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A Borophagine canid (Carnivora: Canidae: Borophaginae) from the middle Miocene Chesapeake Group of eastern North America

  • Steven E. Jasinski (a1) (a2) (a3) and Steven C. Wallace (a3) (a4)


A tooth recovered from the middle Miocene Choptank Formation (Chesapeake Group) of Maryland is identified as a new cynarctin borophagine (Canidae: Borophaginae: Cynarctina), here called Cynarctus wangi n. sp. The tooth, identified as a right upper second molar, represents the first carnivoran material reported from the Choptank Formation and part of a limited record of borophagine canids from eastern North America. As ?Cynarctus marylandica (Berry, 1938), another cynarctin borophagine from the older Calvert Formation, is known only from lower dentition, its generic affinities are uncertain. However, features of this new material are compared to features of ?C. marylandica through occlusal relationships, allowing for referral to a distinct species. Even so, the Choptank Formation material still offers two possible scenarios regarding its identification. In one, its geographic and stratigraphic provenance could imply that it belongs to ?C. marylandica. If this were correct, then the generic placement of ?C. marylandica would be correct and the taxon would be more derived than some other Cynarctus species in regard to hypocarnivory, and less derived than others. The second possibility, and the one believed to be most probable, is that a distinct cynarctin borophagine is present in the Chesapeake Group in strata younger than the type specimen of ?C. marylandica. This new borophagine canid expands the sparse fossil record of this group in northeastern North America and furthers our knowledge of the fossil record of terrestrial taxa in this region.



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Barbour, E.H., and Cook, H. J., 1914, Two new fossil dogs of the genus Cynarctus from Nebraska: Nebraska Geological Survey, v. 4, p. 225227.
Berry, C.T., 1938, A Miocene dog from Maryland: Proceedings of the United States National Museum, v. 85, p. 159161.
Blackwelder, B.W., and Ward, L.W., 1976, Stratigraphy of the Chesapeake Group of Maryland and Virginia Guidebook 7b (Northeast-Southeast Sections Joint Meeting 1976): Arlington, Geological Society of America, 52 p.
Bowdich, T.E., 1821, An Analysis of the Natural Classifications of Mammalia, Paris, J. Smith, 115 p.
Cope, E.D., 1867, An addition to the vertebrate fauna of the Miocene period, with a synopsis of the extinct Cetacea of the United States: Proceedings of the Academy of Natural Sciences of Philadelphia, v. 19, p. 138156.
Dahlberg, A.A., (ed.)., 1971, Dental Morphology and Evolution, Chicago, The University of Chicago Press, 350 p.
Fischer de Waldheim, G., 1817, Adversaria zoologica. Fasciculus primus: Quaedam ad Mammalium systema et genera illustranda: Mémoires de la Société impériale des naturalistes de Moscou, v. 5, p. 357446.
Flower, W.H., 1869, On the value of the characters of the base of the cranium in the classification of the order Carnivora, and on the systematic position of Bassaris and other disputed forms: Proceedings of the Zoological Society of London, v. 37, p. 437.
Gazin, C.L., and Collins, R.L., 1950, Remains of land mammals from the Miocene of the Chesapeake Bay region: Smithsonian Miscellaneous Collections, v. 116, p. 121.
Harlan, R., 1842, Notice of two fossil mammals from Brunswick Canal, Georgia; with observations on some of the fossil quadrupeds of the United States: American Journal of Science, v. 42, p. 141144.
Hayden, F.V., 1873, Report of the United States Geological Survey of the Territories, Washington, DC, US Government Printing Office, p. 259260.
Hershkovitz, P., 1971, Basic crown patterns and cusp homologies of mammalian teeth, in Dahlberg, A.A., ed., Dental Morphology and Evolution, Chicago, Illinois, The University of Chicago Press, p. 95150.
Kretzoi, M., 1943, Kochitis centenii n. g. n. sp., ein altertümlicher Creodonte aus dem Oberoligozän Siebenbürgens: Földtany Közlöny, v. 73, p. 1017. 190–195.
Linnaeus, C., 1758, Systema Naturae, 10th ed., v. 1: Stockholm, Laurentii Sylvii, 824 p.
Matthew, W.D., 1902, New Canidae from the Miocene of Colorado: Bulletin of the American Museum of Natural History, v. 16, p. 281290.
Matthew, W.D., 1924, Third contribution to the Snake Creek fauna: Bulletin of the American Museum of Natural History, v. 50, p. 59210.
Matthew, W.D., 1930, The phylogeny of dogs: Journal of Mammalogy, v. 11, p. 117138.
McGrew, P.O., 1937, The genus Cynarctus: Journal of Paleontology, v. 11, p. 444449.
McGrew, P.O., 1938, Dental morphology of the Procyonidae with a description of Cynarctoides, gen. nov: Geological Series of Field Museum of Natural History, v. 6, p. 323339.
McKenna, M.C., and Bell, S.K., 1997, Classification of Mammals: Above the Species Level, New York, Columbia University Press, 631 p.
Merriam, J.C., 1911, Tertiary mammal beds of Virgin Valley and Thousand Creek in northwestern Nevada: University of California Publications, Bulletin of the Department of Geology, Berkeley, v. 6, p. 199306.
Petuch, E.J., and Drolshagen, M., 2010, Molluscan Paleontology of the Chesapeake Miocene, Boca Raton, CRC Press, 180 p.
Shattuck, G.B., 1904, Geologic and paleontological relations, with a review of earlier investigations, in Clark, W.B., Shattuck, G.B., and Dan, W.H., eds., The Miocene Deposits of Maryland, Baltimore, Maryland Geological Survey, p. 3394.
Simpson, G.G., 1932, Miocene land mammals from Florida: Bulletin of the Florida State Geological Survey, v. 10, p. 141.
Simpson, G.G., 1945, The principles of classification and a classification of mammals: Bulletin of the American Museum of Natural History, v. 8, p. 1350.
Tedford, R.H., and Hunter, M.E., 1984, Miocene marine-nonmarine correlations, Atlantic and Gulf Coastal Plains, North America: Palaeogeography, Palaeoclimatology, Palaeoecology, v. 47, p. 129151.
Vogt, P.R., and Eshelman, R.E., 1987, Maryland’s Cliffs of Calvert: A fossiliferous record of mid-Miocene inner shelf and coastal environments, in Roy, D.C., ed., Geological Society of America Centennial Field Guide No. 5, Northeastern Section, Boulder, Colorado, Geological Society of America, p. 914.
Wang, X., Tedford, R.H., and Taylor, B.E., 1999, Phylogenetic systematics of the Borophaginae (Carnivora, Canidae): Bulletin of the American Museum of Natural History, v. 243, p. 1391.
Weems, R.E., and George, R.E., 2013, Amphibians and nonmarine turtles from the Miocene Calvert Formation of Delaware, Maryland, and Virginia (USA): Journal of Paleontology, v. 87, p. 570588.
Wright, D.B., and Eshelman, R.E., 1987, Miocene Tayassuidae (Mammalia) from the Chesapeake Group of the mid-Atlantic coast and their bearing on marine-nonmarine correlation: Journal of Paleontology, v. 61, p. 604618.

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A Borophagine canid (Carnivora: Canidae: Borophaginae) from the middle Miocene Chesapeake Group of eastern North America

  • Steven E. Jasinski (a1) (a2) (a3) and Steven C. Wallace (a3) (a4)


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