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  • R. R. Mill (a1)

The living species of the genus Retrophyllum C.N. Page (Podocarpaceae) are revised. A key to the six species recognised is given. Retrophyllum filicifolium (N.E.Gray) R.R.Mill comb. nov. from New Guinea and the Moluccas is recognised as a species distinct from R. vitiense (Seemann) C.N. Page, which is restricted to material from Fiji and the Solomon Islands. Additional characters are given by which to separate Retrophyllum piresii (Silba) C.N. Page (Brazil) from R. rospigliosii (Pilg.) C.N. Page (Andes). The six species fall into three species-pairs that differ in reproductive characters: one pair in Papuasia and Melanesia, a second on New Caledonia and a third in South America. In the New Caledonian species-pair, the adult leaves are flattened in four ranks with only minimal heterofacial twisting, whereas in the species-pair inhabiting Fiji, New Guinea and neighbouring areas, as well as the pair inhabiting South America, the adult leaves are arranged in two ranks with significant heterofacial twisting. The names Podocarpus filicifolius N.E.Gray and Nageia minor Carrière, respectively the basionyms of Retrophyllum filicifolium and R. minus, are lectotypified, and the typification of Nageia minor comprehensively discussed in an appendix. Adult female epitypes are additionally designated for Podocarpus filicifolius, which was originally based on juvenile foliage of Retrophyllum mixed with a detached seed of Nageia wallichiana, and for Nageia minor, which Carrière described on the basis of sterile material. Two other appendices provide a list of accepted names and synonyms, and a list of exsiccatae. Illustrations and distribution maps are provided for each species.

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Abdillahi H. S., Stafford G. I., Finnie J. F. & Van Staden J. (2010). Ethnobotany, phytochemistry and pharmacology of Podocarpus sensu latissimo (s.l.). S. African J. Bot. 76: 124.
Amaro-Luis J. M. & Carroz U. D. (1988). Norditerpene dilactones from Decussocarpus rospigliosii. J. Nat. Prod. 51 (6): 12491250.
Amaro Luis J. M., Amesty Á., Montealegre R. & Bahsas A. (2006). Rospiglioside, a new totarane diterpene from the leaves of Retrophyllum rospigliosii. J. Mex. Chem. Soc. 50 (3): 9699. Online. Available: (downloaded 10 February 2011).
Amaro-Luis J. M., Amesty A., Bahsas A. & Montealegre R. (2008; published online 2007). Biflavones from the leaves of Retrophyllum rospigliosii. Biochem. Syst. Ecol. 36 (3): 235237.
Aranguren A. & Márquez N. J. (2011). Etnoecología de las especies vegetales de los bosques estacionalmente secos del Estado Mérida. Ethnobot. Res. Applic. 9: 307323. Online. Available: (downloaded 8 February 2012).
Ataroff S. M. (2002). Precipitación e intercepción en ecosistemas boscosos de los Andes Venezolanos. [Precipitation and interception in forest ecosystems of the Venezuelan Andes.] Ecotropicos 15 (2): 195202. Online. Available: (downloaded 23 November 2005).
Barragan Bedoya J. M. & Valdés Torres R. D. (2011). Lineamientos de participación comunitaria en el manejo de un área protegida, como alternativa de manejo incluyente del territorio: una aproximación desde el conflicto ambiental por la conservación de la naturaleza en la cuenca media del Río Otún. Pereira: Programa Administración Ambiental, Universidad Tecnológica de Pereira.
Barre N., Wichatitsky M. de G., Lecoq R. & Maillard J.-C. (2003). Contribution to the knowledge of the New Caledonian imperial pigeon Ducula goliath (Gray 1859) with emphasis on sexual dimorphism. Notornis 50 (3): 155160.
Barrie F. R. (2006). Report of the General Committee: 9. Taxon 55 (3): 795800.
