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Biolinguistics and the human language faculty

Published online by Cambridge University Press:  01 January 2026

Anna Maria Di Sciullo  and
Lyle Jenkins
Anna Maria Di Sciullo*
Affiliation:
University of Quebec in Montreal
Lyle Jenkins*
Affiliation:
Biolinguistics Institute
*
[di_sciullo.anne-marie@uqam.ca]
[ljenkins2@comcast.net]
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Abstract

In recent years linguists have gained new insight into human language capacities on the basis of results from linguistics and biology. The so-called biolinguistic enterprise aims to fill in the explanatory gap between language and biology, on both theoretical and experimental grounds, hoping to reach a deeper understanding of language as a phenomenon rooted in biology. This research program is taking its first steps, and it has already given rise to new insights on the human language capacity, as well as to controversies, echoing debates that go back to the earlier days of generative grammar. The present discussion piece provides a high-level characterization of biolinguistics. It highlights the main articulation of this research program and points to recent studies linking language and biology. It also compares the biolinguistic program, as defined in Chomsky 2005 and Di Sciullo & Boeckx 2011, to the view of the human language faculty presented in Jackendoff 2002 and Culicover & Jackendoff 2005, and to the discussion in Jackendoff 2011.

Keywords

biolinguisticsbiologyhuman language facultyuniversal grammarMergelanguage acquisitionminimalist programsymmetryassymetries

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Type
Discussion
Information
Language , Volume 92 , Issue 3 , September 2016 , pp. e205 - e236
Copyright
Copyright © 2016 Linguistic Society of America

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Footnotes

*

We wish to thank the participants of the Biolinguistic Conferences organized by Anna Maria Di Sciullo in 2010 and 2011 at UQAM, in 2013 at the GLOW meeting in Lund, and in 2015 at the IUSS Institute in Pavia, as well as three anonymous referees for valuable comments on an earlier version of this paper. This work has been supported in part by funding from the Social Sciences and Humanities Research Council of Canada to the Major Collaborative Research Initiative on Interface Asymmetries (http://www.interfaceasymmetry.uqam.ca/) and the Fonds Québécois de la recherche sur la société et la culture for the research on Dynamic Interfaces; see also http://www.biolinguistics.uqam.ca/.

References

Albertini, Silvia, Tettamanti, Marco; and Moro, Andrea. 2012. The impossible chaos: When the mind cannot eliminate language structure. Pavia: IUSS Center for Neurolinguistics and Theoretical Syntax, ms.Google Scholar
Alcock, Katherine J., Passingham, Richard E., Watkins, Kate E.; and Vargha-Khadem, Faraneh. 2000. Oral dyspraxia in inherited speech and language impairment and acquired dysphasia. Brain and Language 75. 17–33. DOI: 10.1006/brln.2000.2322.CrossRefGoogle ScholarPubMed
Anderson, P. W. 1972. More is different. Science 177. 393–96. DOI: 10.1126/science.177.4047.393.CrossRefGoogle ScholarPubMed
Berwick, Robert C. 2011. Syntax facit saltum redux: Biolinguistics and the leap to syntax. In Di Sciullo & Boeckx, 65–99.Google Scholar
Berwick, Robert c., Beckers, Gabriël J. L., Okanoya, Kazuo; and Bolhuis, Johan J.. 2012. A bird's eye view of human language evolution. Frontiers in Evolutionary Neuroscience 4:5. DOI: 10.3389/fnevo.2012.00005.CrossRefGoogle ScholarPubMed
Berwick, Robert C., and Chomsky, Noam. 2011. The biolinguistic program: The current state of its evolution. In Di Sciullo & Boeckx, 19–41.Google Scholar
Berwick, Robert C., and Chomsky, Noam. 2016. Why only us: Language and evolution. Cambridge, MA: MIT Press.CrossRefGoogle Scholar
Berwick, Robert c., Friederici, Angela D., Chomsky, Noam, Bolhuis, Johan J.. 2013. Evolution, brain, and the nature of language. Trends in Cognitive Sciences 17. 89–98. DOI: 10.1016/j.tics.2012.12.002.CrossRefGoogle ScholarPubMed
Berwick, Robert c., Pietroski, Paul, Yankama, Beracah; and Chomsky, Noam. 2011. Poverty of the stimulus revisited. Cognitive Science 35. 1207–42. DOI: 10.1111/j.1551-6709.2011.01189.x.CrossRefGoogle ScholarPubMed
Berwick, Robert C., and Weinberg, Amy S.. 1984. The grammatical basis of linguistic performance: Language use and acquisition. Cambridge, MA: MIT Press.Google Scholar
Bever, Thomas G. 1970. The cognitive basis for linguistic structures. Cognition and the development of language, ed. by Hayes, John R., 279–362. New York: Wiley.Google Scholar
Bever, Thomas G. 1981. Normal acquisition processes explain the critical period for language learning. Individual differences and universals in language learning aptitude, ed. by Diller, Karl C., 176–98. Rowley, MA: Newbury House.Google Scholar
Biberauer, Theresa (ed.) 2008. The limits of syntactic variation. Amsterdam: John Benjamins.CrossRefGoogle Scholar
Biberauer, Theresa, Holmberg, Anders; and Roberts, Ian. 2014. A syntactic universal and its consequences. Linguistic Inquiry 45. 169–225. DOI: 10.1162/LING_a_00153.CrossRefGoogle Scholar
