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Upper Homerian (upper Wenlock, Silurian) graptolites from arctic Canada

Published online by Cambridge University Press:  14 July 2015

A. C. Lenz  and
A. Kozłowska-Dawidziuk
A. C. Lenz
Affiliation:
Department of Earth Sciences, University of Western Ontario, London, Ontario, Canada N6A 5B7,
A. Kozłowska-Dawidziuk
Affiliation:
Institute of Paleobiology, Polish Academy of Sciences, Twarda 51/55, 00-818 Warszawa, Poland,
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Abstract

An extraordinarily well-preserved upper Homerian (uppermost Wenlock) post-extinction (post-lundgreni Event) graptolite fauna is described from Arctic Canada. The fauna of dendroids, retiolitids and monograptids, predominantly of uncompressed and isolated material, demonstrates almost totally new morphological features in comparison with the pre-extinction fauna. The two species of the lowest post-extinction fauna are followed by a rapid proliferation in the middle part of the interval, resulting in the maximum diversity, 21 species, for the entire upper Homerian. The upper Homerian is divided into three biostratigraphic units: the lower nassadubius Biozone, middle praedeubeli-deubeli Biozone, and the upper ludensis Biozone. The total fauna consists of ten species of monograptids and 11 species of retiolitids; of the latter group, three (Baculograptus, Papiliograptus and Doliograptus) are new genera, and six (B. batesi, P. papilio,P.? petilus,D. latus,Doliograptus sp. A, and Spinograptus praerobustus) are new species. Among the recognized species of monograptids, only Colonograptus schedidoneus and Lobograptus? cornuatus are confined to the Arctic, while five species of retiolitids (B. batesi,P.? petilus,D. latus,Doliograptus sp. A, and Spinograptus praerobustus) appear to be endemic.

Type
Research Article
Information
Journal of Paleontology , Volume 76 , Issue 2 , March 2002 , pp. 321 - 346
Copyright
Copyright © The Paleontological Society

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References

Barrande, J. 1850. Graptolites de Bohême. Published by the author (Prague), p. 1–74.Google Scholar
Bates, D. E. B. 1990. Retiolite nomenclature and relationships. Journal of the Geological Society, London, 147:717–723.CrossRefGoogle Scholar
Bates, D. E. B., and Kirk, N. H. 1992. The ultrastructure, mode of secretion and functioning of a number of Llandovery ancorate diplograptid and retiolitid graptolites. Modern Geology, 17:1–270.Google Scholar
Berry, W. B. N., and Murphy, M. A. 1975. Silurian and Devonian graptolites of central Nevada. University of California Publications in Geological Sciences, 110:1–109.Google Scholar
Bouček, B., and Münch, A. 1952. Retioliti stredoevropského svrchního wenlocku a ludlowu. Sborník ústředního ústavu Geologického, XIX. Oddíl paleontolický, p. 1–151.Google Scholar
Churkin, M., and Carter, C. 1996. Stratigraphy, structure, and graptolites of an Ordovician and Silurian sequence in the Terra Cotta Mountains, Alaska Range, Alaska. U.S. Geological Survey Professional Paper 1555, p. 1–84.Google Scholar
Cuerda, A. 1974. Monograpten aus dem Ludlow Boliviens. Neues Jahrbuch für Geologie und Paläontologie, Monatshefte, 1974:321–335.Google Scholar
Eisenack, A. 1951. Retioliten aus dem Graptolithengestein. Palaeontographica, 100:129–163.Google Scholar
Erdtmann, B.-D., and Prezbindowski, D. R. 1973. Niagaran (Middle Silurian) interreef fossil burial environments in Indiana. Neues Jahrbuch für Geologie und Paläontologie, Abhandlugen, 144:342–372.Google Scholar
Frech, F. 1897. Lethaea geognostica—1, Lethaea paleozoica, 1, Graptolithen. Schweizerbart, Stuttgart, p. 544–684.Google Scholar
Gutiérrez-Marco, J. C., Lenz, A. C., Robardet, M., and Picarra, J. M. 1996. Wenlock-Ludlow graptolite biostratigraphy and extinction: a reassessment from the southwestern Iberian Peninsula (Spain and Portugal). Canadian Journal of Earth Sciences, 33:656–663.CrossRefGoogle Scholar
