Brachiopods of the orders Strophomenida, Orthotetida, and Athyridida from the late middle Frasnian–early Famennian interval (hassi to triangularis conodont zones) in the Namur-Dinant Basin (southeastern margin of Laurussia; southern Belgium) are described. Nine genera and subgenera represented by 12 species are recognized here. One new genus, Retrorstrophia, and one new species, Douvillina area, are erected. Crinisarina stainbrooki is proposed to solve the homonymy between C. reticulata (Gosselet, 1877) and Cleiothyridina reticulata Stainbrook, 1947, the type species of Crinisarina Cooper and Dutro, 1982. A lectotype is selected and illustrated for C. reticulata. A new name is also proposed for Athyris reticulata Chen and Xu, 2000, which is assigned here to Crinisarina: C. shashishanensis nom. nov. In southern Belgium, the last representatives of the families Douvillinidae and Leptostrophiidae (Strophomenida) are from the Upper rhenana Zone (late Frasnian). Athyridid brachiopods, especially the representatives of the subfamily Cleiothyridininae constituted a significant part of the early Famennian recovery faunas with spiriferids and rhynchonellids, just after the late Frasnian mass extinction.

We present a critical review of the alpha taxonomy and evolution of Eocene North American paromomyid primates, based on analysis of more than 570 stratigraphically controlled dental and gnathic specimens from the early Eocene of the southern Bighorn Basin, Wyoming (Wasatchian, Willwood Formation). In addition to documenting numerous previously unpublished specimens of known taxa (including deciduous teeth), we also describe a new species, Phenacolemur willwoodensis n. sp., from the upper part of the Willwood Formation (Wa 5 and 6). The new species is intermediate in size between Phenacolemur simonsi and Phenacolemur citatus and has both primitive features (e.g., retention of m2–3 paraconids, relatively long molar trigonids) and derived traits (e.g., relatively reduced paraconid on m1 and no p4 paracristid, unlike Paromomys). Overall patterns of dental evolution in southern Bighorn Basin paromomyids provide some support for previously hypothesized periods of faunal change (Biohorizons). In particular, Phenacolemur praecox evolves into the similarly sized but morphologically distinct Phenacolemur fortior at Biohorizon A, and P. fortior is replaced by P. citatus just below Biohorizon B. Two taxa previously believed to have become extinct at Biohorizon A (Ignacius graybullianus, P. simonsi) are shown to have persisted about a million years longer than previously thought. The Bighorn Basin paromomyids are of general interest in comprising a very dense sample that allows for the study of patterns of evolution against the backdrop of well-understood patterns of change in other mammalian lineages, and in climatic variables.

Black shales of the Coroico Formation are part of a thick succession of Lower and Middle Ordovician strata that were deposited in the Cordillera Oriental foreland basin along the margin of West Gondwana. The basin was inhabited primarily by a cool-water, Atlantic-type graptolite fauna. Newly discovered material from rocks that crop out near the town of Consata in NW Bolivia include unexpected warm-water or Pacific-type elements such as Parisograptus caduceus and Pseudotrigonograptus within an assemblage dominated by abundant pendent Didymograptus specimens, Cryptograptus schaeferi, and diplograptaceans such as Oelandograptus oelandicus and Hustedograptus bulmani n. sp. Parisograptus caduceus has generally been considered to be restricted to oceanic depths below those of the epipalagic realm that occupied continental shelves, whereas the remainder of the assemblage is more characteristic of relatively shallow water, epicratonic sites. We interpret this mixed assemblage to be the Atlantic Province, West Gondwanan equivalent of the off-shore isograptid biofacies that is much more widely known from low latitude sites around the globe. These results suggest that in this region of West Gondwana, the properties of local water masses (productivity and physical features such as temperature, salinity, or oxygenation) strongly influenced graptolite species distribution and led to biofacies differentiation among coeval assemblages. It also suggests that some isograptids inhabited the epipelagic biotope in mid to high latitude regions. The new species Hustedograptus bulmani is described herein.

