neues Jahrbuch für Mineralogie, Geologie und Paläontologie, Beilage-Band, 13:331–388.Google Scholar
Hall, J. 1843. Geology of New York, Pt. IV, comprising the survey of the fourth geological district. Natural History of New York, 4, 683 p., 19 pls.Google Scholar
Haude, R. 1980. Constructional morphology of the stems of Pentacrinitidae, and mode of life of Seirocrinus , p. 17–23. In Jangoux, M. (ed.), Echinoderms: Present and Past. A. A. Balkema, Rotterdam.Google Scholar
House, M. R. 1985. Correlation of mid-Paleozoic ammonoid evolutionary events with global sedimentary perturbations. Nature, 313:17–22.Google Scholar
House, M. R., and Price, J. D. 1985. New Late Devonian genera and species of tornoceratid goniatites. Palaeontology, 28:159–188.Google Scholar
Hyatt, A. 1884. Genera of fossil cephalopods. Proceedings of the Boston Society of Natural History, 22:253–338.Google Scholar
Jaekel, O. 1898. Über einen neuen devonischen Pentacrinoiden. Zeitschridft der Deutschen Geologischen Gesellschaft, Verhandlungen Jahrgang, 1898:5028–32.Google Scholar
Jell, P. A., and Jell, J. S. 1999. Crinoids, a blastoid and a cyclocystoid from the Upper Devonian reef complex of the Canning Basin, western Australia. Memoirs of the Queensland Museum, 43(1):201–236.Google Scholar
Jell, P. A., Jell, J. S., Johnson, B. D., Mawson, R., and Talent, J. A. 1988. Crinoids from Devonian limestones of eastern Australia. Memoirs of the Queensland Museum, 25:355–402.Google Scholar
Kaufmann, B. 1998. Facies, stratigraphy and diagenesis of Middle Devonian reef- and mud-mounds in the Mader (eastern Anti-Atlas, Morocco). Acta Geologica Polonica, 48:43–106.Google Scholar
Keupp, H. 2000. Ammoniten. Paläobiologische Erfolgsspiralen. Jan Thorbecke Verlag, Stuttgart, 164 p.Google Scholar
Klug, C. 2002. Quantitative stratigraphy and taxonomy of late Emsian and Eifelian ammonoids of the eastern Anti-Atlas Morocco. Courier Forschungsinstitut Senckenberg, 238, 109 p.Google Scholar
Korn, D. 1999. Famennian ammonoid stratigraphy of the Mader and Tafilalt (eastern Anti-Atlas, Morocco). Abhandlungen der Geologischen Bundesanstalt, 54:147–179.Google Scholar
Korn, D. 2002. Die Ammonoideen-Fauna der Platyclymenia annulata-Zone vom Kattensiepen (Oberdevon, Rheinishches Schiefergebirge). Senckenbergiana Lethaea, 82(2):557–608.Google Scholar
Korn, D., and Klug, C. 2002. Ammoneae Devonicae. Fossilis Catalogus Animalia, 138, 375 p.Google Scholar
Korn, D., and Luppold, F. W. 1987. Nach Clymenien und Conodonten gegliederte Profile des oberen Famennium im Rheinischen Schiefergebirge. Courier Forschungsinstitut Senckenberg, 92:199–223.Google Scholar
Korn, D., and Price, J. 1987. Taxonomy and phylogeny of the Kosmoclymeniinae subfam. nov. (Cephalopoda, Ammonoidea, Clymeniida). Courier Forschungsinstitut Senckenberg, 92:5–75.Google Scholar
Korn, D., and Ziegler, W. 2003. The ammonoid and conodont zonation at Enkenberg (Famennian, Late Devonian; Rhenish Mountains). Senckenbergiana Lethaea, 82(2):453–462.Google Scholar
Korn, D., Klug, C., and Reisdorf, A. 2000. Middle Famennian ammonoid stratigraphy in the Amessoui Syncline (Late Devonian; eastern Anti-Atlas, Morocco). Travaux de lInstitut Scientifique, Rabat, Série Géologie and Géographie Physique, 20:69–77.Google Scholar