Hall, J. 1843. Geology of New York, IV, Survey of the Fourth Geological District. Natural History of New York. New York.Google Scholar
Hall, J. 1851. New genera of fossil corals. American Journal of Science, 11:398–401.Google Scholar
Hall, J. 1858. Descriptions of Canadian graptolites. Geological Survey of Canada, Report for 1857, p. 111–145.Google Scholar
Holland, C. H., Rickards, R. B., and Warren, P. T. 1969. The Wenlock graptolites of the Ludlow district, Shropshire, and their stratigraphical significance. Palaeontology, 12:663–683.Google Scholar
Holm, G. 1890. Gotlands graptoliter. Svenska Vetenskaps-Akademiens handlingar, 16:1–34.Google Scholar
Jaeger, H. 1959. Graptolithen und Stratigraphie des jüngsten Thüringer Silurs. Abhandlugen der deutschen Akademie der Wissenschaften zu Berlin. Klasse fur Chemie, Geologie und Biologie, 2:1–197.Google Scholar
Jaeger, H. 1990. New geological and biostratigraphical data on the Silurian of SE-Sardinia. Close affinity with Thuringia, p. 101–117. In Barca, S., and Jaeger, H. (eds.), Bolletino della Societa Geologica Italiana, 108.Google Scholar
Jaeger, H. 1991. Neue Standard-Graptolithenzonenfolge nach der “Grossen Krise” an der Wenlock/Ludlow-Grenze (Silur). Neues Jahrbuch für Geologie und Paläontologie Abhandlungen, 182:303–354.CrossRefGoogle Scholar
Jaekel, O. 1889. Ueber das Alter des sogenanten Graptolithen-Gesteins. Zeitschrift der deutsches geologischen Gesellshaft, 41:653–716.Google Scholar
Koren', T. N. 1991. The lundgreni extinction event in central Asia and its bearing on graptolite biochronology within the Homerian. Proceedings of the Estonian Academy of Sciences, Geology, 40:74–78.Google Scholar
Koren', T. N. 1992. Novye pozdnevenlokski monograptidy Alayskogo khrebta. Paleontologicheskiy zhurnal, 2:21–33.Google Scholar
Koren', T. N. 1994. The Homerian monograptid fauna of central Asia: zonation, morphology and phylogeny, p. 140–148. In Chen, X., Erdtmann, B.-D., and Ni, Y.-N. (eds.), Graptolite Research Today. University Press, Nanjing.Google Scholar
Koren', T. N., and Rinenberg, R. E. 1988. The lundgreni event in the late Wenlock pelagic facies of South Tien Shan, p. 15–18. In Graptolites in Earth History, Abstracts of the Fifth Symposium on Investigations on Graptolites in the USSR, Vilnius, Lithuania.Google Scholar
Koren', T. N., and Suyarkova, A. A. 1994. Monograptus deubeli and praedeubeli (Wenlock, Silurian) in the Asian part of the former Soviet Union. Alcheringa, 18:85–101.CrossRefGoogle Scholar
Koren', T. N., and Urbanek, A. 1994. Adaptive radiation of monograptids after the Late Wenlock crisis. Acta Palaeontologica Polonica, 39:137–167.Google Scholar
Kozłowska-Dawidziuk, A. 1995. Silurian retiolitids of the East European Platform. Acta Palaeontologica Polonica, 40:261–326.Google Scholar
Kozłowska-Dawidziuk, A. 1997. Retiolitid graptolite Spinograptus from Poland and its membrane structure. Acta Palaeontologica Polonica, 42:391–412.Google Scholar
Kozłowska-Dawidziuk, A. 1999. Wydarzenie Cyrtograptus lundgreni a rozwój retiolitów (Graptolithina) na platformie wschodnioeuropejskiej. Przegląd Geologiczny, 47:354–358.Google Scholar
Kozłowska-Dawidziuk, A., Lenz, A. C., and Štorch, P. 1998. Late Homerian-early Ludlow post-extinction graptolites, Všeradice, Czech Republic, p. 97–100. In Gutiérrez-Marco, J. C. and Rábano, I. (eds.), Proceedings 6th International Graptolite Conference (GWG-IPA) and 1998 Field Meeting, IUGS Subcommission on Silurian Stratigraphy. Temas Geológico-Mineros ITGE.Google Scholar
Kozłowska-Dawidziuk, A., Lenz, A. C., and Štorch, P. 2001. Upper Wenlock and lower Ludlow (Silurian) post-extinction graptolites, Všeradice section, Barrandian area, Czech Republic. Journal of Paleontology, 75:147–164.CrossRefGoogle Scholar
Kozłowska-Dawidziuk, A., Porębska, E., and Masiak, M. 2000. Geochemistry, acritarchs, and graptolites of the East European Platform. Palaeontology Down Under 2000, Geological Society of Australia, Abstracts 61:49.Google Scholar
