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Quaternary dryland dynamics: perspectives and prospects

Published online by Cambridge University Press:  16 July 2025

Abi Stone*
Affiliation:
Department of Geography, The University of Manchester , M13 9PL, UK
*
Corresponding author: Abi Stone; Email: abi.stone@manchester.ac.uk
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Abstract

Reconstructing Quaternary-timescale environmental change in drylands provides insights into styles and rates of change in response to direct insolation forcing and variations in global temperature, ice volume and sea level. Changes to the relatively inhospitable environments presented by drylands are also central to debates about the migration and adaptability of hominin species. This review outlines approaches used for reconstructing past environments, which use dated sequences of environmental proxies – the properties of physical sediments, chemical precipitates and biological materials. In addition, climate model simulations can explore responses to known climatic forcing factors. Advances in both approaches remain situated within conceptual frameworks about dryland responses to: (i) cyclical changes in the Earth’s orbit around the sun, either mediated via the global cryosphere during glacial–interglacial cycles (~100 ka periodicity during the last ~0.9 Ma), or mediated via the response of the global monsoon with ~23 ka periodicity (precession) and (ii) millennial-scale climatic shifts, which are thought to originate outside drylands, within the North Atlantic.

In this review, three examples are outlined to demonstrate areas of emerging consensus and remaining contradictions: (1) speleothem and tufa growth records that span hundreds of thousands of years (ka), (2) conditions at the last glacial maximum (LGM) and (3) proxies recording millennial-scale events. Precessional-scale forcing of the monsoon is not always observed, with apparent mediating roles from glacial boundary conditions in parts of the northern hemisphere (NH) and from interglacial boundary conditions in parts of the southern hemisphere (SH). The LGM in drylands was initially conceptualised as experiencing pluvial (wet) conditions, which shifted to a glacial aridity paradigm, and is again shifting to a pattern of global heterogeneity, in which some drylands were wetter-than-present and others drier. However, there remain contradictions between environmental proxies and model simulations, and spatial heterogeneity is observed within many drylands. Dryland proxies that record millennial-scale events demonstrate clearly the importance of climatic teleconnections across the globe in influencing dryland hydroclimate. The records of Quaternary dryland change across a range of timescales get used alongside simulations of ecosystem response through time, hominin habitat and even hominin physiology to better understand likely hominin dispersals through dryland regions.

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Review
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This is an Open Access article, distributed under the terms of the Creative Commons Attribution-NonCommercial-NoDerivatives licence (http://creativecommons.org/licenses/by-nc-nd/4.0), which permits non-commercial re-use, distribution, and reproduction in any medium, provided that no alterations are made and the original article is properly cited. The written permission of Cambridge University Press must be obtained prior to any commercial use and/or adaptation of the article.
Copyright
© The Author(s), 2025. Published by Cambridge University Press
Figure 0

Table 1. Archives and their proxies for past climatic and environmental conditions in drylands, giving an overview of archives and their proxies for past climatic and environmental conditions in drylands, and key modelling simulations from which changes in dryland aridity can be inferred. (abbreviations are used: N – north, N-cen – north central, NW – northwest, S – south, Sn – southern, W – West, Wn – western, Kal – Kalahari)

Figure 1

Figure 1. Map of current global dryland distribution, alongside the locations of the major proxy records explored in the three key examples in this paper and an inset of the TraCE-21 ka simulation for the LGM from Liu et al. (2021) demonstrating which dryland regions are simulated to be wetter-than-present, or drier-than-present. Site numbers (s) are referred to in the text, and (1) North Atlantic dust records, ODP659 (deMenocal, 1995), MD03–2705 and ODP659 (Skonieczny et al., 2019), (2) Mukalla Cave (Nicholson et al., 2020), (3) Hoti Cave (Nicholson et al., 2020), (4) North Arabian Sea dust record ODP721/722 (deMencoal, 1995), (5) Red Sea dust record K11 (Ehrmann et al., 2024), (6) Gulf of Aden RC09–166 (Tierney et al., 2017), (7) Wadi Dabsa tufa record (Stone et al., 2023), (8) Soreq Cave (Bar-Matthews et al., 1997), (9) Egyptian tufa growth (Kele et al., 2021), (10) Rössing Cave, Namibia (Geyh and Heine, 2014), (11) Tswaing Crater (Partridge et al., 1997), (12) CD15410–06 (Simon et al., 2015), (13) Naracoorte Caves (Weij et al., 2024), (14) Leeuwin-Naturaliste-region caves (Weij et al., 2024), (15–17) speleothems on the northwest seaboard of Australia (Denniston et al., 2017), (18) Kati Thanda-Lake Eyre Basin, (19) Murray Darling Basin, (20) Tasman Sea dust record (Hesse, 1994), (21) Native Companion Lagoon, eastern Australia (Petherick et al., 2008), (22) Lake Chilwa, Malawi (Thomas et al., 2009), (23) Pella hydrax midden (Chase et al., 2019), (24) Zizou hyrax midden (Chase et al., 2019), (25) Spitzkoppe midden (Chase et al., 2019), (26) De Rif hyrax midden (Chase et al., 2011).