Kutscher, R. 1976. Beiträge zur Sedimentation und Fossilführung des Hunsrückschiefers. 42. Die Crinoideen-Arten des Hunsrückschiefers. Geologisches Jahrbuch, 104:9–24.Google Scholar
Lane, N. G. 1963. Meristic variation in the dorsal cup of monobathrid camerate crinoids. Journal of Paleontology, 37:917–930.Google Scholar
Lane, N. G., Maples, C. G., and Waters, J. A. 2001. Revision of Late Devonian (Famennian) and some Early Carboniferous (Tournaisian) crinoids and blastoids from the type Devonian area of North Devon. Palaeontology, 44(6): 1043–1080.Google Scholar
Lane, N. G., Waters, J. A., and Maples, C. G. 1997. Echinoderm faunas of the Hongguleleng Formation, Late Devonian (Famennian), Xinjiang-Uygur Autonomous Region, People's Republic of China. Journal of Paleontology, 71 Memoir (Supplement to no. 2), 43 p.Google Scholar
Lange, W. 1929. Zur Kenntnis des Oberdevons am Enkeberg und bei Balve (Sauerland). Abhandlungen der Preußischen Geologischen Landesanstalt, Neue Folge, 119, 32 p.Google Scholar
Le Maitre, D. 1958. Contribution a l'etude des faunes devoniennes d'Afrique du Nord, I Echinodermes. Publications du Service de la Carte géologique de L'Algerie, Bulletin, 20:114–151, 3 pls.Google Scholar
Le Menn, J. 1992. Evolution du genre Haplocrinites: Crinoide Inadunata atypique du Devonien moyen, p. 105–112. In Gayet, M. (ed.), Premier Congrés National de Paléontologie, Paris 17–19 Mai 1990, Geobios Special Mémoire, 14.Google Scholar
Le Menn, J., and Regnault, S. 1993. Découverte de microcrinoïdes dans le Dévonien inférieur de Maroc. Compte Rendus Academie des Sciences Paris, ser. 2, 316:251–256.Google Scholar
Macurda, D. B. Jr. 1968. Ontogeny of the crinoid Eucalyptocrinites . Journal of Paleontology, 32,suppl.5(2 of 2):99–118.Google Scholar
McIntosh, G. C. 2001. Devonian cladid crinoids: Families Glossocrinidae Goldring, 1923, and Rutkowskicrinidae new family. Journal of Paleontology, 75:783–807.Google Scholar
Messing, C. G., Rosesmyth, M. C., Mailer, M. S. T., and Miller, J. E. 1988. Relocation movement in a stalked crinoid (Echinodermata). Bulletin of Marine Science, 42(3):480–487.Google Scholar
Meyer, D. L. 1965. Plate growth in some platycrinid crinoids. Journal of Paleontology, 39:1207–1209.Google Scholar
Meyer-Berthaud, B., Wendt, J., and Galthier, J. 1997. First record of a large Callixylon trunk from the late Devonian of Gondwana. Geological Magazine, 134(6):847–853.Google Scholar
Miller, J. S. 1821. A natural history of the crinoidea, or lily-shaped animals; with observations on the genera, Asteria, Eurayale, Comatula and Marsupites . Bryan & Co., Bristol, England, 150 p.Google Scholar
Moore, R. C., and Laudon, L. R. 1943. Evolution and classification of Paleozoic crinoids. Geological Society of America Special Paper, 46, 151 p., 14 pls.Google Scholar
Moore, R. C., and Teichert, C. (eds.). 1978. Treatise on Invertebrate Paleontology. Pt. T. Echinodermata 2, Crinoidea. 3 vols. Geological Society of America and University of Kansas, Lawrence, 1026 p.Google Scholar
Mu, A. T. 1954. On the occurrence of Pisocrinus in China. Acta Palaeontologica Sinica, 2(3):323–332, 1 pl.Google Scholar
Münster, G., Zu, Graf. 1832. Ueber die Planuliten und Goniatiten im Uebergangs-Kalk des Fichtelgebirges. Birner, Bayreuth, 38 p.Google Scholar
