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Adaptive Herbivore Ecology
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Book description

The adaptation of herbivore behaviour to seasonal and locational variations in vegetation quantity and quality is inadequately modelled by conventional methods. Norman Owen-Smith innovatively links the principles of adaptive behaviour to their consequences for population dynamics and community ecology, through the application of a metaphysiological modelling approach. The main focus is on large mammalian herbivores occupying seasonally variable environments such as those characterised by African savannahs, but applications to temperate zone ungulates are also included. Issues of habitat suitability, species coexistence, and population stability or instability are similarly investigated. The modelling approach accommodates various sources of environmental variability, in space and time, in a simple conceptual way and has the potential to be applied to other consumer-resource systems. This text highlights the crucial importance of adaptive consumer responses to environmental variability and is aimed particularly at academic researchers and graduate students in the field of ecology.


Review of the hardback:' … an enjoyable and informative addition to the ecological literature. We are convinced others will find the book equally stimulating and useful.'

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Abrams P. A. 1994. The fallacies of “ratio dependent” predation. Ecology 75: 1842–50
Abrams P. 1997. Anomalous predictions of ratio-dependent models of predation. Oikos 80: 163–71
Abrams P. A. & Ginzburg L. R.. 2000. The nature of predation: prey dependent, ratio dependent or neither. Trends Ecol. Evol. 15: 337–41
Adamczewski J. Z., Gates C. C., Hudson R. J. & Price M. A.. 1987. Seasonal changes in body composition of mature female caribou and their calves on an Arctic island with limited winter resources. Can. J. Zool. 65: 1149–57
Adamczewski J. Z., Hudson R. J. & Gates C. C.. 1993. Winter energy balance and activity of female caribou on Coats Island, Northwest Territories: the relative importance of foraging and body reserves. Can. J. Zool. 71: 1221–9
Akcakaya H. R. 1992. Population cycles of mammals: evidence for ratio-dependent predation hypothesis. Ecol. Monogr. 62: 119–42
Akcakaya H. R., Arditi R. & Ginzburg L. R.. 1995. Ratio-dependent predation: an abstraction that works. Ecology 76: 995–1004
Alexander R. McN. 1991. Optimization of gut structure and diet for higher vertebrate herbivores. Phil. Trans. R. Soc. Lond. B. 333: 249–55
Allden W. G. & Whittaker I. A. D. McD.. 1970. Determinants of herbage intake by grazing sheep: the interrelationships of factors influencing herbage intake and availability. Aust. J. Agric. Res. 21: 755–66
Allen-Rowlandson, T. S. 1980. The social and spatial organisation of the greater kudu in the Andries Vosloo Kudu Reserve, Eastern Cape. MSc thesis, Rhodes University
Anderson E. W. & Scherzinger R. J.. 1975. Improving quality of winter forage for elk by cattle grazing. J. Range Mgmt 28: 120–5
Andrew M. H. 1988. Grazing impact in relation to livestock watering points. Trends Ecol. Evol. 3: 336–9
Arditi R & Dacorogna B.. 1988. Optimal foraging on arbitrary patch distributions and the definition of habitat patches. Am. Nat. 131: 837–46
Arditi R. & Ginzburg L. R.. 1989. Coupling in predator-prey dynamics: ratio dependence. J. Theor. Biol. 139: 311–26
Ash A. J., McIvor J. G., Corfield J. P. & Winter W. H.. 1996. How land condition alters plant-animal relationships in Australia's tropical rangelands. Agric. Ecosyst. Environ. 56: 77–92
Astrom A., Lundberg P. & Danell K.. 1990. Partial prey consumption by browsers: trees as patches. J. Anim. Ecol. 59: 287–300
Baile C. A. & Forbes J. M.. 1974. Control of feed intake and regulation of energy balance in ruminants. Physiol. Rev. 54: 160–216
Bailey D. W., Gross J. E., Laca E. A., Rittenhouse L. R., Coughenour M. B., Swift D. M. & Sims P. L.. 1996. Mechanisms that result in large herbivore grazing distribution patterns. J. Range Mgmt 49: 386–400
Bailey D. W., Walker J. W. & Rittenhouse L. R.. 1990. Sequential analysis of cattle location: day to day movement patterns. Appl. Anim. Behav. Sci. 25: 137–48
Baker D. L. & Hobbs N. T.. 1987. Strategies of digestion: digestive efficiency and retention time of forage diets in montane ungulates. Can. J. Zool. 65: 1978–84
Bartmann, R. M., G. C. White & L. H. Carpenter. 1992. Compensatory mortality in a Colorado mule deer population. Wildl. Monogr. No. 121, 39pp
Bednekoff P. A. & Houston A. I.. 1994. Optimizing fat reserves over the entire winter: a dynamic model. Oikos 71: 408–15
Begon, M., J. Harper & C. Townsend. 1996. Ecology: Individuals, Populations and Communities, 3rd edition. Blackwell, Oxford
Behnke, R. H., I. Scoones & C. Kerven (eds). 1993. Range Ecology at Disequilibrium. ODI Publications, London
Bell, R. H. V. 1970. The use of the herb layer by grazing ungulates in the Serengeti. In Animal Populations in Relation to their Food Resources, ed. A. Watson (Symp. Brit. Ecol. Soc.), pp. 111–24. Blackwell, Oxford
Bell R. H. V. 1971. A grazing ecosystem in the Serengeti. Scient. Am. 225: 86–93
Bellows T. S. 1981. The descriptive properties of some models for density dependence. J. Anim. Ecol. 50: 139–56
Belovsky G. E. 1978. Diet optimization in a generalist herbivore: the moose. Theor. Popul. Biol. 14: 105–34
Belovsky G. E. 1984. Herbivore optimal foraging: a comparative test of three models. Am. Nat. 124: 97–115
Belovsky G. E. 1986. Optimal foraging and community structure: implications for a guild of generalist grassland herbivores. Oecologia 70: 35–52
Belovsky, G. E. 1990. How important are nutrient constraints in optimal foraging models or are spatial/temporal factors more important? In Behavioral Mechanisms of Food Selection, ed. R. N. Hughes, pp. 255–80. Springer-Verlag, Berlin and Heidelberg
Belovsky G. E. 1994. How good must models and data be in ecology?Oecologia 100: 475–80
Belovsky G. E. & Schmitz O. J.. 1993. Owen-Smith's evaluation of herbivore foraging models: what is constraining?Evol. Ecol. 7: 525–9
Belovsky G. E. & Slade J. B.. 1986. Time budgets of grassland herbivores: body size similarities. Oecologia 70: 53–62
Bergman C. M., Fryxell J. M. & Gates C. C.. 2000. The effect of tissue complexity and sward height on the functional response of Wood Bison. Funct. Ecol. 14: 61–9
Berryman A. A. 1992. The origins and evolution of predator-prey theory. Ecology 73: 1530–5
Berryman A. A. 1999. Alternative perspectives on consumer-resource dynamics: a reply to Ginzburg. J. Anim. Ecol. 68: 1263–6
Berryman A. A., Gutierrez A. P. & Arditi R.. 1995. Credible, parsimonious and useful predator-prey models — a reply to Abrams, Gleeson and Sarnelle. Ecology 76: 1980–5
Boggs C. L. 1992. Resource allocation: exploring connections between foraging and life history. Funct. Ecol. 6: 508–18
Boomker, E. A. 1987. Fermentation and digestion in the kudu. PhD thesis, University of Pretoria
Borner M., FitzGibbon C. D., Borner M., Caro T. M., Lindsay W. K., Collins D. A. & Holt M. E.. 1987. The decline of the Serengeti Thomson's gazelle population. Oecologia 73: 32–40
Botkin, D. B. 1990. Discordant Harmonies. A New Ecology for the Twenty First Century. Oxford University Press, Oxford
Bradbury J. W., Vehrencamp S. L., Clifton K. E. & Clifton L. M.. 1996. The relationship between intake rate and local forage abundance in wild Thomson's gazelles. Ecology 77: 2237–55
Brooks, P. M. & J. A. W. Macdonald. 1983. The Hluhluwe-Umfolozi Reserve: An ecological case history. In Management of Large Mammals in African Conservation Areas, ed. R. N. Owen-Smith, pp. 51–77. Haum, Pretoria
Brown J. H., Marquet P. A. & Taper M. L.. 1993. Evolution of body size: consequences of an energetic definition of fitness. Am. Nat. 142: 573–84
Burgman, M., S. Ferson & H. R. Akcakaya. 1992. Risk Assessment in Conservation Biology. Chapman & Hall, London
Burlison A. J., Hodgson J. & Illius A. W.. 1991. Sward canopy structure and the bite dimensions and bite weight of grazing sheep. Grass Forage Sci. 46: 29–38
Cahill G. F. 1970. Starvation in man. New Engl. J. Med. 282: 668–75
Calder, W. A. III. 1984. Size, Function and Life History. Harvard University Press, Cambridge, MA
Campling, R. C. 1970. Physical regulation of voluntary intake. In Physiology of Digestion and Metabolism in the Ruminant, ed. A. J. Phillipson, pp. 226–34. Oriel Press, Newcastle
Case, T. J. 2000. An Illustrated Guide to Theoretical Ecology. Oxford University Press, Oxford
Caughley G. 1970. Eruption of ungulate populations with emphasis on Himalayan Thar in New Zealand. Ecology 51: 53–72
Caughley, G. 1976a. Plant-herbivore systems. In Theoretical Ecology, ed. R. M. May, pp. 94–113. Blackwell, Oxford