Figure 2

Figure 2. Key proxy records globally and for drylands, where: (A) the LR04 oxygen isotope record, with Marine Oxygen Isotope Stages (MIS) above (Lisiecki and Raymo, 2005), (B) Antarctica Vostok (Petit et al., 1999) and EPICA DomeC (Lambert et al., 2012) dust records, (C) North Atlantic dust records, ODP659 (deMenocal, 1995) and 230Th corrected MD03–2705 and ODP659 (Skonieczny et al., 2019) (sites near 1), (D) North Arabian Sea dust record ODP721/722 (deMencoal, 1995) (site 4), (E) Red Sea dust record K11 (Ehrmann et al., 2024) (site 5), (F) Gulf of Aden RC09–166 terrestrial leaf wax δD (Tierney et al., 2017) (site 6), (G) speleothem-growth record derived South Arabian Humid Periods (SAHPS) (Nicholson et al., 2020) (sites 2 and 3), (H) Soreq Cave δ18O record (Bar-Matthews et al., 1997) (site 8), (I) Tufa growth records from Wadi Dabsa, SW Arabia (Stone et al., 2023) (site 7) and N Africa (summarised in Kele et al., 2021) (within box 9), (J) Rössing Cave speleothem growth record (Geyh and Heine, 2014) (site 10), (K) Tswaing Crater rainfall proxy (Partridge et al., 1997) (site 11), (L) CD154–10-06P southwest Africa humidity record derived from Fe/K ratios for highly eroded terrestrial sediment (Simon et al., 2015) (site 12), (M) KDE (Kernel Density Estimates) for speleothem U-Th ages using 5 ka bandwidth for Naracoorte (east) (site 13) and Leeuwin-Naturliste region (west) (site 14) speleothems in southern Australia (Weij et al., 2024), and (N) insolation curves for summer in each hemisphere to demonstrate precession-paced forcing, where solid orange line is 30°N June and dashed red line is 30°S December (from Laskar et al., 2004).

Figure 3

Figure 3. (A) Timing of highstands of Lake Chilwa from luminescence dating of shorelines, compared to the timing of Heinrich Events, (B) HadCM3 simulation of terrestrial moisture balance over Africa caused by a ‘freshwater hosing’ 50–70°N to simulate a North Atlantic Heinrich Event, displayed as an anomaly (modelled years 275–300 minus the average of 0–100 years), where the inset shows modelled reduction in Atlantic Meridional Overturning Circulation (modified from Thomas et al., 2009).

Figure 4

Figure 4. Hominin species distributions, expressed as time-averaged habitat suitability (intensity of blue shading) over 2 million years from Timmerman et al. (2022), also showing (with coloured circles) the locations of fossil and/or archaeological artefact evidence associated with each of the five hominin groups, collated by Timmerman et al. (2022).

Author comment: Quaternary dryland dynamics: perspectives and prospects — R0/PR1

Comments

Dear Professors David Eldrdige and Osvaldo Sala

Please find the manuscript “Quaternary Dryland Dynamics: perspectives and prospects” (of 5,000 words), and with 1 table and 3 figures for consideration for peer review for your journal. It is designed as a Review (Stand Review) article type.

This is written in response to the invitation in September 2024 from Laetitia Beck (Senior Editorial Assistant, Open Access, STM Journals, Academic) on behalf of Professors Osvaldo Sala and David Eldridge “to author a commissioned article for the journal on a topic of your choice. Drylands provides a home for high-impact, globally relevant reviews and original research on all topics associated with the natural sciences, engineering, social sciences, and management of global drylands.”, which was confirmed on 20-Dec-2024 by Kim Marello (Cambridge Prisms: Drylands Editorial Office) “Dear Dr. Stone,

Thank you for agreeing to submit the manuscript with the working title “” to the Cambridge Prisms: Drylands…”

In identifying Key Words and Topics on the online system, I note there is not anything labelled as “Quaternary” or “Past Climate” and this might be beneficial as additional fields for the journal in the future, in order to encourage engagement with the journal from the palaeo-Drylands community.

Yours sincerely

Dr Abigail (Abi) Stone

Reader in Physical Geography

Review: Quaternary dryland dynamics: perspectives and prospects — R0/PR2

Conflict of interest statement

Reviewer declares none.