Münster, G., Zu, Graf. 1840. Die Versteinerungen des Uebergangskalkes mit Clymenien und Orthoceratiten von Oberfranken. Beiträge zur Petrefactenkunde, 3:33–121.Google Scholar
Pfeiffer, H. 1954. Der Bohlen bei Saalfeld/Thür. Geologie, Beihefte, 11, 105 p.Google Scholar
Prokop, R. J. 1973. Elicrinus n. gen. from the Lower Devonian of Bohemia (Crinoidea). Véstnik Ustrédniho Ustavu Geologickeho, 48(4): 221–224, 2 pls.Google Scholar
Prokop, R. J., and Petr, V. 1991. New microcriniods from the Trebotov Limestone of the Bohemian Lower Devonian (Dalejan = upper Emsian). Casopis pro Mineralogii a Geologii, 36(2–3):131–136, 4 pls.Google Scholar
Prokop, R. J., and Petr, V. 1998. Isolated skeletal ossicles of the crinoid family Catillocrinidae Wachsmuth et Springer, 1866 in the Devonian of Bohemia. Casopis Národního Nuzea Rada Prírodovedná, 167(1–4):52–53.Google Scholar
Richter, R., and Unger, F. 1856. Beitrag zur Paläontologie des Thuringer Waldes (1855). Denckschriften der Akademie der Wissenschaften Wien, Mathematisch-naturwissenschaftliche Klasse, 11:87–138, 3 pls.Google Scholar
Rozhnov, S. V. 1981. Morskie lilii nadsemeistva Pisocrinacea [The crinoid superfamily Pisocrinacea]. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Institut, 192, 127 p., 24 pls.Google Scholar
Rzehak, A. 1910. Der Brünner Clymenienkalk. Zeitschrift des mährischen Landesmuseums, 10(2):149–216.Google Scholar
Sandberger, G. 1855. Clymenia subnautilina (nova species), die erste und bis jetzt einzige Art aus Nassau. Jahrbücher des Vereins für Naturkunde im Herzogthum Nassau, 10:127–136.Google Scholar
Sartenaer, P. 1998. The presence in Morocco of the late Famennian genus Hadyrhynchia Havlícek, 1979 (rhynchonellid, brachiopod). Bulletin de lInstitut Royal des Sciences Naturelles de Belgique, Sciences de la Terre, 68:115–120.Google Scholar
Sartenaer, P. 1999. Tetragonorhynchus, new late Famennian rhynchonellid genus from Ma'der, southern Morocco, and Tetragonorhynchidae n. fam. Bulletin de lInstitut Royal des Sciences Naturelles de Belgique, Sciences de la Terre, 70:75–88.Google Scholar
Sartenaer, P. 2000. Phacoiderhynchus, a new middle Famennian rhynchonellid genus from the Anti-Atlas, Morocco, and Phacoiderhynchidae n. fam. Bulletin de lInstitut Royal des Sciences Naturelles de Belgique, Sciences de la Terre, 68:115–120.Google Scholar
Schindewolf, O. H. 1923. Beiträge zur Kenntnis des Palaeozoikums in Oberfranken, Ostthüringen und dem Sächsischen Vogtlande. I. Stratigraphie und Ammoneenfauna des Oberdevons von Hof a. S. Neues Jahrbuch für Geologie und Paläontologie, Beilage-Band, 49:250–357, 393–509.Google Scholar
Schindewolf, O. H. 1933. Vergleichende Morphologie und Phylogenie der Anfangstammern tetrabranchiater Cephalopoden. Abhandlungen der Preußischen Geologischen Landesanstalt, Neue Folge, 148, 115 p.Google Scholar
Schindewolf, O. H. 1949. Zur Nomenklatur der Clymenien (Ceohalop., Ammon.). Neues Jahrbuch für Mineralogie, Geologie und Paläontologie, Monatshefte, (B) 1949:197–209.Google Scholar
Schindler, E. 1990. Die Kellwasser-Krise (hohe-Frasne-Stufe, Ober-Devon). Göttinger Arbeiten zur Geologie und Palaeontologie, 46, 115 p.Google Scholar
Schmidt, H. 1924. Zwei Cephalopodenfaunen an der Devon-Carbongrenze im Sauerland. Jahrbuch der Preußischen Geologischen Landesanstalt, (für 1923), 44:98–171.Google Scholar