Lapworth, C. 1873. On an improved classification of the Rhabdophora . Geological Magazine, 10:500–504,555–560.CrossRefGoogle Scholar
Lenz, A. C. 1975. Silurian graptolites from eastern Gaspé, Quebec. Canadian Journal of Earth Sciences, 12:77–89.CrossRefGoogle Scholar
Lenz, A. C. 1980. Wenlockian graptolite reference section, Clearwater Creek, Nahanni National Park, Northwest Territories. Canadian Journal of Earth Sciences, 17:1075–1086.CrossRefGoogle Scholar
Lenz, A. C. 1992. Some late Wenlock and Ludlow monograptids graptolites, Cornwallis Island, Arctic Canada. Canadian Journal of Earth Sciences, 29:200–207.CrossRefGoogle Scholar
Lenz, A. C. 1993a. Late Wenlock-Ludlow (Silurian) extinction, evolution, and biostratigraphy: perspectives from Arctic Canada. Canadian Journal of Earth Sciences, 30:491–498.CrossRefGoogle Scholar
Lenz, A. C. 1993b. Late Wenlock and Ludlow (Silurian) Plectograptinae (retiolitid graptolites), Cape Phillips Formation, Arctic Canada. Bulletins of American Paleontology, 104:1–52.Google Scholar
Lenz, A. C. 1994a. Extinction and opportunistic evolution among late Wenlock graptolites. Lethaia, 27:111–117.CrossRefGoogle Scholar
Lenz, A. C. 1994b. The graptolites “Pristiograptus“ praedeubeli (Jaeger) and “Pristiograptus“ ludensis (Murchison) (uppermost Wenlock, Silurian) from Arctic Canada: taxonomy and evolution. Canadian Journal of Earth Sciences, 31:1419–1426.CrossRefGoogle Scholar
Lenz, A. C. 1994c. New upper Homerian (uppermost Wenlock; Silurian) monograptids from Arctic Canada. Canadian Journal of Earth Sciences, 31:1770–1784.CrossRefGoogle Scholar
Lenz, A. C. 1994d. Uppermost Wenlock and lower Ludlow plectograptine graptolites, Arctic Islands, Canada: new isolated material. Journal of Paleontology, 68:851–860.CrossRefGoogle Scholar
Lenz, A. C. 1994e. A sclerotized retiolitid, and its bearing on the origin and evolution of Silurian retiolitids graptolites. Journal of Paleontology, 68:1344–1349.CrossRefGoogle Scholar
Lenz, A. C. 1995. Upper Homerian (Wenlock, Silurian) graptolites and graptolite biostratigraphy, Arctic Archipelago, Canada. Canadian Journal of Earth Sciences, 32:1378–1392.CrossRefGoogle Scholar
Lenz, A. C., Chen, X., and Ni, Y.-N. 1996. Wenlock, Ludlow and Pridoli? graptolites from Yulin, Guangxi, China. Canadian Journal of Earth Sciences, 33:1390–1401.CrossRefGoogle Scholar
Lenz, A. C., and Kozłowska-Dawidziuk, A. 2000. The Upper Wenlock Graptolite extinction (lundgreni extinction): perspectives from the Arctic Canada. Palaeontology Down Under 2000, Geological Society of Australia, Abstracts 61:55.Google Scholar
Lenz, A. C., and Kozłowska-Dawidziuk, A. 2001. Upper Wenlock (Silurian) graptolites of Arctic Canada: pre-extinction, lundgreni Biozone fauna. Palaeontographica Canadiana.CrossRefGoogle Scholar
Lenz, A. C., and Melchin, M. J. 1987. Silurian retiolitids from the Cape Phillips Formation, Arctic Canada. Bulletin of the Geological Society of Denmark, 35:161–170.Google Scholar
Lenz, A. C., and Melchin, M. J. 1991. Wenlock (Silurian) graptolites, Cape Phillips Formation, Canadian Arctic Islands. Transactions of the Royal Society of Edinburgh: Earth Sciences, 27:211–237.CrossRefGoogle Scholar
Lenz, A. C., and Melchin, M. J. 1997. Graptolite faunas, p. 22–24. In Norford, B. S. (ed.), Correlation and biostratigraphy of the Silurian rocks of Canada. International Union of Geological Sciences, Publication 33.Google Scholar
Maletz, J., Königshof, P., Meço, S., and Schindler, E. 1998. Late Wenlock to Early Ludlow graptolites from Albania. Senckenbergiana Lethaea, 78:141–151.CrossRefGoogle Scholar
Murchison, R. I. 1839. The Silurian System. John Murray, London, p. i–xxxii + 1–768.Google Scholar
Nicholson, H. A. 1872. Monograph of British Graptolitidae. Blackwood and Sons, Edinburgh-London, 133 p.Google Scholar
Obut, A. M., and Zaslavskaya, N. 1983. [The families of retiolitids and their phylogenetic relationships]. Trudy Instytut Geologii i Geofiziki, SSSR, Sibirskoe Otdelenie, 538:103–113. (In Russian)Google Scholar