Bickerton, Derek. 1990. Language and species. Chicago: University of Chicago Press.CrossRefGoogle Scholar
Bickerton, Derek. 1998. Catastrophic evolution: The case for a single step from protolanguage to full human language. Approaches to the evolution of language: Social and cognitive bases, ed. by Hurford, James R., Kennedy, Michael Studdert, and Knight, Chris, 341–58. Cambridge: Cambridge University Press.Google Scholar
Bickerton, Derek. 2000. How protolanguage became language. The evolutionary emergence of language: Social function and the origins of linguistic form, ed. by Knight, Chris, Studdert-Kennedy, Michael, and Hurford, James R., 264–84. Cambridge: Cambridge University Press.Google Scholar
Bickerton, Derek. 2008. Darwin's last word: How words changed cognition. Behavioral and Brain Science 31. 132. DOI: 10.1017/S0140525X08003579.CrossRefGoogle Scholar
Bickerton, Derek. 2014. More than nature needs: Language, mind and evolution. Cambridge, MA: Harvard University Press.CrossRefGoogle Scholar
Bishop, Dorothy V. Μ. 2015. The interface between genetics and psychology: Lessons from developmental dyslexia. Proceedings of the Royal Society B: Biological Sciences 282:20143139. DOI: 10.1098/rspb.2014.3139.CrossRefGoogle ScholarPubMed
Bishop, Dorothy v. m., Adams, Caroline v.; and Norbury, Courtenay F.. 2005. Distinct genetic influences on grammar and phonological short-term memory deficits: Evidence from 6-year-old twins. Genes, Brain and Behavior 5. 158–69. DOI: 10.1111/j.1601-183X.2005.00148.X.Google Scholar
Bishop, Dorothy V. M., North, T.; and Donlan, C.. 1995. Genetic basis of specific language impairment: Evidence from a twin study. Developmental Medicine and Child Neurology 37.1.56–71. DOI: 10.1111/j.1469-8749.1995.tb11932.x.CrossRefGoogle ScholarPubMed
Bishop, Dorothy V. Μ., and Snowling, Margaret J.. 2004. Developmental dyslexia and specific language impairment: Same or different? Psychological Bulletin 130. 858–86. DOI: 10.1037/0033-2909.130.6.858.CrossRefGoogle ScholarPubMed
Blanco-Elorrieta, Esti, and Pylkkänen, Liina. 2015. Composition of complex numbers: Delineating the computational role of the left anterior temporal lobe. NeuroImage 124. 194–203. DOI: 10.1016/j.neuroimage.2015.08.049.CrossRefGoogle ScholarPubMed
Bobaljik, Jonathan. 2012. Universals in comparative morphology: Suppletion, superlatives, and the structure of words. Cambridge, MA: MIT Press.CrossRefGoogle Scholar
Boeckx, Cedric. 2015. Elementary syntactic structures. Cambridge: Cambridge University Press.Google Scholar
Boeckx, Cedric, and Grohmann, Kleanthes К. (eds.) 2013. The Cambridge handbook of biolinguistics. Cambridge: Cambridge University Press.CrossRefGoogle Scholar
Bolhuis, Johan J., and Everaert, Martin (eds.) 2013. Birdsong, speech, and language: Exploring the evolution of mind and brain. Cambridge, MA: MIT Press.CrossRefGoogle Scholar
Bolhuis, Johan J., Tattersall, Ian, Chomsky, Noam; and Berwick, Robert С.. 2014. How could language have evolved? PLoS Biology 12. 8.e1001934. DOI: 10.1371/journal.pbio.1001934.CrossRefGoogle ScholarPubMed
Borer, Hagit. 1984. Parametric syntax: Case studies in Semitic and Romance languages. Dordrecht: Foris.CrossRefGoogle Scholar
Briscoe, Josie, Chilvers, Rebecca, Baldeweg, Torsten; and Skuse, David. 2012. A specific cognitive deficit within semantic cognition across a multi-generational family. Proceedings of the Royal Society B: Biological Sciences 279. 3652–61. DOI: 10.1098/rspb.2012.0894.Google ScholarPubMed
Cavalli-Sforza, Luigi, and Feldman, Marcus. 1981. Cultural transmission and evolution. Princeton, NJ: Princeton University Press.Google ScholarPubMed
Chesi, Cristiano, and Moro, Andrea. 2012. Computational blindness in the brain. Pavia: NeTS, IUSS Pavia, ms.Google Scholar
Chomsky, Noam. 1955. The logical structure of linguistic theory. Cambridge, MA: MIT, ms. Distributed by Indiana University Linguistics Club.Google Scholar
Chomsky, Noam. 1956. Three models for the description of language. IRE Transactions on Information Theory 2. 113–24.Google Scholar
Chomsky, Noam. 1957. Syntactic structures. The Hague: Mouton.CrossRefGoogle Scholar
Chomsky, Noam. 1965. Aspects of the theory of syntax. Cambridge, MA: MIT Press.Google Scholar
Chomsky, Noam. 1970. Remarks on nominalization. Readings in English transformational grammar, ed. by Jacobs, Roderick and Rosenbaum, Peter, 184–221. Waltham, MA: Ginn and Co.Google Scholar
Chomsky, Noam. 1976. On the biological basis of language capacities. The neuropsychology of language: Essays in honor of Eric Lenneberg, ed. by Rieber, Robert W., 1–24. New York: Plenum.Google Scholar
Chomsky, Noam. 1981. Lectures on government and binding: The Pisa lectures. Dordrecht: Foris.Google Scholar
Chomsky, Noam. 1986. Knowledge of language. New York: Praeger.Google Scholar
Chomsky, Noam. 1995. The minimalist program. Cambridge, MA: MIT Press.Google Scholar
Chomsky, Noam. 2000. Minimalist inquiries: The framework. Step by step: Essays in minimalist syntax in honor of Howard Lasnik, ed. by Martin, Roger, Michaels, David, and Uriagereka, Juan, 89–155. Cambridge, MA: MIT Press.Google Scholar
Chomsky, Noam. 2005. Three factors in language design. Linguistic Inquiry 36. 1–22. DOI: 10.1162/0024389052993655.CrossRefGoogle Scholar