Comments

This is all very very confused. I am really not clear what the purpose of this study is. If it is a literature review, say so, and be very specific on what it focuses on. The abstract talks about climate model simulations but no results of this are presented. The introduction lacks focus, I am really not clear what the main point is. CMIP is not mentioned. “exploring the timescales over which these changes have been investigated” makes no sense. Name the different drylands examined. Table 1, what is Sn? Section on climate in the Quaternary does not mention any evidence at all, the sections later on dealing with precession and millennial-scale need to be moved here. The back half of the paper dissipates and doesn’t go anywhere. The link to hominins really doesn’t work and is just an add-on, so take this out. Overall none of this works, it needs to be done again, so please find a FOCUS first and structure the paper accordingly.

Review: Quaternary dryland dynamics: perspectives and prospects — R0/PR3

Conflict of interest statement

None

Comments

This is an excellent treatment of global dryland dynamics which will be of immediate interest to archaeologists trying to understand why and how different hominins have adapted to specific environments over the last 2 Ma. The discussion of proxies for reconstructing past climatic and environmental conditions is thorough, as is the treatment of different scales of Quaternary climatic fluctuations. The section on hominin response to Quaternary climate change is useful and salutary, inevitably noting that local processes must always be considered beyond larger climate drivers. The hub of the critical analysis appears at page 11 where it is noted that “There remain many details to be understood and resolved, particularly interpreting which combination of precipitation and/or temperature changes is responsible for driving hydroclimatic response, and understanding why different drylands undergoing similar forcing within the same hemisphere of the Earth, or even the same continental mass, responded differently”. An example is then given of how global glacial phases will not necessarily be accompanied by ‘pluvial’ events. I feel this discussion about the need to focus on regional and local context should be expanded on here with at least one or two additional examples. The current Conclusions do not discuss hominin dispersals and yet these are mentioned in the abstract and main body of the text. Reference should be made to the recent paper by Mercarder et al. 2025 outlining the evidence for 1 Ma occupation of the Oldupai Gorge environs when hyper arid (Mercarder, J., Akuku, P., Boivin, N. et al. Homo erectus adapted to steppe-desert climate extremes one million years ago. Communications earth and environment (2025)6:1). It appears that erectus and archaic sapiens were arid adapted and that deserts have not necessarily represented biogeographic barriers. This observation is directly relevant to ‘green corridor’ arguments for Africa and the Arabian Peninsula.

There are some additional comments/typos, as follow:

L 27 Should read ‘particularly’

P3 L 17 Add Williams, A.N., Veth, P., Steffen, W., Ulm, S., Turney, C.S.M., Phipps, S., Smith, M., Reeves, J. 2015 A Continental Narrative: Human Settlement Patterns and Australasian Climate Change over the last 35,000 Years. Quaternary Science Reviews 123: 91 – 112 as an example of archaeological patterning and drylands at the continental scale.

P 3 L 53 Author is Jankowski et al. and misspelt elsewhere

P 5 L 30 Should read Monsoon

P 7 L 49 - 51 This sentence needs to be rephrased so it is clear that there are no age age clusters recorded for dune building over 120 ka; and specifically not for the LGM (c. 24 - 18 ka).

Review: Quaternary dryland dynamics: perspectives and prospects — R0/PR4

Conflict of interest statement

Reviewer declares none.

Comments

The paper reviews the driving mechanisms and spatiotemporal patterns of environmental changes in Quaternary arid regions, with a focus on exploring the impact of orbital scales (glacial-interglacial cycles, precession cycles) and millennium scale events on global arid hydrological and climatic conditions. Combined with proxy indicators and climate model simulations, the response of arid systems to climate forcing is analyzed. The article reveals the heterogeneity of dryland environments through multiple regional cases (Arabian Peninsula, Sahara, Australia, Southern Africa, etc.) and looks forward to the research prospects of the correlation between ancient human migration and climate change.

Main issues and suggestions:

1. As a review article addressing arid land climate change, the authors should include a map illustrating the spatial extent of global drylands to provide readers with an intuitive geographic context for the regions under discussion. If the boundaries of arid zones vary across different climatic periods, presenting the current distribution would serve as a critical reference baseline. Against this background, the locations of the paleoclimate reconstruction records listed in Table 1 could be overlaid on this map to spatially contextualize the proxy data discussed in the text.

2. The dryland change during the Quaternary has been extensively simulated. These modelling studies have elucidated response mechanisms under various climate scenarios with various external forcings. However, this article provides limited coverage of this topic. Below are some examples, and it is suggested that the authors enhance the summary of the advancements in simulation research.