A bone bed in the middle part of the Javelina Formation (Maastrichtian) in Texas yielded parts of about 37 identifiable ceratopsid dinosaur bones, mostly appendicular and limb girdle elements belonging to one juvenile and two adult individuals of Torosaurus cf. utahensis. The bone bed is a lag assemblage comprising large immobile parts of the skeletons accumulated in an abandoned stream channel. In general form and proportions the postcranial bones are similar to those in Pentaceratops sternbergi and are not as robust as those in Torosaurus latus or Triceratops horridus. A few cranial elements are preserved, including parts of a parietal, squamosal, maxilla, and two dentaries. The form of the parietal fragment is comparable to that of a more nearly complete specimen of Torosaurus cf. utahensis collected nearby at about the same stratigraphic level. The bone bed material provides a basis for the first skeletal reconstruction of this enigmatic horned dinosaur. Most characters used in diagnoses of T. utahensis and T. latus are inadequate. Only the raised bar along the squamosal/parietal suture, present in T. latus; and the midline epiparietal, absent in T. latus, may discriminate the two species.

The Amphoracrinidae Bather, 1899 is redefined, and all genera of this family are diagnosed with objective characters. Dilatocrinus Webster and Lane, 1987 and Pimlicocrinus Wright, 1943 are transferred into the Amphoracrinidae. Type specimens that define Displodocrinus Webster and Lane, 1987 and Dilatocrinus Webster and Lane, 1987 are clarified. The emphasis here is on genus definition, and comprehensive species-level systematics is not attempted. However, Amphoracrinus blairi Miller and Gurley, 1896a and Amphoracrinus jessieae Miller and Gurley, 1896b are designated nomina dubia; Sampsonocrinus sp. Webster and Lane, 1987 is reassigned to Amphoracrinus rupinus Webster and Lane, 1987; Amphoracrinus divergens var. multiramosus (Meek and Worthen, 1866) is reassigned to Dilatocrinus multiramosus (Meek and Worthen, 1866); and Sunwaptacrinus nevadensis Webster and Lane, 1987 is reassigned to Displodocrinus nevadensis (Webster and Lane, 1987).

Bivalves from the Late Devonian pelagic or Hercynian Facies of classical regions such as eastern North America and Europe have not been investigated for almost a century.

A group of small, radially ribbed bivalves frequently occurs in association with ammonoids and conodonts in pelagic cephalopod limestones and shales of the latest Frasnian and early Famennian. These bivalves have traditionally been assigned to the Late Silurian genus Praecardium Barrande, 1881. Re-studying the types of the Late Devonian taxa, Cardium? vetustum Hall, 1843, Cardiola duplicata Münster, 1840, Praecardium clymeniae Beushausen, 1895, Praecardium melletes Clarke, 1904, and Praecardium multicostatum Clarke, 1904 shows that they differ significantly from Praecardium.

As a result, two new genera, Vetupraeca n. gen. and Mucopraeca n. gen, are established. Furthermore, neotypes are designated for Cardiola nehdensis, Kayser, 1873 and Vetupraeca clymeniae (Beushausen, 1895), and lectotypes are chosen for Mucopraeca multicostata (Clarke, 1904) and Vetupraeca duplicata (Münster, 1840). These bivalve taxa were widely distributed in the subtropical to tropical, latest Frasnian/early Famennian outer shelf habitats of Laurussia and Gondwana.

Based on a thorough examination of field and museum Climactichnites specimens, two species of this trace are recognized, each representing a unique behavioral variant produced by a soft-bodied animal in Late Cambrian intertidal environments. C. wilsoni represents surface-produced trails, whereas C. youngi is re-erected for burrows produced below the surface. Burrowing behavior is supported by: 1) the presence of C. youngi within, rather than on, the surface of beds; 2) the orientation of some burrows inclined to bedding; and 3) the occasional presence of distinct burrow fills. Burrows can also be distinguished morphologically from surface traces by the absence of lateral ridges and the presence of fine, mm-scale striations or grooves superimposed on the transverse bars and furrows. Burrowing behavior for the Climactichnites trailmaker was previously unknown and thus represents a new, although not entirely unexpected, behavior for this mollusk or mollusk-like animal. The body impression of the sedentary animal is removed to Musculopodus sedentarius n. igen. and isp. In the future, Musculopodus may be expanded to include the resting traces of other soft-bodied animals known from the fossil record. Currently, Climactichnites is known only from very shallow to emergent strata of North America; reports of this fossil in other parts of the world are misidentified trails produced by other animals.