Becerra-Montalvo V. & Zevallos-Pollito P. A. (2014). Determinación del turno de corta de Cedrela odorata L., Retrophyllum rospigliosii Pilger y Prumnopitys harmsiana Pilger a través del estudio dendrocronológico en San Ignacio, región Cajamarca-Perú. El Ceprosimad 02 (2): 3347.
Benny G. L., Samuelson D. A. & Kimbrough J. W. (1985). Studies on the Coryneliales. III. Taxa parasitic on Podocarpaceae: Lagenulopsis and Tripospora. Bot. Gaz. 146 (3): 431436.
Berry E. W. (1922). The flora of the Concepción-Arauco Coal Measures of Chile. Johns Hopkins Univ. Stud. Geol. 4 (3): 73143.
Bertrand C. E. (1874). Anatomie comparée des tiges et des feuilles chez les Gnétacées et les Conifères. Ann. Sci. Nat., Bot., Sér. 5 20: 5153.
Biffin E., Conran J. G. & Lowe A. J. (2011). Podocarp evolution: a molecular phylogenetic perspective. In: Turner B. L. & Cernusak L. A. (eds) Ecology of the Podocarpaceae in Tropical Forests, pp. 120. Smithsonian Contributions to Botany, no. 95. Washington, District of Columbia: Smithsonian Institution Scholarly Press.
Biffin E., Brodribb T. J., Hill R. S., Thomas P. & Lowe A. J. (2012). Leaf evolution in Southern Hemisphere conifers tracks the angiosperm ecological radiation. Proc. Roy. Soc. Biol. Sci. Ser. B 279 (1727): 341348.
Blackburn D. T. (1981). Tertiary megafossil flora of Maslin Bay, South Australia: numerical study of selected leaves. Alcheringa 5: 928.
Bobrov A. V. F. C. & Kostrikin D. S. (2000). Sistematicheskoe polozhenie roda Retrophyllum C.N. Page Podocarpaceae (Dumort.) Endl. s.l. i filogeniya semeistva Nageiaceae Fu s. l. Problemy filogenii izbrannykh grupp vysshikh rastenij, Sochi, 2000, 1: 1012.
Botero-Delgadillo E. & Páez C. A. (2011). Estado actual del conocimiento y conservación de los loros amenazados de Colombia. Conservación Colombiana no. 14: 86151.
Brako L., & Zarucchi J. L. (1993). Catalogue of the flowering plants and gymnosperms of Peru. Monogr. Syst. Bot. Missouri Bot. Gard. 45. St. Louis, Missouri: Missouri Botanical Garden.
Brongniart A. & Gris A. (1869). Nouvelle note sur les Conifères néo-calédoniennes. Bull. Soc. Bot. France 16: 325331.
Brongniart A. & Gris A. (1871). Observations sur diverses plantes nouvelles ou peu connues de la Nouvelle Calédonie (suite). Supplément aux Conifères (2). Ann. Sci. Nat., Bot. Sér. 5 13: 340404.
Brummitt R. K. (2001). World Geographical Scheme for Recording Plant Distributions, Edition 2. Plant Taxonomic Database Standards No. 2. Pittsburgh, Pennsylvania: Hunt Institute for Botanical Documentation, Carnegie Mellon University, for the International Working Group on Taxonomic Databases for Plant Sciences.
Brummitt R. K. (2006 [‘2005’]). Report of the Committee for Spermatophyta: 57. Taxon 54 (4): 10931103.
Brummitt R. K. (2011). Report of the Nomenclature Committee for Vascular Plants: 63. Taxon 60 (4): 12021210.
Buchholz J. T. (1949). Additions to the coniferous flora of New Caledonia. Bull. Mus. Natl. Hist. Nat. (Paris), Sér. 2 21 (2): 279286.
Buchholz J. T. (1955). Conifères recoltées en Nouvelle Calédonie. Bull. Mus. Natl. Hist. Nat. (Paris), Sér. 2 27: 151152. [Posthumous, with introduction by A. Guillaumin.]