Greve 2017 Simulated changes in aridity from the last glacial maximum to 4xCO2

Liu 2018 A multi-model analysis of moisture changes during the last glacial maximum

Liu 2021 Time-varying responses of dryland aridity to external forcings over the last 21 ka

Liu 2019 Mid-Holocene drylands A multi-model analysis using Paleoclimate Modelling Intercomparison Project Phase III (PMIP3) simulations

Xu 2020 The PMIP3 Simulated Climate Changes over Arid Central Asia during the Mid-Holocene and Last Glacial Maximum

3. The authors’ descriptions for each period lack unified conclusions regarding the dryland changes, with excessive reliance on compiling existing literature instead of synthesizing critical insights. The author may consider categorizing chapters based on different arid regions, rather than different time periods. In this way, conclusions may be better obtained.

Minor Issues:

Page 1 Line 21: “Unpicking the changes to atmospheric and oceanic circulation systems that may have driven the hydroclimatic response of drylands at each of these scales develops via an interrogation of the proxy record in both the terrestrial and the marine sedimentary record, alongside the use of numerical climate model simulations.” This sentence doesn’t seem to make sense. “Unpecking” itself does not “develop”.

Page 1 Line 41: “The emplacement and migration of dune features XXX influences XXX” should be “influence”. Please address similar issues in other paragraphs as well.

Table 1: “photsynthetic” should be “photosynthetic”. Please address similar issues in other paragraphs as well.

Figure: Figure 1a and Figure 3 are not cited in the text. The citing of figures is not consistent. On page 10, “Figure 2B” is used; In other cases, “Fig. x” is used. On pages 10 and 11, the author leaves “Figure x” in the paper to let me guess.

Recommendation: Quaternary dryland dynamics: perspectives and prospects — R0/PR5

Comments

Dear Dr Stone,

Thank you for submitting your paper to the journal. We have now received 3 reviews and, as you will see, the reviewers suggest some changes to the manuscript before it can be published. I especially invite you to consider comments 2 and 3 by rev. #3.

Thank you again for submitting your work to Drylands.

Best regardard.

Decision: Quaternary dryland dynamics: perspectives and prospects — R0/PR6

Comments

No accompanying comment.

Author comment: Quaternary dryland dynamics: perspectives and prospects — R1/PR7

Comments

Dear Professors David Eldrdige and Osvaldo Sala

Please find the revised manuscript “Quaternary Dryland Dynamics: perspectives and prospects”, which is revised in line with the comments of the three reviewers.

This is written in response to the invitation in September 2024 from Laetitia Beck (Senior Editorial Assistant, Open Access, STM Journals, Academic) on behalf of Professors Osvaldo Sala and David Eldridge “to author a commissioned article for the journal on a topic of your choice.

This review provides an entry point for understanding the major subtopic of Quaternary drylands. The paper provides a broad, introductory overview of this topic, and outline some significant areas in the current state of knowledge, areas of uncertainty and future directions. The paper uses three key examples that exemplify advancements in the field, that illustrate evolving interpretations, significant developments in thought, and areas where uncertainty persists. The paper emphasizes the complexity and variability of dryland changes—both across different regions and within individual landscapes—evident in both reconstructions and model simulations. In its final section, the review explores recent progress in understanding how hominins responded to climatic and environmental shifts in drylands during the Quaternary. The interplay between environmental change and hominin adaptation, migration, and evolution is currently revitalizing this area of dryland research.

I note that the file upload window didn’t offer clearly the opportunity to upload the track changes version of the manuscript. I did not add it as a second “main manuscript”. I am very happy to provide the track changes version of the manuscript over email as required.

Yours sincerely

Dr Abi Stone

Review: Quaternary dryland dynamics: perspectives and prospects — R1/PR8

Conflict of interest statement

Reviewer declares none.

Comments

The authors have addressed the major concerns raised in the initial review. The revised manuscript presents a clearer structure and improved articulation of key themes. The inclusion of additional examples, clarification of the role of model simulations, and more explicit summary statements enhance the overall coherence of the paper.

I find the revisions satisfactory and have no further substantive comments. I recommend that the manuscript be accepted for publication.

Recommendation: Quaternary dryland dynamics: perspectives and prospects — R1/PR9

Comments

Dear Dr Stone,

First of all, please accept my apologies for the long delay in getting back to you. I know how frustrating it is to wait for months for a decision on a paper submitted, especially after revision.

I have now received comments on the revised version and I am happy to report that the reviewers were fully satisfied with the changes made. The paper can now be accepted.

Thank you again for submitting your work to the journal and for your incredible patience.

Hope to read more of your work soon.

Best regards,

Decision: Quaternary dryland dynamics: perspectives and prospects — R1/PR10

Comments

No accompanying comment.