Chomsky, Noam. 2007. Approaching UG from below. Interfaces + recursion = language? Chomsky s minimalism and the view from syntax-semantics, ed. by Sauerland, Uli and Gärtner, Hans-Martin, 1–30. Berlin: Mouton de Gruyter.Google Scholar
Chomsky, Noam. 2008. The biolinguistic program: Where does it stand today? Cambridge, MA: MIT Press.Google Scholar
Chomsky, Noam. 2011. Poverty of stimulus: Unfinished business. Transcript of lecture presented in series ‘Sprache und Gehirn—Zur Sprachfähigkeit des Menschen’ organized by Angela D. Friederici, Summer 2010. Cambridge, MA: MIT, ms.Google Scholar
Chomsky, Noam. 2013. Problems of projection. Lingua 130. 33–49. DOI: 10.1016/j.lingua.2012.12.003.CrossRefGoogle Scholar
Chomsky, Noam. 2015a. The minimalist program. 20th anniversary edn. Cambridge, MA: MIT Press.Google Scholar
Chomsky, Noam. 2015b. Problems of projection: Extensions. Structures, strategies and beyond: Studies in honour of Adriana Belletti, ed. by Domenico, Elisa Di, Hamann, Cornelia, and Matteini, Simona, 1–16. Amsterdam: John Benjamins.Google Scholar
Chomsky, Noam, and Miller, George. 1963. Introduction to the formal analysis of natural languages. Handbook of mathematical psychology, vol. 2, ed. by Luce, Duncan, Bush, Robert T., and Galanter, Eugene, 269–323. New York: Wiley.Google Scholar
Cinque, Guglielmo, and Kayne, Richard S. (eds.) 2005. The Oxford handbook of comparative syntax. Oxford: Oxford University Press.Google Scholar
Cinque, Guglielmo, and Rizzi, Luigi. 2010. The cartography of syntactic structures. The Oxford handbook of linguistic analysis, ed. by Heine, Bernd and Narrog, Heiko, 51–65. Oxford: Oxford University Press.Google Scholar
Clark, Eve v., and Barron, Brigid J. S.. 1988. A thrower-button or a button-thrower? Children's judgments of grammatical and ungrammatical compound nouns. Linguistics 26. 3–19. DOI: 10.1515/ling.1988.26.1.3.CrossRefGoogle Scholar
Concha, Miguel L., Bianco, Isaac H.; and Wilson, Stephen W.. 2012. Encoding asymmetry within neural circuits. Nature Reviews Neuroscience 13. 832–43. DOI: 10.1038/nrn3371.CrossRefGoogle ScholarPubMed
Culicover, Peter W., and Jackendoff, Ray S.. 2005. Simpler syntax. Oxford: Oxford University Press.CrossRefGoogle Scholar
de Villiers, Jill, and Roeper, Tom (eds.) 2011. Handbook of generative approaches to language acquisition. New York: Springer.CrossRefGoogle Scholar
Di Sciullo, Anna Maria. 2005. Asymmetry in morphology. Cambridge, MA: MIT Press.CrossRefGoogle Scholar
Di Sciullo, Anna Maria. 2011. A biolinguistic approach to variation. In Di Sciullo & Boeckx, 305–28.Google Scholar
Di Sciullo, Anna Maria. 2012a. An evolutionary developmental universal: Evidence from the morpho-syntactic evolution of the nominal domain. Paper presented at the Typology and Universals in Word Formation Conference, Pavol Jozef Safarik University, Slovakia.Google Scholar
Di Sciullo, Anna Maria. 2012b. Biolinguistics, minimalist grammars, and the emergence of complex numerals. Five approaches to language evolution: Proceedings of the workshops of the 9th International Conference on the Evolution of Language (EvolangIX), Kyoto, 13–18. Online: http://kyoto.evolang.org/sites/default/files/EvolangWorkshopProceedings.pdf.Google Scholar
Di Sciullo, Anna Maria. 2012c. Perspectives on morphological complexity. Morphology: (Ir)regularity, frequency, typology, ed. by Kiefer, Ferenc, Ladányi, Mária, and tár, Péter Sip-, 105–35. Amsterdam: John Benjamins.Google Scholar
Di Sciullo, Anna Maria. 2013. Exocentric compounds, language and proto-language. Language and Information Society 20. 1–26.CrossRefGoogle Scholar
Di Sciullo, Anna Maria. 2014. Minimalism and I-morphology. Minimalism and beyond: Radicalizing the interfaces, ed. by Kosta, Peter, Franks, Steven, Radeva-Bork, Teodora, and Schürcks, Lilia, 267–86. Amsterdam: John Benjamins.Google Scholar
Di Sciullo, Anna Maria. 2015. On the domain specificity of the human language faculty and the effects of principles of computational efficiency: Contrasting language and mathematics. Revista Linguíʃtica 11. 28–56. DOI: 10.17074/2238-975X.2015v11n1p28.Google Scholar
Di Sciullo, Anna Maria, and Agüero-Bautista, Calixto. 2008. The delay of principle B effect (DPBE) and its absence in some languages. Language and Speech 51. 77–100. DOI: 10.1177/00238309080510010601.CrossRefGoogle ScholarPubMed
Di Sciullo, Anna Maria, and Boeckx, Cedric (eds.) 2011. The biolinguistic enterprise: New perspectives on the evolution and nature of the human language faculty. Oxford: Oxford University Press.Google Scholar
Di Sciullo, Anna Maria, and Isac, Dana. 2008. The asymmetry of Merge. Biolinguistics 2. 260–90. Online: http://www.biolinguistics.eu/index.php/biolinguistics/article/view/53.CrossRefGoogle Scholar
Di Sciullo, Anna Maria, and Nicolis, Marco. 2012. Third factor and the development of P. Paper presented at North East Linguistic Society (NELS) 42, University of Toronto.Google Scholar
Di Sciullo, Anna Maria, Nicolis, Marco; and Somesfalean, Stanca. 2017. Comitative P. Variation in prepositions (Oxford studies in comparative syntax), ed. by Garzonio, Jacopo and Rossi, Silvia. Oxford: Oxford University Press, to appear.Google Scholar