Caughley, G. 1976b. Wildlife management and the dynamics of ungulate populations. In Advances in Applied Biology, Vol. 1, ed. T. H. Coaker, pp. 183–245. Academic Press, London
Caughley G. 1976c. The elephant problem — an alternative hypothesis. E. Afr. Wildl. J. 14: 265–83
Caughley G. 1982. Vegetation complexity and the dynamics of modelled grazing systems. Oecologia 54: 309–12
Caughley G. 1994. Directions in conservation biology. J. Anim. Ecol. 63: 215–44
Caughley, G. & J. H. Lawton. 1981. Plant-herbivore systems. In Theoretical Ecology, 2nd edition, ed. R. M. May, pp. 132–66. Blackwell, Oxford
Caughley, G. & A. R. E. Sinclair. 1994. Wildlife Ecology and Management. Blackwell, Oxford
Chan-McLeod A. C. A., White R. G. & Holleman D. F.. 1994. Effects of protein and energy intake, body condition, and season on nutrient partitioning and milk production in caribou and reindeer. Can. J. Zool. 72: 938–47
Chapin F. S.. 1980. The nutrition of wild plants. A. Rev. Ecol. Syst. 11: 233–60
Charnov E. L. 1976. Optimal foraging: attack strategy of a mantid. Am. Nat. 110: 141–51
Clark, C. W. 1990. Mathematical Bioeconomics. The Optimal Management of Renewable Resources. 2nd edition. John Wiley, New York
Clark C. W. 1993. Dynamic models of behavior: an extension of life history theory. Trends Ecol. Evol. 8: 205–9
Clark, C. W. & M. Mangel. 1999. Dynamic State Variable Models in Ecology. Oxford University Press, Oxford
Clutton-Brock T. H., Albon S. D. & Guinness F. E.. 1989. Fitness costs of gestation and lactation in wild mammals. Nature 337: 260–2
Clutton-Brock T. H., Illius A. W., Wilson K., Grenfell B. T., MacColl A. & Albon S. D.. 1997. Stability and instability in ungulate populations: an empirical analysis. Am. Nat. 149: 196–219
Clutton-Brock T. H., Major M. & Guinness F. E.. 1985. Population regulation in male and female red deer. J. Anim. Ecol. 54: 831–46
Clutton-Brock T. H., Price O. F., Albon S. D. & Jewell P. A.. 1991. Persistent instability and population regulation in Soay sheep. J. Anim. Ecol. 60: 593–608
Coe M. J., Cumming D. H. M. & Phillipson J.. 1976. Biomass and production of large herbivores in relation to rainfall and primary production. Oecologia 22: 341–54
Coley P. D. & Barone J. A.. 1996. Herbivory and plant defenses in tropical forests. A. Rev. Ecol. Syst. 27: 305–35
Coley P. D., Bryant J. P. & Chapin F. S.. 1985. Resource availability and plant anti-herbivore defense. Science 230: 895–9
Cooper, S. M. 1985. Factors influencing the utilization of woody plants and forbs by ungulates. PhD thesis, University of the Witwatersrand
Cooper S. M. 1990. The hunting behaviour of spotted hyenas in a region containing both sedentary and migratory prey. Afr. J. Ecol. 28: 131–41
Cooper S. M. & Owen-Smith N.. 1986. Effects of plant spinescence on large mammalian herbivores. Oecologia 68: 446–55
Cooper S. M., Owen-Smith N. & Bryant J. P.. 1988. Foliage acceptability to browsing ruminants in relation to seasonal changes in the leaf chemistry of woody plants in a South Africa savanna. Oecologia 75: 336–42
Coppock D. L., Swift D. M., Ellis J. E. & Galvin K.. 1986. Seasonal patterns of energy allocation to basal metabolism, activity and production for livestock in a nomadic pastoral ecosystem. J. Agric. Sci., Camb. 107: 357–65
Coughenour M. B. 1991. Spatial components of plant herbivore interactions in pastoral, ranching and natural ungulate ecosystems. J. Range Mgmt 44: 530–42
Coughenour M. B. & Singer F. J.. 1996. Elk population processes in Yellowstone National Park under the policy of natural regulation. Ecol. Appl. 6: 573–93
Crawley, M. J. 1983. Herbivory: The Dynamics of Animal-Plant Interactions. (Studies in Ecology Series No. 10.) Blackwell, Oxford
Danckwerts J. E., Aucamp A. J. & Barnard H. J.. 1983. Herbaceous species preference by cattle in the false thornveld of eastern Cape. Proc. Grass. Soc. Sth. Afr. 18: 89–94
Dean W. R. J. & Macdonald I. A. W.. 1994. Historical changes in stocking rates of domestic livestock as a measure of semi-arid and arid rangeland degradation in the Cape Province, South Africa. J. Arid Envir. 26: 281–98
DeAngelis, D. L. 1992. Dynamics of Nutrient Cycling and Food Webs. Chapman & Hall, London
DeAngelis D. L., Goldstein R. A. & O'Neil R. V.. 1975. A model for trophic interaction. Ecology 56: 881–92
DeAngelis, D. L. & L. J. Gross (eds). 1992. Individual-Based Models and Approaches in Ecology. Chapman & Hall, London
Detling J. K., Parton W. J. & Hunt H. W.. 1979. A simulation model of Bouteloua gracilis biomass dynamics on the North American shortgrass prairie. Oecologia 38: 167–91
de Vos, V., R. C. Bengis & H. J. Coetzee. 1983. Population control of large mammals in the Kruger National Park. In Management of Large Mammals in African Conservation Areas, ed. R. N. Owen-Smith, pp. 213–31. Haum, Pretoria
Dinius D. A. & Baumgardt B. R.. 1970. Regulation of food intake in ruminants: 6. Influence of caloric density of pelleted rations. J. Dairy Sci. 53: 311–16
Distel R. A., Laca E. A., Griggs T. C. & Demment M. W.. 1995. Patch selection by cattle: maximization of intake rate in horizontally heterogeneous pastures. Appl. Anim. Behav. Sci. 45: 11–21
Duncan, P. 1975. Topi and their food supply. PhD thesis, University of Nairobi
Duncan P., Foose T. J., Gordon I. J., Gakahu C. G. & Lloyd M.. 1990. Comparative nutrient extraction from forages by bovids and equids: a test of the nutritional model of equid/bovid competition and coexistence. Oecologia 84: 411–18
Dunham K. M. 1982. The foraging behaviour of impala. S. Afr. J. Wildl. Res. 12: 36–40
Dunham K. M. & Murray M. G.. 1982. The fat reserves of impala. Afr. J. Ecol. 20: 81–7
du Toit, J. T. 1988. Patterns of resource use within the browsing ruminant guild in the central Kruger National Park. PhD thesis, University of the Witwatersrand
Toit J. T. & Owen-Smith N.. 1989. Body size, population metabolism and habitat specialization among African large herbivores. Am. Nat. 133: 736–40
Dyer M. I., Turner C. L. & Seastedt T. R.. 1993. Herbivory and its consequences. Ecol. Applic. 3: 10–16
Edwards G. R., Parsons A. J., Penning P. D. & Newman J. A.. 1995. Relationship between vegetation state and bite dimensions of sheep grazing contrasting plant species and its implications for intake rate and diet selection. Grass Forage Sci. 50: 378–88
Ellis J. E., & Swift D. M.. 1988. Stability of African pastoral ecosystems: alternative paradigms and implications for development. J. Range Mgmt 41: 450–9
Elser J. J., Sterner R. W., Gorokhova E., Fagan W. F., Markow T. A., Cotner J. B., Harrison J. F., Hobbie S. E., Odell G. M. & Weides L. J.. 2000. Biological stoichiometry from genes to ecosystems. Ecol. Lett. 3: 540–50
Elton, C. 1927. Animal Ecology. Sidgwick & Jackson, London
Eltringham S. K. 1974. Changes in the large mammal community of Mweya Peninsula, Rwenzori National Park, following removal of hippopotamus. J. Appl. Ecol. 11: 855–65
Eltringham S. K. 1980. A quantitative assessment of range usage by large African mammals with particular reference to the effects of elephants on trees. Afr. J. Ecol. 18: 53–71
Farnsworth K. D. & Illius A. W.. 1996. Large grazers back in the fold: generalizing the prey model to incorporate mammalian herbivores. Funct. Ecol. 10: 678–80
Farnsworth K. D. & Illius A. W.. 1998. Optimal diet choice for large herbivores: an extended contingency model. Funct. Ecol. 12: 74–81
Ferrar A. A. & Walker B. H.. 1974. An analysis of herbivore/habitat relationships in Kyle National Park, Rhodesia. J. S. Afr. Wildl. Mgmt Ass. 4: 137–47
Finch V. A. 1972. Thermoregulation and heat balance of the East African eland and hartebeest. Am. J. Physiol. 222: 1374–9
FitzGibbon C. D. & Fanshawe J. H.. 1988. Stotting in Thomson's gazelles: an honest signal of condition. Behav. Ecol. Sociobiol. 23: 69–74
Forbes T. D. A. 1988. Researching the plant-animal interface: the investigation of ingestive behaviour in grazing animals. J. Anim. Sci. 66: 2369–79
Fowler C. W. 1981. Density dependence as related to life history strategy. Ecology 62: 602–10
Fowler, C. W. 1987. A review of density dependence in populations of large mammals. In Current Mammalogy, Vol. 1, ed. H. H. Genoways, pp. 401–41. Plenum Press, New York
Freeland W. J. & Janzen D. H.. 1974. Strategies in herbivory by mammals: the role of plant secondary compounds. Am. Nat. 108: 269–89
Fretwell, S. D. 1972. Populations in a Seasonal Environment. Princeton University Press, Princeton, NJ
Fritz H. & Garine-Witchatitsky M.. 1996. Foraging in a social antelope: effects of group size on foraging choices and resource perception in impala. J. Anim. Ecol. 65: 736–42
Fritz H., Garine-Witchatitsky M. & Letessier G.. 1996. Habitat use by sympatric wild and domestic herbivores in an African savanna woodland: the influence of cattle spatial behaviour. J. Appl. Ecol. 33: 589–98