Buchholz J. T. & Gray N. E. (1948). A taxonomic revision of Podocarpus. I. The sections of the genus and their subdivisions with special reference to leaf anatomy. J. Arnold Arbor. 29: 4963.
Buchholz J. T. & Gray N. E. (1957). Podocarpaceae. In: Steyermark J. al. Flora of Venezuela. Fieldiana, Bot. 28 (4): 759772. Chicago, Illinois: Chicago Natural History Museum.
Butin H. (1970). Drei neue koniferenbewohnende Hypodermataceae aus Chile. Phytopathol. Z. 68: 6372.
Cambie R. C., Cox R. E., Croft K. D. & Sidwell D. (1983). Phenolic diterpenoids of some podocarps. Phytochemistry 22: 11631166.
Cambie R. C., Cox R. E. & Sidwell D. (1984). Phenolic diterpenoids of Podocarpus ferrugineus and other podocarps. Phytochemistry 23: 333336.
Carlquist S. (1974). Chapter 7, Adaptive radiation: New Caledonia and New Zealand. In: Carlquist S.Island Biology. New York, New York: Columbia University Press.
Carrière É.-A. (1867). Traité général des Conifères. Nouvelle édition. Paris: chez l'auteur, rue de Buffon 53.
Ceballos-Freire A. J. & López-Ríos J. A. (2007). Conservación de la calidad de semillas forestales nativas en almacenamiento. Cenicafé 58 (4): 265292.
Chaabi M., Antheaume C., Weniger B., Justiniano H., Lugnier C. & Lobstein A. (2007). Biflavones of Decussocarpus rospigliosii as phosphodiesterases inhibitors. Pl. Med. (Stuttgart) 73 (12): 12841286.
Chevalier L. (1957). Les conifères actuellement connus en Nouvelle Calédonie. Études Mélanés. n.s. 8e/9e année, 10/11 (II(1)): 105–118.
Compton R. H. (1922). A systematic account of the plants collected in New Caledonia and the Isle of Pines by Mr R. H. Compton, M. A., in 1914. – Part II. Gymnosperms and Cryptogams. By Prof. Compton and others. J. Linn. Soc. London, Bot. 45 (304): 421462.
Conn B. J., Lee L. L. & Kiapranis R. (2004 onwards). PNGplants Database: Plant Collections from Papua New Guinea. Online. Available: (accessed 18 December 2014).
Conran J. G., Wood G. G., Martin P. G., Dowd J. M., Quinn C. J., Gadek P. A. & Price R. A. (2000). Generic relationships within and between the Gymnosperm families Podocarpaceae and Phyllocladaceae based on an analysis of the chloroplast gene rbcL. Austral. J. Bot. 48: 715724.
Cornu A., Sarrailh J.-M. & Marion F. (2001). Espèces endémiques et restauration écologique en Nouvelle-Calédonie. Bois Foréts Trop. 268 (2): 5769.
de Kok R. P. J. & Mabberley D. J. (1999). A synopsis of Oxera Labill. (Labiatae). Kew Bull. 54 (2): 265300.
de Laubenfels D. J. (1968 [‘1967’]). Podocarpus vitiensis in the Moluccas (Taxaceae). Blumea 15 (2): 440.
de Laubenfels D. J. (1969). A revision of the Malesian and Pacific rainforest conifers, I. Podocarpaceae, in part. J. Arnold Arbor. 50: 274369. [Published in two consecutive issues as pp. 274–314 (15 Apr 1969) and pp. 315–369 (15 Jul 1969).]
de Laubenfels D. J. (1972). Gymnospermes. In: Aubreville A. & Leroy J.-F. (eds) Flore de la Nouvelle-Calédonie et Dépendances, 4. Paris: Muséum national d'histoire naturelle.
de Laubenfels D. J. (1982). Podocarpaceae. In: Luces de Febres Z. & Steyermark J. A. (eds) Flora de Venezuela 11 (2): 741. Caracas: Instituto Nacional de Parques, Dirección de Investigaciones Biológicas, Ediciones Fundación Educación Ambiental.