Palmer, D. 1970. Monograptus ludensis zone graptolites from the Devilsbit Mountain District, Tipperary. Scientific Proceedings of the Royal Dublin Society, Series A, 3:335–342.Google Scholar
Poˇta, P. 1894. Bryozoaires, Hydrozoaires et partie des Anthozoaires. In Barrande, J. (ed.), Systême Silurien du centre de la Bohême, 8:1–230.Google Scholar
Příbyl, A. 1942. Revision der Pristiograpten aus den Untergattugen Colonograptus n. subg. and Saetograptus n. subg. Mitteilungen der tschechischen Akademie der Wissenschaften, 52:1–22.Google Scholar
Příbyl, A. 1943. Revision aller Vertreter der Gattung Pristiograptus aus der Gruppe P. dubius und P. vulgaris aus dem böhmischen und ausländischen Silur. Mitteilungen der tschechischen Akademie der Wisseschaften, Jahrgang, 53:1–49.Google Scholar
Prout, H. G. 1851. Description of a new graptolite fauna in the Lower Silurian rocks near the Falls of St. Croix River. American Journal of Science, 23:187–191.Google Scholar
Rickards, R. B. 1967. The Wenlock and Ludlow succession in the Howgill Fells (north-west Yorkshire and Westmoreland.). Quarterly Journal of the Geological Society of London, 123:215–251.CrossRefGoogle Scholar
Rickards, R. B., and Sandford, A. C. 1998. Llandovery-Ludlow graptolites from central Victoria: new correlation perspectives of the major formations. Australian Journal of Earth Sciences, 45:743–763.CrossRefGoogle Scholar
Rickards, R. B., and Wright, A. J. 1997a. Graptolite zonation in the late Wenlock (Early Silurian), with a new graptolite-brachiopod fauna from New South Wales. Records of the Australian Mueum, 49:229–248.CrossRefGoogle Scholar
Rickards, R. B., and Wright, A. J. 1997b. Graptolites of the Barnby Hills shale (Silurian, Ludlow), New South Wales, Australia. Proceedings of the Yorkshire Geological Society 51:209–227.CrossRefGoogle Scholar
Rickards, R. B., Packham, G. H., Wright, A. J., and Williamson, P. L. 1995. Wenlock and Ludlow graptolite faunas and biostratigraphy of the Quarry Creek district, New South Wales. Association of Australasian Palaeontologists Memoir 17:1–68.Google Scholar
Ruedemann, R. 1904. Graptolites of New York, Pt. 1, New York State Museum Bulletin, Memoir, 7:457–803.Google Scholar
Ruedemann, R. 1925. Some Silurian (Ontarian) faunas of New York. New York State Museum Bulletin 265:1–134.Google Scholar
Sherwin, L. 1975. Silurian graptolites from the Forbes district, New South Wales. Records of the Geological Survey of New South Wales, 16:227–237.Google Scholar
Skevington, D. 1963. Graptolites from the Ontikan limestones (Ordovician) of Öland, Sweden. Publications from the Palaeontological Institution of the University of Uppsala, No. 46:1–62.Google Scholar
Spencer, J. W. 1878. Graptolites of the Niagara Formation. Canadian Naturalist, 8:457–463.Google Scholar
Suess, E. 1851. Über böhmische Graptoliten. Naturwissenschaftliche Abhandlungen von W. Haidinger, 4:87–134.Google Scholar
Thorsteinsson, R. 1958. Cornwallis and Little Cornwallis Islands, District of Franklin, Northwest Territories. Geological Survey of Canada, Memoir 294, 134 p.Google Scholar
Törnquist, S. L. 1887. Anteckningar om de alder Paleozoiska Leden i Osthuringen och Vogtland. Geologiska Föreningens Förhandlingar, 9:471–491.CrossRefGoogle Scholar
Torsvik, T. H., Smethurst, M. A., Meert, J. G., Van Der Voo, R., McKerrow, W. S., Brasier, M. D., Strut, B. A., and Walderhaug, H. J. 1996. Continental break-up and collision in the Neoproterozoic and Palaeozoic: a tale of Baltica and Laurentia. Earth Sciences Reviews, 40:229–258.CrossRefGoogle Scholar
Tullberg, S. A. 1883. Skånes graptoliter, I and II. Sveriges Geologiska Undersökning, Series C, 50:1–43.Google Scholar
Urbanek, A. 1958. Monograptidae from erratic boulders of Poland. Palaeontologia Polonica, 9:1–105.Google Scholar
Wiman, C. 1895. Über die Graptolithen. University of Uppsala, Geological Institute, Bulletin 2:239–316.Google Scholar
Wood, E. M. R. 1900. The Lower Ludlow Formation and its graptolite fauna. Quarterly Journal of the Geological Society of London, 56:415–492.CrossRefGoogle Scholar