Fritz H. & Duncan P.. 1994. On the carrying capacity for large ungulates of African savanna ecosystems. Proc. R. Soc. Lond. B 256: 77–82
Fryxell J. M. 1991. Forage quality and aggregation by large herbivores. Am. Nat. 138: 478–98
Fryxell J. M., Greever J. & Sinclair A. R. E.. 1988. Why are migratory ungulates so abundant?Am. Nat. 131: 781–98
Fryxell J. M. & Lundberg P.. 1994. Diet choice and predator prey dynamics. Evol. Ecol. 8: 407–21
Fryxell, J. M. & P. Lundberg. 1998. Individual Behavior and Community Dynamics. Chapman & Hall, London
Futuyma, D. J. 1979. Evolutionary Biology. Sinauer Associates, Sunderland, MA
Gaillard J. M., Festa-Bianchet M., Yoccoz N. G. & Loison A.. 2000. Temporal variation in vital rates and population dynamics of large herbivores. A. Rev. Ecol. Syst. 31: 367–93
Gallivan G. I., Culverwell J. & Girdwood R.. 1995. Body condition indices of impala: effect of age class, sex, season and management. S. Afr. J. Wildl. Res. 25: 23–31
Garten C. T. Jr., 1978. Multivariate perspectives on the ecology of plant mineral element composition. Am. Nat. 112: 533–44
Geist V. 1974. On the relationship of social evolution and ecology in ungulates. Am. Zool. 14: 205–20
Gerhart K. L., White R. G., Cameron R. D. & Russell D. E.. 1996. Body composition and nutrient reserves of arctic caribou. Can. J. Zool. 74: 136–46
Getz W. M. 1984. Population dynamics: a per-capita resource approach. J. Theor. Biol. 108: 623–43
Getz W. M. 1991. A unified approach to multispecies modelling. Natur. Res. Model. 5: 393–421
Getz W. M. 1993. Metaphysiological and evolutionary dynamics of populations exploiting constant and interactive resources: r-K selection revisited. Evol. Ecol. 7: 287–305
Getz, W. M. 1994. A metaphysiological approach to modelling ecological populations and communities. In Frontiers in Mathematical Biology, ed. S. A. Levin, pp. 411–42. Springer-Verlag, Berlin
Getz W. M. & Owen-Smith N.. 1999. A metaphysiological population model of storage in variable environments. Natur. Res. Model. 12: 197–230
Giesecke D. & Gylswyk N. O.. 1975. A study of feeding types and certain rumen functions in six species of South African wild ruminants. J. Agric. Sci., Camb. 85: 75–83
Gillard P. & Monypenny R.. 1990. A decision support model to evaluate the effects of drought and stocking rate on beef cattle properties in Northern Australia. Agric. Syst. 34: 37–52
Gillingham M. P., Parker K. L. & Hanley T. A.. 1997. Forage intake by black tailed deer in a natural environment: bout dynamics. Can. J. Zool. 75: 1118–28
Ginnett T. F., Dankosky J. A., Deo G. & Demment M. W.. 1999. Patch depression in grazers: the roles of biomass distribution and residual stems. Funct. Ecol. 13: 37–44
Ginnett T. F. & Demment M. W.. 1995. The functional response of herbivores: analysis and test of a simple mechanistic model. Funct. Ecol. 9: 376–84
Ginnett T. F. & Demment M. W.. 1997. Sex differences in giraffe foraging behaviour at two scales. Oecologia 110: 291–300
Ginzburg L. R. 1998. Assuming reproduction to be a function of consumption raises doubts about some popular predator-prey models. J. Anim. Ecol. 67: 325–7
Gleeson S. K. 1995. Density dependence is better than ratio dependence. Ecology 75: 1834–5
Goddard J. 1968. Food preferences of two black rhinoceros populations. E. Afr. Wildl. J. 6: 1–18
Gordon I. J. 1988. Facilitation of red deer grazing by cattle and its impact on red deer performance. J. Appl. Ecol. 25: 1–10
Gordon I. J. & Illius A. W.. 1988. Incisor arcade structure and diet selection in ruminants. Funct. Ecol. 2: 15–22
Gordon I. J. & Illius A. W.. 1994. The functional significance of the browser-grazer dichotomy in African ruminants. Oecologia 98: 167–75
Gotelli, N. J. 1995. A Primer of Ecology. Sinauer, Sunderland, MA
Greenwood G. B. & Demment M. W.. 1988. The effect of fasting on short-term cattle grazing behaviour. Grass Forage Sci. 43: 377–86
Grenfell B. T. 1992. Parasitism and the dynamics of grazing systems. Am. Nat. 139: 907–29
Grenfell B. T., Price O. F., Albon S. D. & Clutton-Brock T. H.. 1992. Overcompensation and population cycles in an ungulate. Nature 355: 823–6
Grenfell B. T., Wilson K., Finkenstadt B. F., Coulson T. N., Murray S., Albon S. D., Pemberton J. M., Clutton-Brock T. H. & Crawley M. J.. 1998. Noise and determinism in synchronized sheep dynamics. Nature 394: 674–7
Grime J. P. 1977. Evidence for the existence of three primary strategies in plants and its relevance to ecological and evolutionary theory. Am. Nat. 111: 1169–94
Gross J. E., Alkon O. V. & Demment M. W.. 1996. Nutritional ecology of dimorphic herbivores: digestion and passage rates in Nubian ibex. Oecologia 102: 170–8
Grossman, D., P. L. Holden & R. F. H. Collinson. 1999. Veld management on the game ranch. In Veld Management in South Africa, ed. N. M. Tainton, pp. 261–79. University of Natal Press, Pietermaritzburg
Grunow J. O., Groeneveld H. T. & Toit H. C.. 1980. Above ground dry matter dynamics of the grass layer in the Nylsvley tree savanna. J. Ecol. 68: 877–89
Gulland F. M. D. 1992. Role of nematode infections in mortality of Soay sheep during a population crash on St. Kilda. Parasitology 105: 493–503
Gurney, W. S. C. & R. M. Nisbet. 1998. Ecological Dynamics. Oxford University Press, Oxford
Gustafson L. L., Franklin W. L., Sarno R. J., Hunter R. L., Young K. M., Johnson W. E. & Behl M. J.. 1998. Predicting early mortality of newborn guanaco by birth mass and haematological parameters: a provisional model. J. Wildl. Mgmt 62: 24–35
Hagerman A. E., Robbins C. T., Weerasuriya Y., Wilson T. C. & McArthur C.. 1992. Tannin chemistry in relation to digestion. J. Range Mgmt 45: 57–62
Hairston N. G., Smith F. E. & Slobodkin L. B.. 1960. Community structure, population control and competition. Am. Nat. 94: 421–5
Hamilton E. W., Giovannini M. S., Moses S. A., Coleman J. S. & McNaughton S. J.. 1998. Biomass and mineral element responses of a Serengeti short-grass species to nitrogen supply and defoliation: compensation requires a critical [N]. Oecologia 116: 407–18
Hammond K. A. & Diamond J.. 1997. Maximal sustained energy budgets in humans and animals. Nature 386: 457–62
Hanley T. A. 1997. A nutritional view of understanding and complexity in the problem of diet selection by deer (Cervidae). Oikos 79: 209–18
Hanski J. 1989. Metapopulation dynamics: does it help to have more of the same?Trends Ecol. Evol. 4: 113–14
Hanski, J. 1999. Metapopulation Ecology. Oxford University Press
Hanson J. D., Parton W. J. & Innis G. S.. 1985. Plant growth and production of grassland ecosystems: a comparison of modelling approaches. Ecol. Model. 29: 131–44
Hansson, L., L. Fahrig & G. Merriam (eds). 1995. Mosaic Landscapes and Ecological Processes. Chapman & Hall, London
Hardy, M. B., C. R. Hurt & O. J. H. Bosch. 1999. Veld condition assessment. Grassveld. In Veld Management in South Africa, ed. N. M. Tainton, pp. 194–206. University of Natal Press, Pietermaritzburg
Harper, J. L. 1977. Population Biology of Plants. Academic Press, London and New York
Hassell M. P., Lawton J. H. & May R. M.. 1976. Patterns of dynamical behaviour in single-species populations. J. Anim. Ecol. 45: 471–86
Hastings, A. 1997. Population Biology. Concepts and Models. Springer-Verlag, New York
Hatch G. P. & Stafford Smith D. M.. 1997. The bioeconomic implications of various drought management strategies for a communal cattle herd in a semi-arid savanna of KwaZulu-Natal. Afr. J. Range Forage Sci. 14: 17–25
Heady, H. F. & R. D. Child. 1994. Rangeland Ecology and Management. Westview Press, Boulder, CO
Heitschmidt, R. K. & J. W. Stuth (eds). 1991. Grazing Management: An Ecological Perspective. Timber Press, Portland, OR
Herbers J. M. 1981. Time resources and laziness in animals. Oecologia 49: 252–62
Hewison A. J. M., Angibault J. M., Boutin J., Bideau E., Vincent J. P. & Sempere A.. 1996. Annual variation in body composition of roe deer in moderate environmental conditions. Can. J. Zool. 74: 245–53
Hik D. S. 1995. Does risk of predation influence population dynamics? Evidence from the cyclic decline of snowshoe hares. Wildl. Res. 22: 115–29
Hirakawa H. 1997. Digestion — constrained optimal foraging in generalist mammalian herbivores. Oikos 78: 37–47
Hobbs, N. T. 1990. Diet selection by generalist herbivores: a test of the linear programming model. In Behavioural Mechanisms of Food Selection, ed. R. N. Hughes, pp. 395–413. Springer-Verlag, Heidelberg
Hobbs N. T., Baker D. L., Bear G. D. & Bowden D. C.. 1996a. Ungulate grazing in sagebrush grassland: mechanisms of resource competition. Ecol. Applic. 6: 200–17
Hobbs N. T., Baker D. L., Bear G. D. & Bowden D. C.. 1996b. Ungulate grazing in sagebrush grassland: effects of resource competition on secondary production. Ecol. Applic. 6: 218–27