Di Sciullo, Anna Maria, Piattelli-Palmarini, Massimo, Wexler, Ken, Berwick, Robert c., Boeckx, Cedric, Jenkins, Lyle, Uriagereka, Juan, Stromswold, Karin, Cheng, Lisa, Harley, Heidi, Wedel, Andrew, McGilvray, James, Gelderen, Elly van; and Bever, Thomas. 2010. The biological nature of human language. Biolinguistics 4. 4–34. Online: http://www.biolinguistics.eu/index.php/biolinguistics/article/view/110.CrossRefGoogle Scholar
Di Sciullo, Anna Maria, and Somesfalean, Stanca. 2013. The definite determiner in Romanian: A biolinguistic perspective. Australian Journal of Linguistics (Special issue: Romance linguistics in the Pacific: Variation in time and space) 33. 121–39. DOI: 10.1080/07268602.2013.814526.Google Scholar
Di Sciullo, Anna Maria, and Somesfalean, Stanca. 2015. Object pronouns in the evolution of Romanian: A biolinguistic perspective. Formal approaches to DPs in Old Romanian, ed. by Hill, Virginia, 269–90. Leiden: Brill.Google Scholar
Embick, David, Marantz, Alec, Miyashita, Yasushi, O’Neil;, Wayne and Sakai, Kuniyoshi L.. 2000. A syntactic specialization for Broca's area. Proceedings of the National Academy of Sciences 97. 6150–54. DOI: 10.1073/pnas.100098897.CrossRefGoogle ScholarPubMed
Emonds, Joseph E. 1976. A transformational approach to English syntax. New York: Academic Press.Google Scholar
Enard, Wolfgang, Przeworski, Molly, Fisher, Simon e., Lai, Celica s., Wiebe, Victor, Kitano, Takashi, Monaco, Antony P.; and Pääbo, Svante. 2002. Molecular evolution of FOXP2, a gene involved in speech and language. Nature 418. 869–72. DOI: 10.1038/nature01025.CrossRefGoogle ScholarPubMed
Everett, Daniel L. 2004. Coherent fieldwork. Linguistics today—Facing a greater challenge, ed. by Sterkenburg, Piet van, 141–62. Amsterdam: John Benjamins.Google Scholar
Everett, Daniel L. 2005. Cultural constraints on grammar and cognition in Pirahã: Another look at the design features of human language. Current Anthropology 46. 4–11. DOI: 10.1086/431525.CrossRefGoogle Scholar
Fisher, Simon E., and Marcus, Gary F.. 2006. The eloquent ape: Genes, brains and the evolution of language. Nature Reviews Genetics 7. 9–20. DOI: 10.1038/nrg1747.CrossRefGoogle ScholarPubMed
Fitch, W. Tecumseh. 2010. The evolution of language. New York: Cambridge University Press.CrossRefGoogle ScholarPubMed
Fitch, W. Tecumseh, and Hauser, Mark D.. 2004. Computational constraints on syntactic processing in a nonhuman primate. Science 303. 377–80. DOI: 10.1126/science.1089401.CrossRefGoogle Scholar
Fodor, Jerry, Bever, Thomas G.; and Garrett, Merrill F.. 1974. The psychology of language: An introduction to psycholinguistics and generative grammar. New York: McGraw-Hill.Google Scholar
Fox, Danny. 1999. Local economy. Cambridge, MA: MIT Press.Google Scholar
Frampton, John, and Gutmann, Sam. 2002. Crash-proof syntax. Derivation and explanation in the minimalist program, ed. by Epstein, Samuel D. and Seely, T. David, 90–105. Oxford: Blackwell.CrossRefGoogle Scholar
Friederici, Angela D. 2009. The brain differentiates hierarchical and probabilistic grammars. Of minds and language: A dialogue with Noam Chomsky in the Basque country, ed. by Piattelli-Palmarini, Massimo, Uriagereka, Juan, and Salaburu, Pello, 184–94. New York: Oxford University Press.Google Scholar
Friederici, Angela d., Bahlmann, Jörg, Friedrich, Roland; and Makuuchi, Michiru. 2011. The neural basis of recursion and complex syntactic hierarchy. Biolinguistics 5. 87–104. Online: http://www.biolinguistics.eu/index.php/biolinguistics/article/view/170.CrossRefGoogle Scholar
Friederici, Angela d., Bahlmann, Jörg, Heim, Stefan, Schubotz, Ricarda I.; and Anwander, Alfred. 2006. The brain differentiates human and non-human grammars: Functional localization and structural connectivity. Proceedings of the National Academy of Sciences 103. 2458–63. DOI: 10.1073/pnas.0509389103.CrossRefGoogle ScholarPubMed
Friedrich, Roland, and Friederici, Angela D.. 2009. Mathematical logic in the human brain: Syntax. PLoS ONE 4:e5599. DOI: 10.1371/journal.pone.0005599.CrossRefGoogle Scholar
Friedrich, Roland, and Friederici, Angela D.. 2013. Mathematical logic in the human brain: Semantics. PLoS ONE 8:e53699. DOI: 10.1371/journal.pone.0053699.CrossRefGoogle Scholar
Geschwind, Norman, and Galaburda, Albert Μ. (eds.) 1984. Cerebral dominance: The biological foundations. Cambridge, MA: Harvard University Press.Google Scholar
Geschwind, Norman, and Galaburda, Albert Μ.. 1987. Cerebral lateralization. Cambridge, MA: MIT Press.Google Scholar
Gil, David. 2011. Where does predication come from? Leipzig: Max Planck Institute for Evolutionary Anthropology, ms.Google Scholar
Gopnik, Myrna, and Crago, Martha В.. 1991. Familial aggregation of a developmental language disorder. Cognition 39. 1–50. DOI: 10.1016/0010-0277(91)90058-C.CrossRefGoogle ScholarPubMed
Gould, Stephen Jay. 1977. Ontogeny and phylogeny. Cambridge, MA: Harvard University Press.Google Scholar
Gould, Stephen Jay. 2002. The structure of evolutionary theory. Cambridge, MA: Harvard University Press.Google Scholar
Gowers, Timothy, and Barrow-Green, June (eds.) 2008. The Princeton companion to mathematics. Princeton: Princeton University Press.Google Scholar