de Laubenfels D. J. (1987). Revision of the genus Nageia (Podocarpaceae). Blumea 32: 209211.
de Laubenfels D. J. (1988). Coniferales. In: van Steenis C. G. G. J. & De Wilde W. J. J. O. (‘1986’). Flora Malesiana 10 (3). Dordrecht: Kluwer Academic Publishers.
de Laubenfels D. J. (1994). Las Podocaráceas [sic] del Perú. Bol. Lima 16 (91–96): 3539.
de Oliveira D. M. (2012). Retrophyllum piresii (Silba) C.N. Page. Rio de Janeiro: Instituto de Pesquisas Jardim Botãnico do Rio de Janeiro. Online. Available: (downloaded 29 October 2015).
Degener O. & Degener I. (1953). Nutzpflanzen der Eingeborenen von Fidschi. Mitt. Bot. Gart. Mus. Berlin-Dahlem 1 (1): 131150.
Dinerstein E., Olson D. M., Graham D. J., Webster A. L., Primm S. A., Bookbinder M. P. & Ledec G. (1995). A Conservation Assessment of the Terrestrial Ecoregions of Latin America and the Caribbean. Washington, District of Columbia: The World Bank.
Dorr L. J., Stergios B., Smith A. R. & Cuello A. N. L. (2000). Catalogue of the Vascular Plants of Guaramacal National Park, Portuguesa and Trujillo States, Venezuela. Contrib. U. S. Natl. Herb. 40: 1155. Washington, District of Columbia: Smithsonian Institution.
Doyle M. F. (1999). Regional action plan: conifers of the Oceanic Islands in the insular South Pacific (Fiji, Tonga, Solomon Islands and Vanuatu). In Farjon A. & Page C. N. (comp.) Conifers. Status Survey and Conservation Action Plan, pp. 7274. Gland, Switzerland, and Cambridge: IUCN/SSC Conifer Specialist Group, IUCN.
Eckenwalder J. E. (2009). Conifers of the World: the Complete Reference. Portland, Oregon: Timber Press.
ESRI (2008). ArcGIS 9.3. Redlands, California: Environmental Systems Resource Institute.
Evans W. P. (1937). Note on the flora which yielded the Tertiary lignites of Canterbury, Otago, and Southland. New Zealand J. Sci. Technol. 19: 188193.
Farjon A. (2010). A Handbook of the World's Conifers. Leiden: E.J. Brill.
Feuillet Hurtado C., Macias Pinto D. & Chito Cerón E. (2011). Plantas útiles para la elaboración de artesianías en el Departamento de Cauca (Colombia). Bol. Cient. Mus. Hist. Nat. 15 (2): 4059.
Florin R. (1940). The Tertiary fossil conifers of South Chile and their phytogeographical significance (with a review of the fossil conifers of southern lands). Kungl. Svenska Vetenskapsakad. Handl. 19: 1107.
Furman T. E. (1970). The nodular mycorrhizae of Podocarpus rospigliosii. Amer. J. Bot. 57 (8): 910915.
Fuzzy Gazetteer (no date). Online. Available: (accessed 22 July 2014 etc.)
Gardner M. (2013). Retrophyllum piresii. The IUCN Red List of Threatened Species 2013: e.T34109A2846432. Online. Available: (downloaded 29 October 2015).
Gardner M. & Thomas P. (2013). Retrophyllum rospigliosii. The IUCN Red List of Threatened Species 2013: e.T34110A2846471. Online. Available: (downloaded 29 October 2015).
Garnier J. (1867). Essai sur la géologie et les ressources minérales de la Nouvelle-Calédonie. Ann. Mines Sér. 6, Mém. 12: 192.