Hodgson, J. & A. W. Illius. 1996. The Ecology and Management of Grazing Systems. CAB International, Wallingford
Hofmann, R. R. 1973. The Ruminant Stomach. (East Afr. Monogr. Biol. Vol. 2.) East African Literature Bureau, Nairobi
Hofmann R. R. 1989. Evolutionary steps of ecophysiological adaptation and diversification of ruminants: A comparative review of their digestive system. Oecologia 78: 443–57
Holland O. 1992. Seasonal variation in body composition of European roe deer. Can. J. Zool. 70: 502–4
Holling C. S. 1959. Some characteristics of simple types of predation and parasitism. Can. Entomol. 91: 385–98
Holling C. S. 1965. The functional response of predators to prey density and its role in mimicry and population regulation. Mem. Entomol. Soc. Can. 45: 1–60
Houston, A. J. & J. M. McNamara. 1999. Models of Adaptive Behaviour. An Approach Based on State. Cambridge University Press, Cambridge
Houston A. J., Welton N. J. & McNamara J. M.. 1997. Acquisition and maintenance costs in the long-term regulation of avian fat reserves. Oikos 78: 331–40
Hudson R. J. & Frank S.. 1987. Foraging ecology of bison in aspen boreal habitats. J. Range Mgmt 40: 71–5
Hudson R. J. & Watkins W. G.. 1986. Foraging rates of wapiti on green and cured pastures. Can. J. Zool. 64: 1705–8
Huff D. E. & Varley J. D.. 1999. Natural regulation in Yellowstone National Park's northern range. Ecol. Applic. 9: 17–29
Hungate R. E. 1975. The rumen microbial ecosystem. A. Rev. Ecol. Syst. 6: 39–66
Hunter, M. D., T. Ohgushi & P. W. Price. 1992. Effects of Resource Distribution on Animal-Plant Interactions. Academic Press, London and New York
Hunter M. D. & Price P. W.. 1992. Playing chutes and ladders: heterogeneity and the relative roles of bottom-up and top-down forces in natural communities. Ecology 73: 724–32
Hutchinson G. E. 1958. Concluding remarks. Cold Spring Harbor Symp. Quant. Biol. 22: 415–27
Hutchinson G. E. 1959. Homage to Santa Rosalia, or why are there so many kinds of animals?Am. Nat. 93: 145–59
Illius A. W. & Gordon I. J.. 1987. The allometry of food intake in grazing ruminants. J. Anim. Ecol. 56: 989–99
Illius A. W. & Gordon I. J.. 1991. Prediction of intake and digestion in ruminants by a model of rumen kinetics integrating animal size and plant characteristics. J. Agric. Sci., Cambr. 116: 145–57
Illius A. W. & Gordon I. J.. 1992. Modelling the nutritional ecology of ungulate herbivores: evolution of body size and competitive ability. Oecologia 89: 428–34
Illius, A. W. & I. J. Gordon. 1998. Scaling up from functional response to numerical response in vertebrate herbivores. In Herbivores, Plants and Predators, ed. H. Olff, V. K. Brown & R. T. Drent, pp. 397–427. Blackwell Science, Oxford
Illius A. W., Gordon I. J., Elston D. A. & Milne J. D.. 1999. Diet selection in goats: a test of intake-rate maximization. Ecology 80: 1008–18
Illius A. W. & Jessop N. S.. 1995. Modelling metabolic costs of allelochemical ingestion by foraging ungulates. J. Chem. Ecol. 21: 693–719
Illius A. W. & Jessop N. S.. 1996. Metabolic constraints on voluntary intake in ruminants. J. Anim. Sci. 74: 3052–62
Illius A. W. & O'Connor T. G.. 1999. On the relevance of nonequilibrium concepts to arid and semi-arid grazing systems. Ecol. Applic. 9: 798–813
Illius A. W. & O'Connor T. G.. 2000. Resource heterogeneity and ungulate population dynamics. Oikos 89: 283–94
Irby L. R. 1977. Food habits of Chandler's mountain reedbuck in a Rift Valley ranch. E. Afr. Wildl. J. 15: 289–94
Ivlev, V. S. 1961. Experimental Ecology of the Feeding of Fishes. Yale University Press, New Haven, MA
James C. D., Landsberg J. & Morton S. R.. 1999. Provision of watering points in the Australian Arid zone: a review of effects on biota. J. Arid Envir. 41: 87–121
Janis C. 1976. The evolutionary strategy of the Equidae and the origins of rumen and cecal digestion. Evolution 30: 757–76
Jarman P. J. 1971. Diets of large mammals in the woodlands around Lake Kariba. Oecologia 8: 157–78
Jarman P. J. 1974. The social organization of antelope in relation to their ecology. Behaviour 48: 215–67
Jeschke J. M., Kopp M. & Tollrian R.. 2001. Predator functional responses: discriminating between handling and digesting prey. Ecology. (In press.)
Jiang Z. & Hudson R. J.. 1992. Estimating forage intake and energy requirements of free-ranging wapiti. Can. J. Zool. 70: 675–79
Jiang Z. & Hudson R. J.. 1993. Optimal grazing of wapiti on grassland: patch and feeding station departure rules. Evol. Ecol. 7: 488–98
Jiang Z. & Hudson R. J.. 1994. Bite characteristics of wapiti in seasonal Bromus-Poa swards. J. Range Mgmt 47: 127–32
Jones R. J. & Sandland R. L.. 1974. The relation between animal gain and stocking rate: derivation of the relation from the results of grazing trials. J. Agric. Sci., Cambr. 83: 335–41
Joos-Vandewalle, M. E. 2000. Movement of migratory zebra and wildebeest in northern Botswana. PhD thesis, University of the Witwatersrand
Jorgenson J. T., Festa-Bianchet M., Gaillard J.-M. & Wishart W. D.. 1997. Effects of age, sex, disease and density on survival of bighorn sheep. Ecology 78: 1019–32
Joubert D. M. 1954. The influence of winter nutritional depressions on the growth, reproduction and production of cattle. J. Agric. Sci., Cambr. 44: 5–66
Judson O. P. 1994. The rise of the individual based model in ecology. Trends Ecol. Evol. 9: 9–14
Kaufmann, S. A. 1993. The Origins of Order: Self-Organization and Selection in Evolution. Oxford University Press, Oxford
Kaufmann, S. A. 1995. At Home in the Universe. The Search for the Laws of Complexity. Oxford University Press, New York
Kemeny, J. G. & T. E. Kurtz. 1990. True BASIC. A Powerful Structured Language System by the Original Authors of BASIC. True BASIC Inc., West Lebanon, NH
Klein D. R. 1968. The introduction, increase and crash of reindeer on St. Matthew Island. J. Wildl. Mgmt 32: 350–67
Kooijman, S. A. L. M. 1990. Biomass conversion at the population level. In Individual-Based Models and Approaches in Ecology, ed. D. L. DeAngelis & L. J. Gross, pp. 338–58. Chapman & Hall, New York and London
Kooijman, S. A. L. M. 1993. Dynamic Energy Budgets in Biological Systems. Cambridge University Press, Cambridge
Krebs, J. R. & R. H. McLeery. 1984. Optimization in behavioural ecology. In Behavioural Ecology. An Evolutionary Approach, ed. J. R. Krebs & N. B. Davies, 2nd edn., pp. 91–21. Blackwell, Oxford
Krebs, C. J., A. R. E. Sinclair, R. Boonstra, S. Boutin, K. Martin & J. N. M. Smith. 1998. Community dynamics of vertebrate herbivores: how can we untangle the web? In Herbivores: Between Plants and Predators, ed. H. Olff, V. K. Brown & R. H. Drent, pp. 447–73. Blackwell Science, Oxford
Kruuk, H. 1972. The Spotted Hyena. University of Chicago Press, Chicago
Laca, E. A. & M. W. Demment. 1991. Herbivory: the dilemma of foraging in a spatially heterogeneous food environment. In Plant Defences Against Mammalian Herbivory, ed. R. T. Palo & C. T. Robbins. pp. 29–44. CRC Press, Boca Raton, FL
Laca, E. A. & M. W. Demment. 1993. Modelling intake of a grazing ruminant in a heterogeneous environment. In Proceedings of the International Symposium on Vegetation-Herbivore Relationships, ed. T. Okubo, B. Hubest & G. Arnold, pp. 57–76. Academic Press, London
Laca, E. A. & M. W. Demment. 1996. Foraging strategies of grazing animals. In The Ecology and Management of Grazing Systems, ed. J. Hodgson & A. W. Illius, pp. 137–58. CAB International, Wallingford, UK
Laca E. A., Distel R. A., Griggs T. C. & Demment M. W.. 1994a. Effects of canopy structure on patch depression by grazers. Ecology 75: 706–16
Laca, E. A., R. A. Distel, T. C. Griggs, G. Deo & M. W. Demment. 1993. Field test of optimal foraging with cattle: the marginal value theorem successfully predicts patch selection and utilisation. Proc. XVII Int. Grassl. Congr., pp. 709–10
Laca E. A., Ungar E. D. & Demment M. W.. 1994b. Mechanism of handling time and intake rate of a large mammalian grazer. Appl. Anim. Behav. Sci. 39: 3–19
Laca E. A., Ungar E. D., Seligman N. & Demment M. W.. 1992. Effects of sward height and bulk density on bite dimensions of cattle grazing homogeneous swards. Grass Forage Sci. 47: 91–102
Lange R. T. 1969. The piosphere: sheep track and dung patterns. J. Range Mgmt 22: 396–400
Langvatn R. & Hanley T. A.. 1993. Feeding patch choice by red deer in relation to foraging efficiency. Oecologia 95: 164–70
Laws R. M. & Parker I. S. C.. 1968. Recent studies on elephant populations in East Africa. Symp. Zool. Soc., Lond. No. 21, pp. 319–59
Lechner-Doll M., Rutagwenda T., Schwartz H. J., Schultka W. & Engelhardt W. v.. 1990. Seasonal changes of ingestion mean retention time and forestomach fluid volume in indigenous camels, cattle, sheep and goats grazing a thornbush savannah pasture in Kenya. J. Agric. Sci., Cambr. 115: 409–20