Three fossil taxa of megapodagrionid damselflies are described and figured from the Paleogene localities in Europe on the basis of isolated wings. Eckfeldia superstes (Wappler, 2003) gen. nov. is described from the laminated mudstones of middle Eocene age from Eckfeld Maar, Germany. Furagrion jutlandicus (Henriksen, 1922) gen. nov. is recorded from the laminated claystones of lowermost Eocene age from the Ølst and Fur-Formation, Denmark, and an undetermined megapodagrionid damselfly is recognized from middle Eocene strata. Taphonomy and color preservation in the fossils are briefly considered. Characters used for phylogenetic analyses in extant and fossil Megapodagrionidae are discussed. The biogeographic and paleoecological implications of the new European fossils are briefly discussed.

Crowns of four crinoid species, Talarocrinus planus Weller, 1920, Phacelocrinus longidactylus (McChesney, 1860), Ampelocrinus mundus? Kirk, 1942, and Phanocrinus maniformis (Yandell and Shumard, 1847) are reported from the limestones of the Wymps Gap Member of the Greenbrier Formation in northern West Virginia. Collectively these species indicate an early Chesterian age for the Wymps Gap Member, with the occurrence of T. planus indicating correlation with the Ridenhower (Paint Creek) Formation of the Illinois Basin. A review of the 17 species currently placed in Talarocrinus suggests that the Talarocrinus zone ranges no higher than the Ridenhower Formation in the Upper Gnathodus bilineatus conodont zone of the late Visean. The new occurrence of A. mundus extends its range upwards from the Genevievian substage of the Chesterian.

Previous studies have used sequence stratigraphy to correlate the Wymps Gap Member with the lower part of the Union Limestone Formation of the Greenbrier Group in southern West Virginia, which in turn has been correlated with the Ridenhower Formation. The occurrence of T. planus in the Wymps Gap Member provides independent confirmation of the correlations based on sequence stratigraphy between the Appalachian and Illinois basins. Correlation of the overlying Reynolds Member of the Mauch Chunk Formation with the Glen Dean Formation of the Illinois Basin suggests a significant unconformity between the Greenbrier Formation and Mauch Chunk Formation in northern West Virginia.

Upper Carboniferous and Lower Permian shallow marine rocks in the Pequop Mountains of northeastern Nevada contain an abundance of fossils, especially fusulinids and fasciculate corals. As the composite stratigraphic section in this area represents the most nearly complete sequence of this type of corals anywhere in the Cordilleran miogeocline, this study was undertaken to document their occurrence sequentially. This information should be useful for comparison with other sections in the miogeocline containing similar corals.

Only one species of Durhamina is present in the Gzhelian rocks. Durhamina, Heintzella, and Paraheritschioides occur in the Asselian rocks, and all of these genera plus Wilsonastraea are present in the Sakmarian part of the section. Corals in the Artinskian and Kungurian are represented by an advanced species of Durhamina and the very advanced durhaminid, Sandolasma.

Newly described species are: Durhamina primitiva, D. snyderi, Sandolasma perplexa, Heintzella davydovi, Paraheritschioides nevadaensis, and Wilsonastraea fraseri.

New Mexico contains a significant record of trace fossil assemblages, in terms of both abundance and ichnodiversity, from Lower Permian nonmarine depositional settings. Most notable amongst these are the trace fossil assemblages in the Robledo Mountains Formation of the Robledo Mountains in Doña Ana County, southern New Mexico, as recognized by the recent proposal to designate this area as a national monument. These trace fossil assemblages formed on a tidal flat under largely non-marine conditions (Mack and James, 1986; Hunt et al., 1993; Lucas et al., 1995a, 1998) and are dominated by the trackways of tetrapods and arthropods, yielding important information on the paleoecology, diversity and behavior of late Paleozoic arthropods (Braddy and Briggs, 2002; Minter and Braddy, 2006a), as well as evidence of specialized foraging strategies (Minter et al., 2006). Additional Lower Permian trace fossil assemblages occur at a number of localities in New Mexico and represent a variety of nonmarine depositional settings.