Schuchert, C., and Cooper, G. A. 1931. Synopsis of the brachiopod genera of the suborders Orthoidea and Pentameroidea, with notes on the Telotremata. American Journal of Science, 5/22:241–251.Google Scholar
Schultz, L. 1866. Monographie der Echinodermen des Eifler Kalkes. Denkschriften der Kaiserlich Akademie der Wissenschaften Mathematisch-Naturwissenschaftlichen Classe, Wien, 26(2):113–230, pls. 1–13.Google Scholar
Scotese, C. R., and McKerrow, W. S. 1990. Revised world maps and introduction, p. 1–21. In McKerrow, W. S. and Scotese, C. R. (eds.), Palaeozoic Palaeogeography and Biogeography. Geological Society Memoir, 12.Google Scholar
Seilacher, A. 1961. Ein Füllhorn aus dem Hunsrückschiefer. Natur und Volk, 91(1):15–19.Google Scholar
Sobolev, D. N. 1914. Nabroski po filogenii goniatitov. Warszawa, 199 p.Google Scholar
Springer, F. 1926. American Silurian Crinoids. Smithsonian Institution Publication, 2872:1–239, 33 pls.Google Scholar
Steininger, J. 1849. Die Versteinerungen des Uebergangs-Gebirges der Eifel. Jahresbericht über den Schul-Cursus 1848/49 an dem Gymnasium zu Trier, 50 p.Google Scholar
St. John, O., and Worthen, A. H. 1875. Description of fossil fishes. Geological Survey of Illinois, Paleontology, 6:245–488.Google Scholar
Strimple, H. L. 1966. Some notes concerning the Allagecrinidae. Oklahoma Geology Notes, 26:99–111, 1 pl.Google Scholar
Stukalina, G. A. 1986. Zakonomernosti istoricheskogo rzvitiya krinoidei v rannemi srednem paleozoe SSSR [Laws of historical development of crinoidea in the early and middle Paleozoic of the USSR]. Akademiya Nauk SSSR, Paleontological Institut Moskova, 142 p., 32 pls.Google Scholar
Termier, G., and Termier, H. 1947. Paléontologie Marocaine I. Généralités sur les invertébres fossiles. Service de la Carte géologique Morocco, Notes et Mémoires, 69,391 p., 22 pls.Google Scholar
Termier, G., and Termier, H. 1950. Paléontologie Marocaine II. Invertébres de l'Ere Primaire. 4. Annélides, Arthropodes, Echinodermes, Conularides et Graptolithes. Service de la Carte géologique Morocco, Notes et Mémoires, 79(4), 279 p., pls. 184–241.Google Scholar
Termier, H. 1936. Études géologiques sur le Maroc central et le moyen atlas septentrional. Tome III. Paléontologie Pétrographie. Notes et Mémoires Service des Mines et de la Carte géologique, Algerie, 33:1087–1421, 23 pls.Google Scholar
Termier, H., and Termier, G. 1949. Hiérarchie et corrélations des caractéres chez les Crinoides fossiles. Bulletin Service de la Carte géologique Algerie, 1st Series, Paleontologie, 10:1–91, 8 pls.Google Scholar
Termier, H., and Termier, G. 1974. Une méthode nouvelle: L'utilisation des fragments d'échinoderms contenus dans le sédiments dévoniens et carboniféres du Maroc. Notes et Service géologique de Maroc, 35(255):27–53, 9 pls.Google Scholar
Wachsmuth, C., and Springer, F. 1886. Revision of the Palaeocrino-idea: Proceedings of the Academy of Natural Sciences of Philadelphia, Pt. III, Sec. 2. Discussion of the classification and relations of the brachiate crinoids, and conclusion of the generic descriptions (1886), p. 64–226.Google Scholar
Wanner, J. 1916. Die Permischen echinodermen von Timor, I. Teil. Palaontologie von Timor, 11:1–329, pls. 94–114.Google Scholar
Wanner, J. 1930. Neue Beiträge zur Kenntnis der Permischen Echinodermen von Timor III. Hypocriinae, Paracatillocrinus und Allagecrinus dux . Dienst van den Mijnbouw in Nederlansch-Indië, Wetenschappelijke Mededeelingen, 13:1–30, 2 pls.Google Scholar