Ledger, H. P. 1968. Body composition as a basis for a comparative study of some East African mammals. In Comparative Nutrition of Wild Animals, ed. M. A. Crawford (Symp. Zool. Soc., Lond. No. 21), pp. 289–310
Leirs H., Stenseth N. C., Nicholl J. D., Hines J. E., Verhagen R. & Verhagen W.. 1997. Stochastic seasonality and nonlinear density-dependent factors regulate population size in an African rodent. Nature 389: 176–80
Lemaire, G. & D. Chapman. 1996. Tissue flows in grazed plant communities. In The Ecology and Management of Grazing Systems, ed. J. Hodgson & A. W. Illius, pp. 3–36. CAB International, Wallingford, UK
Loison A., Festa-Bianchet M., Gaillard J.-M., Jorgenson J. T. & Jullien J.-M.. 1999. Age-specific survival in five populations of ungulates: evidence of senescence. Ecology 80: 2539–54
Lotka, A. J. 1925. Elements of Mathematical Biology. Dover, New York
Lott, D. F. 1991. Intraspecific Variation in the Social Systems of Wild Vertebrates. Cambridge University Press, Cambridge
Low W. A., Dudzinski M. L. & Miller W. J.. 1981. The influence of forage and climatic conditions on range community preferences of shorthorn cattle in central Australia. J. Appl. Ecol. 18: 11–26
Ludwig D., Hilborn R. & Walters C.. 1993. Uncertainty, resource exploitation, and conservation: lessons from history. Science 260: 17–36
Mangel, M. & C. W. Clark. 1988. Dynamic Modelling in Behavioral Ecology. Princeton University Press, Princeton, NJ
Manser M. B. & Brotherton P. N. M.. 1995. Environmental constraints on the foraging behaviour of a dwarf antelope. Oecologia 102: 404–12
Martin, R. B., G. C. Craig & V. R. Booth. 1989. Elephant Management in Zimbabwe. Department of National Parks & Wild Life Management, Harare, Zimbabwe
May, R. M. 1973. Stability and Complexity in Model Ecosystems. Princeton University Press, Princeton, NJ
May, R. M. 1981. Models for single populations. In Theoretical Ecology, ed. R. M. May, pp. 5–29. Blackwell, Oxford
May R. & Oster G.. 1976. Bifurcations and dynamic complexity in simple ecological models. Am. Nat. 110: 573–99
Maynard Smith J. & Slatkin M.. 1973. The stability of predator-prey systems. Ecology 54: 384–91
McCabe J. T. 1987. Drought and recovery: Livestock dynamics among the Ngisonyoka Turkana of Kenya. Human Ecol. 15: 317–29
McCullough, D. R. 1992. Concepts of large herbivore population dynamics. In Wildlife 2001. Populations, ed. D. R. McCullough & R. H. Barrett, pp. 967–84. Elsevier, London
McCullough, D. R. 1997. Irruptive behavior in ungulates. In The Science of Overabundance, Deer Ecology and Population Management, ed. W. J. McShea, H. B. Underwood & J. H. Rappole, pp. 69–99. Smithsonian Institution Press, Washington, DC
McLeod S. R. 1997. Is the concept of carrying capacity useful in variable environments?Oikos 79: 529–42
McNamara J. M. & Houston A. I.. 1987a. Partial preferences and foraging. Anim. Behav. 35: 1084–99
McNamara J. M. & Houston A. I.. 1987b. Starvation and predation as factors limiting population size. Ecology 68: 1515–19
McNamara J. M. & Houston A. I.. 1990. The value of fat reserves and the trade-off between starvation and predation. Acta Biotheor. 38: 37–61
McNamara J. M. & Houston A. I.. 1996. State-dependent life histories. Nature 380: 215–21
McNaughton S. J. 1976. Serengeti migratory wildebeest: facilitation of energy flow by grazing. Science 191: 92–3
McNaughton S. J. 1979a. Grazing as an optimization process: grass-ungulate relationships in the Serengeti. Am. Nat. 113: 691–703
McNaughton, S. J. 1979b. Grassland herbivore dynamics. In Serengeti: Dynamics of an Ecosystem, ed. A. R. E. Sinclair & M. Norton-Griffiths, pp. 46–81. University of Chicago Press, Chicago, IL
McNaughton S. J. 1983. Compensatory plant growth as a response to herbivory. Oikos 40: 329–36
McNaughton S. J. 1984. Grazing lawns: animals in herds, plant form and coevolution. Am. Nat. 124: 863–86
McNaughton S. J., Banyikwa F. F. & McNaughton M. M.. 1997. Promotion of the cycling of diet-enhancing nutrients by African grazers. Science 278: 1798–800
McNaughton S. J., Banyikwa F. F. & McNaughton M. M.. 1998. Root biomass and productivity in a grazing ecosystem: the Serengeti. Ecology 79: 587–92
Mduma S. A. R., Sinclair A. R. E. & Hilborn R.. 1999. Food regulates the Serengeti wildebeest: a 40-year record. J. Anim. Ecol. 68: 1101–22
Meissner H. H. & Paulsmeier D. V.. 1995. Plant composition affecting between-plant and animal species prediction of forage intake. J. Anim. Sci. 73: 2447–57
Meissner, H. H., P. J. K. Zacharias & P. J. O'Reagain. 1999. Forage quality (feed value). In Veld Management in South Africa, ed. N. M. Tainton, pp. 139–68. University of Natal Press, Pietermaritzburg
Mentis, M. T. 1981. Acceptability and palatability. In Veld and Pasture Management in South Africa, ed. N. M. Tainton, pp. 186–91. Shuter & Shooter, Pietermaritzburg
Mertens D. R. 1977. Dietary fiber components: relationships to the rate and extent of ruminal digestion. Fedn Am. Socs. Exp. Biol. 36: 187–92
Mertens D. R. 1987. Predicting intake and digestibility using mathematical models of rumen function. J. Anim. Sci. 64: 1548–58
Messier F. 1994. Ungulate population models with predation: a case study with North American moose. Ecology 75: 478–88
Messier F. 1995. Trophic interaction in two northern wolf-ungulate systems. Wildl. Res. 22: 131–46
Metz, J. A. J. & O. Dieckmann. 1986. The Dynamics of Physiologically Structured Populations. Springer-Verlag, New York
Milinski, M. & G. Parker. 1991. Competition for resources. In Behavioural Ecology. An Evolutionary Approach, 3rd edn, ed. J. R. Krebs & N. B. Davies, pp. 137–68. Blackwell, Oxford
Miller M. F. 1996. Dispersal of Acacia seeds by ungulates and ostriches in an African savanna. J. Trop. Ecol. 12: 345–56
Mills M. G. L., Biggs H. C. & Whyte I. J.. 1995. The relationship between rainfall, lion predation and population trends in African herbivores. Wildl. Res. 22: 75–88
Milne J. A., MacRae J. C., Spence A. M. & Wilson S.. 1976. Intake and digestion of hill-land vegetation by the red deer and the sheep. Nature 263: 763–4
Milner, C. & D. Gwynne. 1974. The Soay sheep and their food supply. In Island Survivors. The Ecology of the Soay Sheep of St. Kilda, ed. P. A. Jewell, C. Milner & J. Morton Boyd, pp. 273–325. Athlone Press, University of London
Milner J. M., Albon S. D., Illius A. W., Pemberton J. M. & Clutton-Brock T. H.. 1999a. Repeated selection of morphometric traits in the Soay sheep on St. Kilda. J. Anim. Ecol. 68: 472–88
Milner J. M., Elston D. A. & Albon S. D.. 1999b. Estimating the contributions of population density and climatic fluctuations to interannual variation in survival of Soay sheep. J. Anim. Ecol. 68: 1235–47
Milner-Gulland, E. J. & R. Mace. 1998. Conservation of Biological Resources. Blackwell Science, Oxford
Moen, A. N. 1973. Wildlife Ecology. Freeman, San Francisco
Molvar E. M., Bowyer R. T. & Ballenbergh V.. 1993. Moose herbivory, browse quality and nutrient cycling in an Alaskan treeline community. Oecologia 94: 472–9
Morton Boyd, J. 1974. Introduction. In Island Survivors. The Ecology of the Soay Sheep of St Kilda, ed. P. A. Jewell, C. Milner & J. Morton Boyd, pp. 1–7. Athlone Press, University of London
Mrosovsky N. & Prowley T. L.. 1977. Set points for body weight and fat. Behav. Biol. 20: 205–23
Murray M. G. 1993. Comparative nutrition of wildebeest, hartebeest and topi in the Serengeti. Afr. J. Ecol. 31: 172–7
Murray M. G. & Brown D.. 1993. Niche separation of grazing ungulates in the Serengeti: an experimental test. J. Anim. Ecol. 62: 380–9
Murray, M. G. & A. W. Illius. 1996. Multispecies grazing in the Serengeti. In The Ecology and Management of Grazing Systems, ed. J. Hodgson & A. W. Illius, pp. 247–72. CAB International, Wallingford, UK
Murray M. G. & Illius A. W.. 2000. Vegetation modification and resource competition in grazing ungulates. Oikos 89: 501–8
Newman J. A., Parsons A. J. & Harvey A.. 1993. Not all sheep prefer clover: diet selection revisited. J. Agric. Sci., Cambr. 119: 275–83
Nicolis G. & I. Prigogine. 1977. Self-organization in Non-equilibrium Systems. Wiley, New York
Novellie P. A. 1978. Comparison of the foraging strategies of blesbok and springbok on the Transvaal highveld. S. Afr. J. Wildl. Res. 8: 137–44
Novellie P. 1990. Habitat use by indigenous grazing ungulates in relation to sward structure and veld condition. J. Grassl. Soc. Sth. Afr. 7: 16–23
Noy-Meir I. 1975. Stability of grazing systems: an application of predator-prey graphs. J. Ecol. 63: 459–81
Noy-Meir I. 1978. Grazing and production in seasonal pastures: analysis of a simple model. J. Appl. Ecol. 15: 809–35
O'Connor T. G. 1991. Local extinction in perennial grasslands: a life history approach. Am. Nat. 137: 753–73
Oftedal, O. T. 1984. Milk composition, milk yield and energy output at peak lactation. In Lactation Strategies. (Symp. Zool. Soc. Lond. No. 51), pp. 33–85
