Boegoeberg 1 (BOG1) is located on the Atlantic coast of South Africa, 850 km north of Cape Town. The site is a shallow rock shelter in the side of a sand-choked gully that was emptied by diamond miners. Abundant coprolites, chewed bones, and partially digested bones implicate hyenas as the bone accumulators. The location of the site, quantity of bones, and composition of the fauna imply it was a brown hyena nursery den. The abundance of Cape fur seal bones shows that the hyenas had ready access to the coast. Radiocarbon dates place the site before 37,000 14C yr ago, while the large average size of the black-backed jackals and the presence of extralimital ungulates imply cool, moist conditions, probably during the early part of the last glaciation (isotope stage 4 or stage 3 before 37,000 14C yr ago) or perhaps during one of the cooler phases (isotope substages 5d or 5b) within the last interglaciation. Comparisons of the BOG1 seal bones to those from regional Middle Stone Age (MSA) and Later Stone Age (LSA) archeological sites suggest (1) that hyena and human seal accumulations can be distinguished by a tendency for vertebrae to be much more common in a hyena accumulation and (2) that hyena and LSA accumulations can be distinguished by a tendency for hyena-accumulated seals to represent a much wider range of individual seal ages. Differences in the way hyenas and people dismember, transport, and consume seal carcasses probably explain the contrast in skeletal part representation, while differences in season of occupation explain the contrast in seal age representation. Like modern brown hyenas, the BOG1 hyenas probably occupied the coast year-round, while the LSA people focused their coastal visits on the August–October interval when nine-to-eleven-month-old seals were abundant. The MSA sample from Klasies River Mouth Cave 1 resembles BOG1 in seal age composition, suggesting that unlike LSA people, MSA people obtained seals more or less throughout the year.

The climate of the Great Plains during the middle Holocene varied considerably, but overall it was marked by a north–south gradient of increasingly warmer and drier conditions, with a reduction in effective moisture, surface water, and resource abundance, and an increase in resource patchiness, sediment weathering, erosion, and aeolian activity. Pronounced drought conditions were most evident on the Southern High Plains. Understanding the human responses to middle Holocene climates is complicated by a lack of archaeological data, which is partly a result of geomorphic processes that removed or deeply buried sites of this age, and by the varying adaptive responses of hunter-gatherers during this period. On the Southern High Plains, where drought was most severe, surface and groundwater sources dried and bison populations were diminished, prompting substantial adaptive changes, including local abandonment, well-digging to tap underground water, and a widening of the diet breadth to incorporate higher-cost, lower-return seed and plant resources. Sites of this age on the Central and Northern Plains also show a possible increase in diet breadth (with the incorporation of plant foods in the diet), and perhaps changes in settlement mobility (including possible shift into higher elevation areas, or mapping-on to extant rivers and springs). But linking those changes to middle Holocene drought is less straightforward.