Wanner, J. 1937. Neue beiträge zur kenntnis der Permischen echinodermen von Timor, VIII-XIII. Palaeontographica, supplement 4, IV Abteilungen, Abschnitt 1, 212 p., 14 pls.Google Scholar
Webster, G. D. 1987. Permian crinoids from the type-section of the Callytharra Formation, Callytharra Springs, western Australia. Alcheringa, 11:95–135.Google Scholar
Webster, G. D. 1998. Palaeobiogeography of Tethys Permian crinoids. Proceedings of the Royal Society of Victoria, 110(1/2):289–308.Google Scholar
Webster, G. D. 2003. Bibliography and index of Paleozoic crinoids, coronates, and hemistreptacrinoids 1758–1999. Geological Society of America Special Paper, 363:1–2235. (GSA Web site, http://crinoid.gsajournals.org/crinoid/crinoid-netscape.css)Google Scholar
Webster, G. D., Maples, C. G., Mawson, R., and Dastanpour, M. 2003. A cladid-dominated Early Mississippian crinoid and conodont fauna from Kerman Province, Iran and revision of the glossocrinids and rhenocrinids. Journal of Paleontology, Memoir, 60, 35 p.Google Scholar
Wedekind, R. 1913. Beiträge zur Kenntnis des Oberdevons am Nor-drande des Rheinischen Schiefergebirges. 2. Zur Kenntnis der Prolobitiden. Neues Jahrbuch für Mineralogie, Geologie und Paläontologie, 1913:78–95.Google Scholar
Wedekind, R. 1918. Die Genera der Palaeoammonoidea (Goniatiten). Palaeontographica, 62:85–184.Google Scholar
Wendt, J. 1988. Condensed carbonate sedimentation in the late Devonian of the eastern Anti-Atlas (Morocco). Eclogae Geologicae Helvetiae, 81:155–173.Google Scholar
Wendt, J. 1989. Facies pattern and paleogeography of the Middle and Late Devonian in the eastern Anti-Atlas (Morocco). Canadian Society of Petroleum Geologists Memoir, 14(1):467–480.Google Scholar
Wendt, J. 2003. Late Devonian pseudoplanktonic crinoids from Morocco. Neues Jahrbuch für Geologie und Paläontologie Monatshefte, 2003(3):153–163.Google Scholar
Wendt, J., and Belka, Z. 1991. Age and depositional environment of Upper Devonian (early Frasnian to early Famennian) black shales and limestone (Kellwasser facies) in the eastern Anti-Atlas, Morocco. Facies, 25:51–90.Google Scholar
Wendt, J., Aigner, T., and Neugebauer, J. 1984. Cephalopod limestone deposition on a shallow pelagic ridge: the Tafilalt Platform (Upper Devonian, eastern Anti-Atlas, Morocco). Sedimentology, 31:601–625.Google Scholar
Weyer, D. 1965. Triacrinus Münster 1839 (Crinoidea) aus der Wock-lumeria -Stufe des thüringischen Oberdevons. Geologie, Deutschland, 24(8):969–981.Google Scholar
Whitfield, R. P. 1905. Notice of a new crinoid and a new mollusk from the Portage rocks of New York. American Museum Natural History Bulletin, 21(2):17–20, pls. 1–4.Google Scholar
Yakovlev, N. N. 1949. Jaekelicrinus bashkivicus Yakovlev n. gen., n. sp. Journal of Paleontology, 23:435.Google Scholar
Zalesskey, M. D. 1911. Etude sur l'anatomie du Dadosylon tohihat-chaffi Göppert sp. Mémoires du Comité géologique, St. Petersbourg, serie 2, 68, 29 p.Google Scholar
Zenker, J. C. 1833. Beiträge zur Naturgeschichte der Urwelt. Organische Reste (Petrefacten) aus der Altenburger Braunkohlen-formation, dem Blankenburger Quadersandstein, jenaischen bunten Sandstein und böhmischen Übergangsgebirge. Jena, 67 p., 7 pls.Google Scholar