Oksanen L., Moen J. & Lundberg P. A.. 1992. The time-scale problem in exploiter-victim models: does the solution lie in ratio-dependent exploitation?Am. Nat. 140: 938–60
Oliver M. D. N., Short N. R. M. & Hanks J.. 1978. Population ecology of oribi, grey rhebuck and mountain reedbuck in Highmoor State Forest Land, Natal. S. Afr. J. Wildl. Res. 8: 95–105
O'Reagain P. J. 1993. Plant structure and the acceptability of different grasses to sheep. J. Range Mgmt 46: 232–6
O'Reagain, P. J. 1994. The effect of sward structure and species composition on dietary quality and intake in cattle and sheep grazing the Dohne sourveld. PhD thesis, University of the Witwatersrand
O'Reagain, P. J. 2001. Foraging strategies on rangelands: Effects of intake and animal performance. In Proceedings of the XIX International Grassland Congress, Sao Pedro, Brazil, Feb. 2001, ed. J. A. Gomide, W. R. S. Mattos and S. C. da Silva, pp. 277–84. Brazilian Society of Animal Production, Sao Paulo, Brazil
O'Reagain P. J., Goetsch B. C. & Owen-Smith R. N.. 1995. Ruminal degradation characteristics of some African rangeland grasses. J. Agric. Sci., Cambr. 125: 189–97
O'Reagain P. J., Goetsch B. C. & Owen-Smith R. N.. 1996. Effect of species composition and sward structure on the ingestive behaviour of cattle and sheep grazing South African sourveld. J. Agric. Sci., Cambr. 127: 271–80
O'Reagain P. J. & Grau E. R.. 1995. Sequence of species selection by cattle and sheep on South African sourveld. J. Range Mgmt 48: 314–21
O'Reagain P. J. & Mentis M. T.. 1989a. Sequence and process of species selection by cattle in relation to optimal foraging theory in an old land in the Natal Sour Sandveld. J. Grassl. Soc. Sth. Afr. 6: 71–6
O'Reagain P. J. & Mentis M. T.. 1989b. The effect of plant structure on the acceptability of different grass species to cattle. J. Grassl. Soc. Sth. Afr. 6: 163–70
O'Reagain P. J. & Mentis M. J.. 1990. The effects of veld condition on the quality of diet selected by cattle grazing the Natal Sour Sandveld. J. Grassl. Soc. Sth. Afr. 7: 190–5
O'Reagain P. J. & Owen-Smith R. N.. 1996. Effect of species composition and sward structure on dietary quality in cattle and sheep grazing South Africa sourveld. J. Agric. Sci., Cambr. 127: 261–70
O'Reagain, P. J. & J. Schwartz. 1995. Dietary selection and foraging strategies of animals on rangeland. Coping with spatial and temporal variability. In Recent Developments in the Nutrition of Herbivores (Proceedings of IV International Symposium on the Nutrition of Herbivores), ed. M. Journet, E. Grenet, M.-H. Farce, M. Thierez & C. Demarquilly, pp. 407–23. INRA Editions, Paris
Orians, G. H. & N. E. Pearson. 1979. On the theory of central place foraging. In Analysis of Ecological Systems, ed. D. J. Horn, R. D. Mitchell & G. R. Stairs, pp. 154–77. Ohio State University Press, Columbus
Owen-Smith, N. 1973. The behavioural ecology of the white rhinoceros. PhD thesis, University of Wisconsin
Owen-Smith, N. 1974. The social system of the white rhinoceros. In The Behaviour of Ungulates and Its Relation to Management (IUCN Public., NS No. 24) ed. V. Geist & F. Walther, pp. 341–51. IUCN, Morges
Owen-Smith N. 1975. The social ethology of the white rhinoceros. Z. Tierpsychol. 38: 337–84
Owen-Smith N. 1979. Assessing the forage efficiency of a large herbivore, the kudu. S. Afr. J. Wildl. Res. 9: 102–10
Owen-Smith, N. 1981. The white rhinoceros overpopulation problem, and a proposed solution. In Problems in Management of Locally Abundant Wild Mammals, ed. P. A. Jewell, S. Holt & D. Hart, pp. 129–50. Academic Press, New York
Owen-Smith, N. 1982. Factors influencing the consumption of plant products by large herbivores. In The Ecology of Tropical Savannas, ed. B. J. Huntley & B. H. Walker, pp. 359–404. Springer-Verlag, Berlin and Hamburg
Owen-Smith, N. 1983. Dispersal and the dynamics of large herbivore populations in enclosed areas. In Management of Large Mammals in African Conservation Areas, ed. R. N. Owen-Smith, pp. 127–43. Haum, Pretoria
Owen-Smith, N. 1985. Niche separation among African ungulates. In Species and Speciation, ed. E. S. Vrba (Transvaal Museum Monograph No. 4), pp. 167–71
Owen-Smith, R. N. 1988. Megaherbivores. The Influence of Very Large Body Size on Ecology. (Cambridge Studies in Ecology). Cambridge University Press, Cambridge
Owen-Smith N. 1989. Megafaunal extinctions: the conservation message from 11 000 years BP. Cons. Biol. 3: 405–12
Owen-Smith N. 1990. Demography of a large herbivore, the greater kudu, in relation to rainfall. J. Anim. Ecol. 59: 893–913
Owen-Smith N. 1991. Veld condition and animal performance: application of an optimal foraging model. J. Grassl. Soc. Sth. Afr. 8: 77–81
Owen-Smith N. 1993a. Evaluating optimal diet models for an African browsing ruminant, the kudu: how constraining are the assumed constraints?Evol. Ecol. 7: 499–524
Owen-Smith N. 1993b. Comparative mortality rates of male and female kudus: the costs of sexual size dimorphism. J. Anim. Ecol. 62: 428–40
Owen-Smith N. 1993c. Age, size, dominance and reproduction among male kudus: mating enhancement by attrition of rivals. Behav. Ecol. Sociobiol. 32: 177–84
Owen-Smith N. 1994. Foraging responses of kudus to seasonal changes in food resources: elasticity in constraints. Ecology 75: 1050–62
Owen-Smith N. 1996a. Ecological guidelines for waterpoints in extensive protected areas. S. Afr. J. Wildl. Res. 26: 107–12
Owen-Smith N. 1996b. Circularity in linear programming models of optimal diet. Oecologia 108: 259–61
Owen-Smith N. 1997. Control of energy balance by a wild ungulate, the kudu, through adaptive foraging behaviour. Proc. Nutr. Soc. 56: 15–24
Owen-Smith N. 1998a. How high ambient temperature affects the daily activity and foraging time of a subtropical ungulate, the greater kudu. J. Zool., Lond. 246: 183–92
Owen-Smith, N. 1998b. Dynamics of herbivore-vegetation systems and the overgrazing issue. In Communal Rangelands in Southern Africa: A Synthesis of Knowledge. Proceedings of a Symposium on Policy-making for the Sustainable Use of Southern African Communal Rangelands, ed. T. D. de Bruyn & P. F. Scogings, pp. 124–34. Dept. of Livestock & Pasture Science, University of Fort Hare, Alice, South Africa
Owen-Smith N. 2000. Modelling the population dynamics of a subtropical ungulate in a variable environment: rain, cold and predators. Natur. Res. Model. 13: 57–87
Owen-Smith N. & Cooper S. M.. 1985. Comparative consumption of vegetation components by kudus, impalas and goats in relation to their commercial potential as browsers in savanna regions. S. Afr. J. Sci. 81: 72–6
Owen-Smith N. & Cooper S. M.. 1987a. Palatability of woody plants to browsing ungulates in a South African savanna. Ecology 68: 319–31
Owen-Smith N. & Cooper S. M.. 1987b. Acceptability indices for assessing the food preferences of ungulates. J. Wildl. Mgmt 51: 372–8
Owen-Smith N. & Cooper S. M.. 1989. Nutritional ecology of a browsing ruminant, the kudu, through the seasonal cycle. J. Zool., Lond. 219: 29–43
Owen-Smith, N. & J. E. Danckwerts. 1997. Herbivory. In Vegetation of Southern Africa, ed. R. M. Cowling, D. M. Richardson & S. M. Pierce, pp. 397–420. Cambridge University Press
Owen-Smith N. & Novellie P.. 1982. What should a clever ungulate eat?Am. Nat. 119: 151–78
Panaretto, B. A. 1968. Estimation of body composition by the dilution of hydrogen isotopes. In Body Composition in Animals and Man, pp. 200–17. National Academy of Sciences, Washington, DC
Papageorgiou, N. 1979. Population energy relationships of the Agrimi on Theodorou Island, Greece. Mammalia Depicta. Verlag Paul Parey, Berlin
Parker K. L. 1988. Effects of heat, cold and rain on coastal black-tailed deer. Can. J. Zool. 66: 2475–83
Parker K. L., Gillingham M. P., Hanley T. A. & Robbins C. T.. 1993. Seasonal patterns in body weight, body composition, and water transfer rates of free ranging and captive black tailed deer in Alaska. Can. J. Zool. 71: 1397–404
Parker K. L., Gillingham M. P., Hanley T. A. & Robbins C. T.. 1996. Foraging efficiency: energy expenditure versus energy gain in free ranging black tailed deer. Can. J. Zool. 74: 442–50
Parker, K. L., M. P. Gillingham, T. A. Hanley & C. T. Robbins. 1999. Energy and protein balance of free-ranging black-tailed deer in a natural forest environment. Wildl. Monogr. No. 143, 48pp
Parsons A. J., Newman J. A., Penning P. D., Harvey A. & Orr R. J.. 1994. Diet preference of sheep: effects of recent diet, physiological state and species abundance. J. Anim. Ecol. 63: 465–78
Pastor J., Moen R. A. & Cohen Y.. 1997. Spatial heterogeneity, carrying capacity and feedbacks in animal-landscape interactions. J. Mamm. 78: 1040–52