Gaussen H. (1976). Les Gymnospermes actuelles et fossiles. Fascicule XIV. Genre Podocarpus. Conclusion des Podocarpines. Trav. Lab. Forest. Toulouse, tome 2, vol. 1, fasc. XIV. Toulouse: Faculté des Sciences.
GeoNames (no date). Online. Available: (accessed 23 July 2014 etc.)
Graham D. (2008). Special to (August 17, 2008). New tree species discovered in Amazon biodiversity hotspot. Online. Available: (accessed 6 December 2011).
Graham D. (2012). Lonely George in love! Proj. Amazonas Newslett. Dec. 2012: 1–2. Online. Available: (downloaded 27 July 2015).
Gray B. (1974). Forest insect problems in the South Pacific islands. Commonw. Forest. Rev. 53 (1): 3948.
Gray N. E. (1960). A taxonomic revision of Podocarpus XII. Section Microcarpus. J. Arnold Arbor. 41: 3639.
Gray N. E. (1962). A taxonomic revision of Podocarpus XIII. Section Polypodiopsis in the South Pacific. J. Arnold Arbor. 43: 6779.
Gray N. E. & Buchholz J. T. (1948). A taxonomic revision of Podocarpus III. The American species of Podocarpus: section Polypodiopsis. J. Arnold Arbor. 29: 117122.
Greenwood D. R. (1987). Early Tertiary Podocarpaceae: megafossils from the Eocene Anglesea locality, Victoria, Australia. Austral. J. Bot. 35: 111133.
Greenwood W. (1977). The food plants or hosts of some Fijian insects. V. Proc. Linn. Soc. New South Wales 101: 237241.
Guerrero Forero E. & Hodson de Jaramillo E. (1988). Micorrizas en Decussocarpus rospigliossi (Pilger) De Laubenfels, una Podocarpacea del bosque andino. Revista Fac. Ci. Univ. Javeriana 1 (2): 5366.
Guillaumin A. (1911). Catalogue des plantes phanérogames de la Nouvelle-Calédonie et dépendances (Iles des Pins et Loyalty). Ann. Mus. Colon. Marseille Sér. 2 9: 77290.
Guillaumin A. (1944). Contribution à la flore de la Nouvelle-Calédonie. LXXXV. Plantes de collecteurs divers (fin). Bull. Mus. Natl. Hist. Nat. Sér. 2 16: 346351.
Hair J. B. & Beuzenberg E. J. (1958). Chromosomal evolution in the Podocarpaceae. Nature 181 (4623): 15841586.
Herbario Nacional Colombiano (no date). Colleciones cientificas en linea. Online. Available: (accessed 5 February 2015 etc.).
Herbert J., Hollingsworth P. M., Gardner M. F., Mill R. R., Thomas P. I. & Jaffré T. (2002). Conservation genetics and phylogenetics of New Caledonian Retrophyllum (Podocarpaceae) species. New Zealand J. Bot. 40 (2): 175188.
Herbier du Centre IRD de Nouméa (NOU) (no date). Database VIROT. Online. Available: (accessed 21 and 22 July 2014).
Hill R. S. & Brodribb T. J. (1999). Southern conifers in time and space. Austral. J. Bot. 47 (5): 639696.
Hill R. S. & Merrifield H. (1993). An Early Tertiary macroflora from West Dale, south-western Australia. Alcheringa 17: 285326.
Hill R. S. & Pole M. S. (1992). Leaf and shoot morphology of extant Afrocarpus, Nageia and Retrophyllum (Podocarpaceae) species, and species with similar leaf arrangement, from Tertiary sediments in Australasia. Austral. Syst. Bot. 5: 337358.
Hokche O., Berry P. E. & Huber O. (2008). Nuevo Catálogo de la Flora Vascular de Venezuela. Caracas: Fundación Instituto Botánico de Venezuela.
Hope G. (2015). Extended vegetation histories from ultramafic karst depressions. Austral. J. Bot. 63 (4): 222233.