Graham, Sarah A., and Fisher, Simon E.. 2015. Understanding language from a genomic perspective. Annual Review of Genetics 49. 131–60. DOI: 10.1146/annurev-genet-120213-092236.CrossRefGoogle ScholarPubMed
Greenberg, Joseph H. 1966. Language universals, with special reference to feature hierarchies. The Hague: Mouton.Google Scholar
Grodzinsky, Yosef, and Amunts, Katrin (eds.) 2006. Brocas region. Oxford: Oxford University Press.CrossRefGoogle ScholarPubMed
Hale, Kenneth L. 1983. Warlpiri and the grammar of non-configurational languages. Natural Language and Linguistic Theory 1. 1. 5–47. DOI: 10.1007/BF00210374.CrossRefGoogle Scholar
Hale, Kenneth L., Laughren, Mary; and Simpson, Jane. 1995. Warlpiri syntax. Syntax: An international handbook of contemporary research, ed. by Jacobs, Joachim, Stechow, Arnim von, Sternefeld, Wolfgang, and Vennemann, Theo, 1430–51. Berlin: Walter de Gruyter.Google Scholar
Halle, Mark, and Marantz, Alec. 1993. Distributed morphology and the pieces of inflection. The view from Building 20: Essays in linguistics in honor of Sylvain Bromberger, ed. by Hale, Kenneth and Keyser, Samuel Jay, 111–76. Cambridge, MA: MIT Press.Google Scholar
Hallgrimsson, Benedikt, and Hall, Brian K. (eds.) 2005. Variation: A central concept in biology. London: Academic Press.Google Scholar
Hancock, Roeland, and Bever, Thomas G.. 2013. Genetic factors and normal variation in the organization of language. Biolinguistics 7. 75–95. Online: http://www.biolinguistics.eu/index.php/biolinguistics/article/view/272/296.CrossRefGoogle Scholar
Hauser, Marc D., Chomsky, Noam; and Fitch, W. Tecumseh. 2002. The faculty of language: What is it, who has it, and how did it evolve? Science 298. 1569–79. DOI: 10.1126/science.298.5598.1569.CrossRefGoogle ScholarPubMed
Hauser, Marc d., Yang, Charles, Berwick, Robert c., Tattersall, Ian, Ryan, Michael J., Watumull, Jeffrey, Chomsky, Noam; and Lewontin, Richard С.. 2014. The mystery of language evolution. Frontiers in Psychology 5:00401. DOI: 10.3389/fpsyg.2014.00401.CrossRefGoogle ScholarPubMed
Hilliard, Austin t., Miller, Julie e., Fraley, Elizabeth r., Horvath, Steve; and White, Stephanie A.. 2012. Molecular microcircuitry underlies functional specification in a basal ganglia circuit dedicated to vocal learning. Neuron 73. 537–52. DOI: 10.1016/j.neuron.2012.01.005.CrossRefGoogle Scholar
Hiramatsu, Kazuko, Snyder, William, Roeper, Thomas, Storrs, Stephanie; and Saccoman, Matthew. 2000. Of musical hand chairs and linguistic swing. Proceedings of the Boston University Conference on Language Development (BUCLD) 24. 409–17.Google Scholar
Hugdahl, Kenneth, and Westerhausen, René (eds.) 2010. The two halves of the brain: Information processing in the cerebral hemispheres. Cambridge, MA: MIT Press.CrossRefGoogle Scholar
Hurford, James. 2001. Protothought had no logical names. New essays on the origins of language, ed. by Trabant, Jürgen and Ward, Sean, 117–30. Berlin: Mouton de Gruyter.Google Scholar
Hurford, James. 2012. Language in the light of evolution. Oxford: Oxford University Press.Google Scholar
Hurford, James. 2014. The origins of language: A slim guide. New York: Oxford University Press.Google Scholar
Hurst, Jane a., Baraitser, Michael, Auger, Elizabeth, Graham, Frances; and Norell, S.. 1990. An extended family with a dominantly inherited speech disorder. Developmental Medicine and Child Neurology 32. 352–55. DOI: 10.1111/j.1469-8749.1990.tb16948.x.CrossRefGoogle ScholarPubMed
Jackendoff, Ray. 1999. Possible stages in the evolution of the language capacity. Trends in Cognitive Sciences 3. 272–79. DOI: 10.1016/S1364-6613(99)01333-9.CrossRefGoogle ScholarPubMed
Jackendoff, Ray. 2002. Foundations of language: Brain, meaning, grammar, evolution. New York: Oxford University Press.CrossRefGoogle Scholar
Jackendoff, Ray. 2011. What is the human language faculty? Two views. Language 87. 586–624. DOI: 10.1353/lan.2011.0063.CrossRefGoogle Scholar
Jarvis, Erich D. 2004. Learned birdsong and the neurobiology of human language. Annals of the New York Academy of Science 1016. 749–77. DOI: 10.1196/annals.1298.038.CrossRefGoogle ScholarPubMed
Jenkins, Lyle. 2000. Biolinguistics: Exploring the biology of language. Cambridge: Cambridge University Press.CrossRefGoogle Scholar
Jenkins, Lyle (ed.) 2004. Variation and universals in biolinguistics. Amsterdam: Elsevier.CrossRefGoogle Scholar
Jenkins, Lyle. 2013a. Biolinguistics: A historical perspective. In Boeckx & Grohmann, 28–43.Google Scholar
Jenkins, Lyle. 2013b. Biolinguistics: Current state and future prospects. English Linguistics 2. 485–508.CrossRefGoogle Scholar
Johnson, Mark. 1988. Attribute-value logic and the theory of grammar. Stanford, CA: CSLI Publications.Google Scholar
Kayne, Richard S. 1994. The antisymmetry of syntax. Cambridge, MA: MIT Press.Google Scholar
Kayne, Richard S. 2005. Some notes on comparative syntax, with special reference to English and French. In Cinque & Kayne, 3–69.Google Scholar
Kayne, Richard S. 2011a. Antisymmetry and the lexicon. In Di Sciullo & Boeckx, 329–53.Google Scholar
Kayne, Richard S. 2011b. Why are there no directionality parameters? West Coast Conference on Formal Linguistics (WCCFL) 28. 1–23. Online: http://www.lingref.com/cpp/wccfl/28/abstract2431.html.Google Scholar