Machaeridians are marine Palaeozoic fossils, known from sediments of early Ordovician to mid Permian age (Bengtson, 1978; Cooper and Grant, 1972; Kaasa, 1992). Rare, completely articulated specimens reveal that they were long, slender, bilaterally symmetrical animals, covered by a dorsal scleritome comprising longitudinally arranged series of calcite sclerites (e.g., Adrain et al., 1991; Högström and Taylor, 2001a, 2001b). However, most known machaeridians, including those described in this paper, comprise isolated disarticulated sclerites. Until recently, the taxonomic affinities of the group were uncertain, and historically they were assigned to a number of different phyla (e.g., see Adrain, 1992). However, the recent account of an exceptionally preserved specimen with soft tissue anatomy, from the Lower Fezouata Formation (Lower Ordovician, Tremadocian) of Morocco, now indicates that machaeridians were annelid worms (Vinther et al., 2008).

Hydrothermal vents and methane seeps sustain unique ecosystems with a highly adapted fauna that largely thrives on chemotrophic endosymbionts. This large biomass attracts not only predators but also parasitic taxa like the recently reported oophagous bivalve Acesta bullisis Vokes, 1963 that lives permanently attached to a vestimentiferan tube worm (Järnegren et al., 2005). Here I report large brachiopods of the Early Cretaceous seep-restricted genus Peregrinella Oehlert, in Fischer, 1887 that were infested by polychaete tubes inside their shells during their lifetime.

The Huntington cluster (USNM 536849; Fig. 1) measures 4.1 mm in total length and is comprised of five elements of the maxillary apparatus; the first and second pair of maxillae (right and left MI and MII) and the basal plate (Bp) fitted into the bight of the right MI (for descriptive terminology for ramphoprionids, see Kielan-Jaworowska, 1966; Eriksson, 2001; Fig. 2). Both the MI are broken and lack approximately a third of the anterior end. The right MI also lacks part of its posterior denticulated ridge; the posteriormost tip (p in Fig. 1.2) is, however, still preserved and partly embedded in the slab. Most of the right MII is missing and only a thin rim is exposed, showing the outline of the element. The left MII is almost complete, lacking only a fifth of the anterodorsal end. The basal plate is well preserved and can be seen in full detail, except for a part of the antero-dextral flange, which is broken off (Fig. 1.3). Adjacent to the maxillary apparatus are other fragments that might be the carriers (Fig. 1.4), i.e., the posteriormost, supporting elements of the apparatus. Except for these possible carriers, all elements are more or less in natural position.

The genus Bicarinella Rode et al., 2003 was erected for a new hipponicharionid bradoriid species described from the early Cambrian of East Antarctica, characterized by a subtriangular carapace with prominent anterior and posterior lobes developed as two distinctive, sharp ridges (bi = two, carina = ridges). Unfortunately, the name Bicarinella is preoccupied by two different gastropod genera: Bicarinella Waterhouse 1966, a Permian gastropod from New Zealand and Australia, and Bicarinella Akopyan 1976, a gastropod from Late Cretaceous strata of Armenia, Serbia, Romania, Tajikistan, and Egypt (Mennessier, 1994; Banjac, 1998; Pana, 1998). Mennessier (1994) transferred Bicarinella Akopyan, 1976 from its original status as an independent taxon to a subgenus of Pseudomesalia Douvillé 1916, but subsequent workers have continued to consider Bicarinella a valid genus (Pena, 1998; Banjac, 1998). Due to the preoccupation, the bradoriid genus is herein renamed in accordance with the requirement of the International Code of Zoological Nomenclature (1999, article 60). It is also noted here that the one of the two distinct gastropod genera should be renamed. The name Bicarinellata (bi = two, carina = ridges) is proposed as a replacement name for Bicarinella Rode et al., 2003. This name retains the original prefixes to preserve taxonomic stability as much as possible. The type species of Bicarinellata is B. evansi by original designation (Rode et al., 2003).