Peel M. J. S., Biggs H. & Zacharias P. J. K.. 1999. The evolving use of stocking rate indices currently based on animal number and type in semi-arid heterogeneous landscapes and complex land-use systems. Afr. J. Range Forage Sci. 15: 117–27
Pellew R. A. 1984. The feeding ecology of a selective browser, the giraffe. J. Zool., Lond. 202: 57–81
Penry, D. L. 1993. Digestive constraints on diet choice. In Diet Selection, ed. R. N. Hughes, pp. 32–55. Blackwell, Oxford
Penry D. L. & Jumars P. A.. 1986. Chemical reactor analysis and optimal digestion. BioScience 36: 310–15
Penry D. L. & Jumars P. A.. 1987. Modelling animal guts as chemical reactors. Am. Nat. 129: 69–96
Perez-Barberia F. J. & Gordon I. J.. 2001. Relationships between oral morphology and feeding style in the Ungulata: a phylogenetically controlled evaluation. Proc. R. Soc. Lond. B 268: 1021–30
Peters, R. H. 1983. The Ecological Implications of Body Size. Cambridge University Press, Cambridge
Peterson C. C., Nagy K. A. & Diamond J.. 1990. Sustained metabolic scope. Proc. Nat. Acad. Sci. USA 87: 2324–8
Peterson R. O. 1999. Wolf-moose interaction on Isle Royale: the end of natural regulation?Ecol. Applic. 9: 10–16
Peterson R. O., Page R. E. & Dodge K. M.. 1984. Wolves, moose and the allometry of population cycles. Science 224: 1350–2
Pielou, E. 1969. An Introduction to Mathematical Ecology. Wiley-Interscience, New York
Prigogine I. & Wiame J. M.. 1946. Biologie et thermodynamique des phenomenes irreversibles. Experientia 2: 451–3
Price, P. W. 1984. Insect Ecology. 2nd edn. John Wiley & Sons, New York
Prins, H. H. T. 1996. Ecology and Behaviour of the African Buffalo. Chapman & Hall, London
Prins, H. H. T. & H. Olff. 1998. Species richness of African grazer assemblages: towards a functional explanation. In Dynamics of Tropical Ecosystems, ed. D. N. Newberry, H. H. T. Prins & N. Brown, pp. 449–90. Blackwell, Oxford
Provenza F. D. 1995. Postingestive feedback as an elementary determinant of food preference and intake in ruminants. J. Range Mgmt 48: 2–17
Pulliam H. R. 1988. Sources, sinks and population regulation. Am. Nat. 132: 652–61
Rasmussen D. I. 1941. Biotic communities of the Kaibab Plateau. Ecol. Monogr. 3: 229–75
Real L. A. 1977. The kinetics of functional response. Am. Nat. 111: 289–300
Reimers E., Ringberg J. & Sorumgard R.. 1982. Body composition of Svalbard reindeer. Can. J. Zool. 60: 1812–21
Renecker L. A. & Hudson R. J.. 1986. Seasonal foraging rates of free-ranging moose. J. Wildl. Mgmt 50: 143–7
Ricklefs R. E., Konarzewski M. & Daan S.. 1996. The relationship between basal metabolic rate and daily energy expenditure in birds and mammals. Am. Nat. 147: 1047–71
Ringberg T. M., White R. G., Holleman D. F. & Luick J. R.. 1981. Body growth and carcase composition of lean reindeer from birth to sexual maturity. Can. J. Zool. 59: 1040–61
Risenhoover, K. L. 1987. Winter foraging strategies of moose in subarctic and boreal forest habitats. PhD thesis, Michigan Technological University
Risenhoover K. L. & Peterson R. O.. 1986. Mineral licks as a sodium source for Isle Royal moose. Oecologia 71: 121–6
Robbins, C. T. 1993. Wildlife Feeding and Nutrition. Academic Press, New York
Robbins C. T., Prior R. L., Moen A. N. & Vesek W. J.. 1974. Nitrogen metabolism of white-tailed deer. J. Anim. Sci. 38: 186–91
Robbins C. T., Podbielancik-Norman R. S., Wilson D. L. & Mould E. P.. 1981. Growth and nutrient consumption of elk calves compared to other ungulate species. J. Wildl. Mgmt 45: 172–86
Robbins C. T., Spalinger D. E. & Hoven W.. 1995. Adaptions of ruminants to browse and grass diets: are anatomical-based browser-grazer interpretations valid?Oecologia 103: 208–13
Rosenzweig M. L. 1971. Paradox of enrichment: destabilization of exploitation systems. Science 171: 385–7
Rosenzweig M. L. 1981. A theory of habitat selection. Ecology 62: 327–35
Rosenzweig M. L. 1991. Habitat selection and population interactions: the search for mechanism. Am. Nat. 137 (Suppl.): 5–28
Rosenzweig M. L. & Abramsky Z.. 1997. Two gerbils of the Negev: a long-term investigation of optimal habitat selection and its consequences. Evol. Ecol. 11: 733–56
Roughgarden, J. 1997. Production functions for ecological populations: a survey with emphasis on spatially explicit models. In Spatial Ecology, ed. D. Tilman & P. Kareiva, pp. 296–317. Princeton University Press, Princeton, NJ
Ruel J. J. & Ayres M. P.. 1999. Jensen's inequality predicts effects of environmental variation. Trends Ecol. Evol. 14: 361–6
Rutherford M. C. 1980. Annual plant production-precipitation relations in arid and semi-arid regions. S. Afr. J. Sci. 76: 53–6
Rutherford M. C. 1984. Relative allocation and seasonal phasing of growth of woody plant components in a South African savanna. Progr. Biometeorol. 3: 200–21
Rutherford M. C. & Panagos M. D.. 1982. Seasonal woody plant shoot growth in Burkea africana — Ochna pulchra savanna. S. Afr. J. Bot. 1: 104–16
Sale P. P. & Tolimieri N.. 2000. Density dependence at some time and place. Oecologia 124: 166–71
Scheffer V. B. 1951. The rise and fall of a reindeer herd. Scient. Monthly 73: 356–62
Schmidt K. A., Eambardt J. M., Brown J. S. & Holt R. D.. 2000. Habitat selection under temporal heterogeneity: exorcizing the ghost of competition past. Ecology 81: 2622–30
Schmidt-Nielsen, K. 1975. Animal Physiology. Adaptation and Environment. Cambridge University Press, Cambridge
Schmitz O. J., Beckerman A. P. & Litman S.. 1997. Functional responses of adaptive consumers and community stability with emphasis on the dynamics of plant-herbivore systems. Evol. Ecol. 11: 773–84
Scholes, R. J. & B. H. Walker. 1993. An African Savanna. Synthesis of the Nylsvley Study. Cambridge University Press, Cambridge
Scoones I. 1992. Coping with drought: responses of herders and livestock in contrasting savanna environments in Southern Zimbabwe. Human Ecol. 20: 293–314
Scoones, I. 1993. Why are there so many animals? Cattle population dynamics in the communal areas of Zimbabwe. In Range Ecology at Disequilibrium, ed. R. H. Behnke, I. Scoones & C. Kerven, pp. 62–76. ODI Publications, London
Scoones I. 1995. Exploiting heterogeneity: habitat use by cattle in dryland Zimbabwe. J. Arid Envir. 29: 221–37
Senft R. L., Stilwell M. A. & Rittenhouse L. R.. 1987. Nitrogen and energy budgets of free-roaming cattle. J. Range Mgmt 40: 421–4
Shackleton C. M. 1993. Are the communal grazing lands in need of saving?Development Southern Africa 10: 65–78
Shipley L. A., Gross J. G., Spalinger D. E., Hobbs N. T. & Wunder B. A.. 1994. The scaling of the functional response of mammalian herbivores. Am. Nat. 143: 1055–82
Shipley L. A. & Spalinger D. E.. 1995. Influence of size and density of browse patches on intake rates and foraging decisions of young moose and white-tailed deer. Oecologia 104: 112–21
Short J. 1985. The functional response of kangaroos, sheep and rabbits in an arid grazing system. J. Appl. Ecol. 22: 435–47
Shugart, H. H. 1998. Terrestrial Ecosystems in Changing Environments. Cambridge University Press, Cambridge
Sibly, R. M. 1981. Strategies of digestion and defecation. In Physiological Ecology, ed. C. R. Townsend & P. Calow, pp. 109–39. Blackwell, Oxford
Sinclair A. R. E. 1974. The natural regulation of buffalo populations in East Africa. IV. The food supply as a regulating factor and competition. E. Afr. Wildl. J. 12: 291–321
Sinclair, A. R. E. 1977. The African Buffalo. A Study of Resource Limitation in Populations. University of Chicago Press, Chicago
Sinclair A. R. E. 1985. Does interspecific competition or predation shape the African ungulate community?J. Anim. Ecol. 54: 899–918
Sinclair, A. R. E. 1989. Population regulation in animals. In Ecological Concepts, ed. J. M. Cherrett, pp. 197–241. Blackwell, Oxford
Sinclair A. R. E. & Arcese P.. 1995. Population consequences of predation-sensitive foraging: the Serengeti wildebeest. Ecology 76: 882–91
Sinclair A. R. E., Dublin H. & Borner M.. 1985. Population regulation of Serengeti wildebeest: a test of the food hypothesis. Oecologia 65: 266–8
Sinclair A. R. E. & Duncan P.. 1972. Indices of condition in tropical ruminants. E. Afr. Wildl. J. 10: 143–9
Sinclair A. R. E. & Norton-Griffiths M.. 1983. Does competition or facilitation regulate migrant ungulate populations in the Serengeti? A test of hypotheses. Oecologia 53: 364–9
Singer F. J., Harting A., Symonds K. K. & Coughenour M. B.. 1997. Density dependence, compensation and environmental effects on elk calf mortality in Yellowstone National Park. J. Wildl. Mgmt 61: 12–25
Skogland T. 1985. The effects of density-dependent resource limitations on the demography of wild reindeer. J. Anim. Ecol. 54: 359–74
Skogland T. 1988. Tooth wear by food limitation and its life history consequences in wild reindeer. Oikos 51: 238–42
Skogland T. 1990. Density dependence in a fluctuating wild reindeer herd: maternal vs offspring effects. Oecologia 84: 442–50