Kimball, John. 1973. Seven principles of surface structure parsing in natural language. Cognition 2. 15–47. DOI: 10.1016/0010-0277(72)90028-5.CrossRefGoogle Scholar
Klein, Felix. 2004 [1939]. Elementary mathematics from an advanced standpoint: Geometry, translated from the German 3rd edn. by E. R. Hedrick and C. A. Noble. Mineola, NY: Dover.Google Scholar
Kobele, Gregory M. 2006. Generating copies: An investigation into structural identity in language and grammar. Los Angeles: University of California, Los Angeles dissertation.Google Scholar
Kolmogorov, A. N. 1965. Three approaches to the quantitative definition of information. Problems in Information Transmission 1. 1–7. DOI: 10.1080/00207166808803030.Google Scholar
Konopka, Genevieve, Bomar, Jamee m., Winden, Kellen, Coppola, Giovanni, Jonsson, Zophonias o., Gao, Fuying, Peng, Sophia, Preuss, Todd m., Wohlschlegel, James a.; and Geschwind, Daniel H.. 2009. Human-specific transcriptional regulation of CNS development genes by FOXP2. Nature 462. 213–17. DOI: 10.1038/nature08549.CrossRefGoogle ScholarPubMed
Kuroda, Reiko. 2015. A twisting story: How a single gene twists a snail? Mechanogenetics. Quarterly Reviews of Biophysics 48. 445–52. DOI: 10.1017/S0033583515000098.CrossRefGoogle ScholarPubMed
Lai, Cecilia S. L., Fisher, Simon E., Hurst, Jane A., Vargha-Khadem, Faraneh; and Monaco, Anthony P.. 2001. A forkhead-domain gene is mutated in a severe speech and language disorder. Nature 413. 519–23. DOI: 10.1038/35097076.CrossRefGoogle Scholar
Legate, Julie A. 2002. Warlpiri theoretical implications. Cambridge, MA: MIT dissertation.Google Scholar
Lenneberg, Eric H. 1967. Biological foundations of language. New York: Wiley.CrossRefGoogle Scholar
Levin, Michael, and Palmer, A. Richard. 2007. Left-right patterning from the inside out: Widespread evidence for intracellular control. BioEssays 29. 271–87. DOI: 10.1002/bies.20545.CrossRefGoogle ScholarPubMed
Lewis, Gwyneth A., Poeppel, David; and Murphy, Gregory L.. 2015. The neural bases of taxonomic and thematic conceptual relations: An MEG study. Neuropsychologia 68. 176–89. DOI: 10.1016/j.neuropsychologia.2015.01.011.CrossRefGoogle ScholarPubMed
Lewontin, R. С. 1974. The genetic basis of evolutionary change. New York: Columbia University Press.Google Scholar
Lewontin, R. С. 2000. The triple helix: Gene, organism, and environment. Cambridge, MA: Harvard University Press.Google Scholar
Li, Rong, and Bowerman, Bruce. 2010. Symmetry breaking in biology. Cold Spring Harbor Perspectives in Biology 2:a003475. DOI: 10.1101/cshperspect.a003475.CrossRefGoogle Scholar
Lightfoot, David. 1984. The language lottery: Toward a biology of grammars. Cambridge, MA: MIT Press.Google Scholar
Lightfoot, David. 1991. How to set parameters: Arguments from language change. Cambridge, MA: MIT Press.Google Scholar
Longobardi, Giuseppe, and Guardiano, Cristina. 2011. The biolinguistic program and historical reconstruction. In Di Sciullo & Boeckx, 266–304.Google Scholar
Longobardi, Giuseppe, Guardiano, Cristina, Silvestri, Giuseppina, tini, Alessio Boat-; and Ceolin, Andrea. 2013. Toward a syntactic phylogeny of modern Indo- European languages. Journal of Historical Linguistics 3. 122–52. DOI: 10.1075/jhl.3.1.07lon.Google Scholar
Longobardi, Giuseppe, and Roberts, Ian. 2010. Universals, diversity and change in the science of language: Reaction to ‘The myth of language universals and cognitive science’. Lingua 120. 2699–2703. DOI: 10.1016/j.lingua.2010.04.006.CrossRefGoogle Scholar
MacDermot, Kay d., Bonora, Elena, Sykes, Nuala, Coupe, Anne-Marie, Lai, Cecilia S., Vernes, Sonja С., Vargha-Khadem, Faraneh, McKenzie, Fiona, Smith, Robert L., Monaco, Anthony P.; and Fisher, Simon E.. 2005. Identification of FOXP2 truncation as a novel cause of developmental speech and language deficits. American Journal of Human Genetics 76. 1074–80. DOI: 10.1086/430841.CrossRefGoogle ScholarPubMed
Magrassi, Lorenzo, Giuseppe, Aromataris, Annovazz-Lodi, Cabrini Valerio; and Moro, Andrea. 2015. Sound representation in higher language areas during language generation. Proceedings of the National Academy of Sciences 18. 1868–73. DOI: 10.1073/pnas.1418162112.Google Scholar
Makuuchi, Michiru, Bahlmann, Jörg, Anwander, Alfred; and Friederici, Angela D.. 2009. Segregating the core computational faculty of human language from working memory. Proceedings of the National Academy of Sciences 106. 8362–67. DOI: 10.1073/pnas.0810928106.CrossRefGoogle ScholarPubMed
Männel, Claudia, Meyer, Lars, Wilcke, Arndt, Boltze, Johannes, Kirsten, Holger; and Friederici, Angela D.. 2015. Working-memory endophenotype and dyslexia- associated genetic variant predict dyslexia phenotype. Cortex 71. 291–305. DOI: 10.1016/j.cortex.2015.06.029.CrossRefGoogle ScholarPubMed
Marcus, Gary F., and Fisher, Simon E.. 2003. FOXP2 in focus: What can genes tell us about speech and language? Trends in Cognitive Sciences 7. 257–62. DOI: 10.1016/S1364-6613(03)00104-9.CrossRefGoogle ScholarPubMed
Miller, George, and Chomsky, Noam. 1963. Finitary models of language users. Handbook of mathematical psychology, vol. 2, ed. by Luce, Robert Duncan, Bush, Robert T., and Galanter, Eugene, 419–92. New York: Wiley.Google Scholar
