Chapter 2 sets out the history of the Province from 1348 to 1559 showing first its resilience in the face of the Black Death with new foundations in Ireland and the establishment of the nuns’ convent at Dartford. Its resilence is then shown against theological attack in anti-fraternal literature, including writings by Wycliff, and the rise of Lollardy. The Province’s continuing value to key supporters is shown through the patronage manifest in church decoration, through lay burials and grants of confraternity, while their secure place in civic life before the Reformation is seen in relation to the guilds associated with their churches. The sudden collapse of the Province at the Henrician Reformation is then examined to identify several factors, the most important of which was the crown’s imposition of its agents as Provincials and Priors.