Smith N. S. 1970. Appraisal of condition estimation models for East African ungulates. E. Afr. Wildl. J. 8: 123–9
Sohal R. S. & Weindruch R.. 1996. Oxidative stress, caloric restriction and aging. Science 273: 59–63
Solomon M. E. 1949. The natural control of animal populations. J. Anim. Ecol. 18: 1–35
Spalinger D. E. & Hobbs N. T.. 1992. Mechanisms of foraging in mammalian herbivores: new models of functional response. Am. Nat. 140: 325–48
Spalinger D. E., Robbins C. T. & Hanley T. A.. 1993. Adaptive rumen function in elk and mule deer. Can. J. Zool. 71: 601–10
Staddon, J. E. R. 1983. Adaptive Behavior and Learning. Cambridge University Press, Cambridge
Stafford Smith, D. M. 1996. Management of rangelands: paradigms at their limits. In The Ecology and Management of Grazing Systems, ed. J. Hodgson & A. Illius, pp. 325–58. CAB International, Wallingford, UK
Starfield, A. M., K. A. Smith & A. L. Bleloch. 1990. How to Model It. Problem Solving for the Computer Age. McGraw-Hill, New York
Stephens, D. W. & J. R. Krebs. 1986. Foraging Theory. Princeton University Press, Princeton, NJ
Sutherland, W. J. 1996. From Individual Behaviour to Population Ecology. Oxford University Press, Oxford
Swift, M. J., O. W. Heal & J. M. Anderson. 1979. Decomposition in Terrestrial Ecosystems. Blackwell Scientific Publications, Oxford
Tainton, N. M. 1981. The ecology of the main grazing lands of South Africa. In Veld and Pasture Management, ed. N. M. Tainton, pp. 25–56. Shuter & Shooter, Pietermaritzburg
Tainton N. M. 1988. A consideration of veld condition assessment techniques for commercial livestock production in South Africa. J. Grassl. Soc. Sth. Afr. 5: 76–9
Tainton, N. M. 1999. Veld Management in South Africa. University of Natal Press, Pietermaritzburg
Thom, R. 1975. Structural Stability and Morphogenesis. W. A. Benjamin Co., Reading, MA
Thornley, J. H. M. & J. R. Johnson. 1990. Plant and Crop Modelling. Oxford University Press, Oxford
Thrash I. & Derry J. F.. 1999. The nature and modelling of piospheres: a review. Koedoe 42: 73–94
Tilman D. 1980. Resources: a graphical-mechanistic approach to competition and predation. Am. Nat. 116: 362–93
Tilman, D. 1982. Resource Competition and Community Structure. Princeton University Press, Princeton, NJ
Tilman D. 1986. A consumer-resource approach to community structure. Am. Zool. 26: 3–22
Tilman, D. 1988. Plant Strategies and the Dynamics and Structure of Plant Communities. (Monographs in Population Biology No. 26.) Princeton University Press, Princeton, NJ
Tilman D. 1996. Biodiversity: population versus ecosystem stability. Ecology 77: 350–63
Tilman D. 1997. Community invasability, recruitment limitation and grassland biodiversity. Ecology 78: 81–92
Tilman, D. & P. Kareiva. 1997. Spatial Ecology. The Role of Space in Population Dynamics and Interspecific Interactions. Princeton University Press, Princeton, NJ
Tolkamp B. J. & Ketelaars J. J. M. H.. 1992. Towards a new theory of feed intake regulation in ruminants 2. Costs and benefits of feed consumption: An optimization approach. Livestock Production Science 30: 97–317
Torbit S. C., Carpenter L. H., Allridge A. W. & Swift D. M.. 1985. Mule deer body composition — a comparison of methods. J. Wildl. Mgmt 49: 86–91
Torbit S. C., Carpenter L. H., Swift D. M. & Alldredge A. W.. 1982. Differential loss of fat and protein by mule deer during winter. J. Wildl. Mgmt 49: 80–5
Trudell J. & White R. G.. 1981. The effect of forage structure and availability on food intake, biting rate, bite size and daily eating time of reindeer. J. Appl. Ecol. 18: 63–81
Turner M. G., Wu Y., Romme W. H. & Wallace L. L.. 1993. A landscape simulation model of winter foraging by large ungulates. Ecol. Model. 69: 163–84
Tyler N. J. C. 1987. Body composition and energy balance of pregnant and non-pregnant Svalbard reindeer during winter. Symp. Zool. Soc. Lond. 57, 203–29
Underwood R. 1982. Vigilance behaviour in grazing African ungulates. Behaviour 79: 81–107
Underwood R. 1983. The feeding behaviour of grazing African ungulates. Behaviour 84: 195–243
Ungar E. D., Seligman N. G. & Demment M. W.. 1992. Graphical analysis of sward depletion by grazing. J. Appl. Ecol. 29: 427–35
Meer J. & Ens B. J.. 1997. Models of interference and their consequences for the spatial distribution of ideal and free predators. J. Anim. Ecol. 66: 846–58
Van Soest, P. J. 1982. Nutritional Ecology of the Ruminant. O & B Books Inc., Corvallis, OR
van Wieren, S. E. 1992. Factors limiting food intake in ruminants and non-ruminants in the temperate zone. In Ongules/Ungulates 91, ed. F. Spitz, G. Janeau, G. Gonzalez & S. Aulagnier, pp. 139–145. SFEPM-IRGM, Paris, Tolouse
Wieren S. E. 1997. Do large herbivores select a diet that maximizes short-term energy-intake rate?For. Ecol. Mgmt 88: 149–56
Verlinden C. & Wiley R. H.. 1989. The constraints of digestive rate: an alternative model of diet selection. Evol. Ecol. 3: 264–73
Vesey-Fitzgerald D. T. 1960. Grazing succession among East African game animals. J. Mamm. 41: 161–72
Vivas H. J., Saether B.-E. & Anderson R.. 1991. Optimal twig-size selection of a generalist herbivore, the moose: implications for plant-herbivore interactions. J. Anim. Ecol. 60: 395–408
Volterra V. 1926. Fluctuations in the abundance of species considered mathematically. Nature 118: 558–60
Walker B. H., Emslie R. H., Owen-Smith N. & Scholes R. J.. 1987. To cull or not to cull: lessons from a Southern African drought. J. Appl. Ecol. 24: 381–402
WallisDeVries M. & Daleboudt C.. 1994. Foraging strategy of cattle in patchy grassland. Oecologia 100: 98–106
WallisDeVries M. F., Laca E. O. & Demment M. W.. 1998. From feeding station to patch: scaling up food intake measurements in grazing cattle. Appl. Anim. Behav. Sci. 60: 301–15
Walters, C. 1986. Adaptive Management of Renewable Natural Resources. Macmillan, New York
Ward D. 1992. The role of satisficing in foraging theory. Oikos 63: 312–17
Watson, L. H. 1997. The feeding ecology of eland in Mountain Zebra National Park. PhD thesis, University of the Witwatersrand
Weber, B. H., D. J. Depew & J. D. Smith (eds). 1988. Entropy, Energy and Evolution. New Perspectives on Physical and Biological Evolution. MIT Press, Cambridge, MA
Weber G. E., Jeltsch F., Rooyen N. & Milton S. J.. 1998. Simulated long-term vegetation response to grazing heterogeneity in semi-arid rangelands. J. Appl. Ecol. 35: 687–99
Western D. 1975. Water availability and its influence on the structure and dynamics of a savannah large mammal community. E. Afr. Wildl. J. 13: 265–86
Westoby M. 1978. What are the biological bases of varied diets?Am. Nat. 112: 627–31
Westoby M., Walker B. & Noy-Meir I.. 1989. Opportunistic management for rangelands not at equilibrium. J. Range Mgmt 42: 266–74
White R. G. 1983. Foraging patterns and their multiplier effects on productivity of northern ungulates. Oikos 40: 377–84
Whyte J., Aarde R. & Pimm S. L.. 1998. Managing the elephants of Kruger National Park. Anim. Conserv. 1: 77–83
Wickstrom M. L., Robbins C. T., Hanley T. A., Spalinger D. E. & Parish S. M.. 1984. Food intake and foraging energetics of elk and mule deer. J. Wildl. Mgmt 48: 1285–301
Wiegand T., Jeltsch F., Bauer S. & Kellner K.. 1999. Simulation models for semi-arid rangelands of Southern Africa. Afr. J. Range Forage Sci. 15: 48–60
Wilmshurst J. F., Fryxell J. M. & Bergman C. M.. 2000. The allometry of patch selection in ruminants. Proc. R. Soc. Lond. B 267: 345–9
Wilmshurst J. F., Fryxell J. M. & Colucci P. E.. 1999a. What constrains daily intake in Thomson's gazelles?Ecology 80: 2338–47
Wilmshurst J. F., Fryxell J. M. & Hudson R. J.. 1995. Forage quality and patch choice by wapiti. Behav. Ecol. 6: 209–17
Wilmshurst J. F., Fryxell J. M., Farm B. P., Sinclair A. R. E. & Henschel C. P.. 1999b. Spatial distribution of Serengeti wildebeest in relation to resources. Can. J. Zool. 77: 1223–32
Wilson, E. O. & W. H. Bossert. 1971. A Primer of Population Biology. Sinauer, Sunderland, MA
Witter M. S. & Cuthill I. C.. 1993. The ecological costs of avian fat storage. Phil. Trans. Roy. Soc. Lond. B 340: 73–92
Woods S. C., Leely R. J., Porte D. & Schwartz M. W.. 1998. Signals that regulate food intake and energy homeostasis. Science 280: 1378–83
Woiwod I. P. & Hanski I.. 1992. Patterns of density dependence in moths and aphids. J. Anim. Ecol. 61: 619–29
Wright S. 1932. The roles of mutation, inbreeding, crossbreeding and selection in evolution. Proc. Ⅺ Int. Congr. Genet. 1: 356–66
Wu J. & Loucks O. L.. 1995. From balance of nature to hierarchical patch dynamics: A paradigm shift in ecology. Q. Rev. Biol. 70: 439–66
Yodzis, P. 1989. Introduction to Theoretical Ecology. Harper & Row, New York
Ziv Y. 2000. On the scaling of habitat specificity with body size. Ecology 81: 2932–38


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