Montell, Denise J. 2008. Morphogenetic cell movements: Diversity from modular mechanical properties. Science 322. 1502–5. DOI: 10.1126/science.1164073.CrossRefGoogle ScholarPubMed
Moro, Andrea. 2000. Dynamic antisymmetry. Cambridge, MA: MIT Press.Google Scholar
Moro, Andrea, Tettamanti, Μ., Perani, D., Donati, С., Cappa, S. F.; and Fazio, F.. 2001. Syntax and the brain: Disentangling grammar by selective anomalies. NeuroImage 13. 110–18. DOI: 10.1006/nimg.2000.0668.CrossRefGoogle ScholarPubMed
Nevins, Andrew, Pesetsky, David; and Rodrigues, Cilene. 2009. Pirahã exceptionality: A reassessment. Language 85. 355–404. DOI: 10.1353/lan.0.0107.CrossRefGoogle Scholar
Nicoladis, Elena. 2007. Acquisition of deverbal compounds by French-speaking preschoolers. The Mental Lexicon 2. 79–102. DOI: 10.1075/ml.2.1.06nic.CrossRefGoogle Scholar
Niyogi, Partha. 2006. The computational nature of language learning and evolution. Cambridge, MA: MIT Press.CrossRefGoogle Scholar
Niyogi, Partha, and Berwick, Robert С.. 2009. The proper treatment of language acquisition and change in a population setting. Proceedings of the National Academy of Sciences 106. 10124–29. DOI: 10.1073/pnas.0903993106.CrossRefGoogle Scholar
Nóberga, Victor A., and Miyagawa, Shigeru. 2015. The precedence of syntax in the rapid emergence of human language in evolution as defined by the integration hypothesis. Frontiers in Psychology 6:00271. DOI: 10.3389/fpsyg.2015.00271.Google Scholar
Nowak, Martin A., Komarova, Natalia L.; and Niyogi, Partha. 2001. Evolution of universal grammar. Science 291. 114–18. DOI: 10.1126/science.291.5501.114.CrossRefGoogle ScholarPubMed
Palmer, A. Richard. 1996. From symmetry to asymmetry: Phylogenetic patterns of asymmetry variation in animals and their evolutionary significance. Proceedings of the National Academy of Sciences 93. 14279–86.CrossRefGoogle ScholarPubMed
Palmer, A. Richard. 2004. Symmetry breaking and the evolution of development. Science 306. 828–33. DOI: 10.1126/science.1103707.CrossRefGoogle ScholarPubMed
Palmer, A. Richard. 2009. Animal asymmetry. Current Biology 19.R473-R477. DOI: 10.1016/j.cub.2009.04.006.CrossRefGoogle Scholar
Palmer, A. Richard. 2012. Developmental origins of normal and anomalous random right- left asymmetry: Lateral inhibition versus developmental error in a threshold trait. Contributions to Zoology 81. 111–24.Google Scholar
Petrides, Michael, and Pandya, Deepak N.. 1994. Comparative architectonic analysis of the human and the macaque frontal cortex. Handbook of neuropsychology, vol. 9, ed. by Boller, François and Grafman, Jordan, 17–58. Amsterdam: Elsevier.Google Scholar
Piattelli-Palmarini, Massimo, and Vitiello, Giuseppe. 2015. Linguistics and some aspects of its underlying dynamics. Biolinguistics 9. 96–115. Online: http://www.biolinguistics.eu/index.php/biolinguistics/article/view/374.CrossRefGoogle Scholar
Pinker, Steven, and Jackendoff, Ray. 2005. The faculty of language: What's special aboutit? Cognition 95. 201–36. DOI: 10.1016/j.cognition.2004.08.004.CrossRefGoogle Scholar
Progovac, Ljiljana. 2010. Syntax: Its evolution and its representation in the brain. Biolinguistics 4. 234–54. Online: http://www.biolinguistics.eu/index.php/biolinguistics/article/view/161/175.CrossRefGoogle Scholar
Progovac, Ljiljana. 2015. Evolutionary syntax. Oxford: Oxford University Press.CrossRefGoogle Scholar
Progovac, Ljiljana, and Locke, John L.. 2009. The urge to merge: Ritual insult and the evolution of syntax. Biolinguistics 3. 337–54. Online: http://www.biolinguistics.eu/index.php/biolinguistics/article/view/81.CrossRefGoogle Scholar
Reinhart, Tanya. 2006. Interface strategies: Optimal and costly computations. Cambridge, MA: MIT Press.CrossRefGoogle Scholar
Rizzi, Luigi. 2000. Comparative syntax and language acquisition. London: Routledge.Google Scholar
Rizzi, Luigi. 2009. Language invariance and variation. Of minds and language: A dialogue with Noam Chomsky in the Basque country, ed. by Piattelli-Palmarini, Massimo, Uriagereka, Juan, and Salaburu, Pello, 211–20. New York: Oxford University Press.Google Scholar
Ross, Danielle S., and Bever, Tom G.. 2004. The time course for language acquisition in biologically distinct populations: Evidence from deaf individuals. Brain and Language 89. 115–21. DOI: 10.1016/S0093-934X(03)00308-0.CrossRefGoogle ScholarPubMed
Scharff, Constance, and Haesler, Sebastian. 2005. An evolutionary perspective on FoxP2: Strictly for the birds? Current Opinion in Neurobiology 15. 694–703. DOI: 10.1016/j.conb.2005.10.004.CrossRefGoogle ScholarPubMed
Shapiro, Kevin A., Moo, Lauren R.; and Caramazza, Alfonso. 2006. Cortical signatures of noun and verb production. Proceedings of the National Academy of Sciences 103. 1644–49. DOI: 10.1073/pnas.0504142103.CrossRefGoogle ScholarPubMed
Sherwood, Chet c., Broadfield, Douglas c., Holloway, Ralph L., Gannon, Patrick J.; and Hof, Patrick R.. 2003. Variability of Broca's area homologue in African great apes: Implications for language evolution. The Anatomical Record Part A: Discoveries in Molecular, Cellular, and Evolutionary Biology 271A.276–85. DOI: 10.1002/ar.a.l0046.CrossRefGoogle Scholar
Shi, Rushen. 2007. Infants’ recognition of function words in continuous speech. Proceedings of the 16th International Congress of Phonetic Sciences (ICPhS), Saarbrücken, 1541–44. Online: http://www.icphs2007.de/conference/Papers/1296/1296.pdf.Google Scholar
Shi, Rushen, and Lepage, Mélanie. 2008. The effect of functional morphemes on word segmentation in preverbal infants. Developmental Science 11. 407–13. DOI: 10.1111/j.1467-7687.2008.00685.X.CrossRefGoogle ScholarPubMed
Shi, Rushen, Werker, Janet F.; and Cutler, Anne. 2006. Recognition and representation of function words in English-learning infants. Infancy 10. 187–98. DOI: 10.1207/s15327078in1002_5.CrossRefGoogle Scholar
Shieber, Stuart Μ. 1985. Evidence against the context-freeness of natural language. Linguistics and Philosophy 8. 333–43. DOI: 10.1007/978-94-009-3401-6_12.CrossRefGoogle Scholar
Simpson, Jane. 1991. Warlpiri morpho-syntax: A lexicalist approach. Dordrecht: Kluwer.CrossRefGoogle Scholar
Singh, Simon. 1998. Fermat's enigma: The epic quest to solve the worlds greatest mathematical problem. New York: Anchor Books.Google Scholar
Stromswold, Karin. 2007. Genetics and the structure, acquisition and evolution of language. Paper presented at Biolinguistic Investigations, Santo Domingo, Dominican Republic, February 2007.Google Scholar
Stromswold, Karin. 2008. The genetics of speech and language impairments. The New England Journal of Medicine 359. 2381–83. DOI: 10.1056/NEJMe0807813.CrossRefGoogle ScholarPubMed
Stromswold, Karin. 2010. Genetics and the evolution of language: What genetic studies reveal about the evolution of language. Evolution of language: Biolinguistic perspectives, ed. by Larson, Richard K., Déprez, Viviane, and Yamakido, Hiroko, 176–93. Cambridge: Cambridge University Press.Google Scholar
Sun, Tao, and Walsh, Christopher A.. 2006. Molecular approaches to brain asymmetry and handedness. Nature Reviews Neuroscience 7. 655–62. DOI: 10.1038/nrn1930.CrossRefGoogle ScholarPubMed
Thompson, D’Arcy Wentworth. 1942. On growth and form. New York: Dover.Google Scholar
Turing, Alan Μ. 1952. The chemical basis of morphogenesis. Philosophical Transactions of the Royal Society of London B 237. 37–72. DOI: 10.1098/rstb.1952.0012.Google Scholar
Ullman, Shimon. 1979. The interpretation of visual motion. Cambridge, MA: MIT Press.CrossRefGoogle Scholar
Ullman, Shimon. 1996. High-level vision: Object recognition and visual cognition. Cambridge, MA: MIT Press.CrossRefGoogle Scholar
Ullman, Shimon. 2006. Object recognition and segmentation by a fragment-based hierarchy. Trends in Cognitive Sciences 11. 58–64. DOI: 10.1016/j.tics.2006.11.009.CrossRefGoogle ScholarPubMed
Uriagereka, Juan. 2012. Spell-out and the minimalist program. Oxford: Oxford University Press.Google Scholar
van Rhijn, Jon-Ruben, and Vernes, Sonja C.. 2015. Retinoic acid signaling: Anew piece in the spoken language puzzle. Frontiers in Psychology 6:01816. DOI: 10.3389/fpsyg.2015.01816.CrossRefGoogle Scholar
Wexler, Kenneth. 2003. Lenneberg's dream: Learning, normal language development and specific language impairment. Language competence across populations: Toward a definition of specific language impairment, ed. by Levi, Yonata and Schaeffer, Jeannette C., 11–61. Mahwah, NJ: Lawrence Erlbaum.Google Scholar
Xiao, Yaqiong, Friederici, Angela D., Margulies, Daniel S.; and Brauer, Jens. 2016. Development of a selective left-hemispheric fronto-temporal network for processing syntactic complexity in language comprehension. Neuropsychologia 83. 274–82. DOI: 10.1016/j.neuropsychologia.2015.09.003.CrossRefGoogle Scholar
Yang, Charles D. 2002. Knowledge and learning in natural language. Oxford: Oxford University Press.Google Scholar
Yang, Charles D. 2004a. Toward a theory of language growth. In Jenkins 2004, 37–56.Google Scholar
Yang, Charles D. 2004b. Universal grammar, statistics or both? Trends in Cognitive Sciences 8. 451–56. DOI: 10.1016/j.tics.2004.08.006.CrossRefGoogle Scholar
Yang, Charles D. 2008. The great number crunch. Journal of Linguistics 44. 205–28. DOI: 10.1017/S0022226707004999.CrossRefGoogle Scholar
Yang, Charles D. 2011. Usage unevenness in child language supports grammar productivity. Paper presented at the Boston University Conference on Language Development (BUCLD), Philadelphia, University of Pennsylvania.Google Scholar
Yang, Charles D. 2013. Who 's afraid of George Kingsley Zipf? Or: Do children and chimps have language? Significance 10. 29–34. DOI: 10.ni1/j.1740-9713.2013.00708.x.CrossRefGoogle Scholar
Yang, Charles D. 2015. Negative knowledge from positive evidence. Language 91.93853. DOI: 10.1353/lan.2015.0054.CrossRefGoogle Scholar
Zabbal, Youri. 2005. The syntax of numeral expressions. Amherst: University of Massachusetts Amherst, ms.Google Scholar
Zaccarella, Emiliano, and Friederici, Angela D.. 2015. Merge in the human brain: A sub-region based functional investigation in the left pars opercularis. Frontiers in Psychology 6:01818. DOI: 10.3389/fpsyg.2015.01818.CrossRefGoogle ScholarPubMed
Zeijlstra, Hedde. 2008. On the syntactic flexibility of formal features. In Biberauer, 143–75.Google Scholar
Zwart, J. W. 2011. Structure and order: Asymmetric Merge. The Oxford handbook of linguistic minimalism, ed. by Boeckx, Cedric, 96–118. Oxford: Oxford University Press.Google Scholar