Hopkins H. C. F. & Bradford J. C. (2009). Nomenclature and typification of names in the endemic genus Pancheria (Cunoniaceae) from New Caledonia. Adansonia 31 (1): 103135.
Huguenin B. (1969). Micromycètes du Pacifique sud (Huitième contribution). Ascomycètes de Nouvelle-Calédonie (II). Cah. Pacifique no. 13: 295308.
Hutchinson J. (as ‘J. H.’) (1920). Polypodiopsis, Carrière. Bull. Misc. Inform. Kew 1920: 372.
Instituto de Ciencias Naturales, Universidad Nacional de Colombia (2004 and continually updated). Colecciones en Línea. Online. Available: (accessed 8 February 2013).
IUCN (2012). IUCN Red List Categories and Criteria, version 3.1, 2nd edition. Gland, Switzerland and Cambridge: International Union for Conservation of Nature.
IUCN Standards and Petitions Subcommittee (2013). Guidelines for using the IUCN Red List Categories and Criteria, version 10.1 (September 2013). Online. Available: (downloaded 19 November 2013).
Jaffré T. (1988). Végétation et flore de la Chute de la Madeleine. (Étude en vue d'une proposition de mise en reserve.) Nouméa: Laboratoire de botanique, ORSTOM.
Jaffré T., Munzinger J. & Lowry P. P. II (2010). Threats to the conifer species found on New Caledonia's ultramafic massifs and proposals for urgently needed measures to improve their protection. Biodivers. & Conserv. 19 (5): 14851502.
JBRJ – Instituto de Pesquisas Jardim Botânico do Rio de Janeiro (no date). Jabot – Banco de Dados da Flora Brasileira. Online. Available: (accessed 15 April 2014).
Jørgensen P. M. & Léon-Yánez S. (eds.) (1999). Catalogue of the vascular plants of Ecuador. / Catálogo de las plantas vasculares del Ecuador. Being: Monogr. Syst. Bot. Missouri Bot. Gard. 75. St. Louis, Missouri: Missouri Botanical Garden Press.
JSTOR Global Plants (no date). Electronic database. Online. Available: (accessed 3 September 2014 etc.).
Keppel G. (2005). Summary Report on Forests of the Mataqali Nadicake Kilaka, Kubulau District, Bua, Vanua Levu. Suva: Wildlife Conservation Program (downloaded from on 23 July 2014).
Keppel G., Tuiwawa M. V., Naikatini Alifereti & Rounds I. A. (2011). Microhabitat specialization of tropical rain-forest canopy trees in the Sovi Basin, Viti Levu, Fiji Islands. J. Trop. Ecol. 27 (5): 491501.
Kew (no date). The Herbarium Catalogue, Royal Botanic Gardens, Kew. Online. Available: (accessed 3 September 2014 etc.).
Knopf P., Schulz C., Little D. P., Stützel T. & Stevenson D. W. (2012). Relationships within Podocarpaceae based on DNA sequence, anatomical, morphological, and biogeographical data. Cladistics 28 (3): 271299.
Koch K. H. E. (1871). Bericht über die neuesten Pflanzen. (Schluss). Wochenschr. Vereines Beförd. Gartenbaues Königl. Preuss. Staaten 14 (25): 195200.
Krypto-S (no date). Botanical Database at the Swedish Museum of Natural History. Online. Available: (accessed July 2014 etc.)
Lapie G. (1928). Les forêts des colonies françaises. Ann. École Natl. Eaux 2 (1): 107136.
Li J. (2007). Gymnosperms of Papua. In: Marshall A. J. (ed.) Ecology of Indonesian Papua, 1, pp. 344348. Singapore: Periplus Editions/Conservation International Foundation.
Lindman C. A. M. (1898). Leguminosae austro-americanae ex itinere Regnelliano primo. Bih. Kongl. Svenska Vetensk.-Akad. Handl. 24 (Afdelning III), no. 7, 61 pp.
Mansano V. de F. & Vianna Filho M. D. M. (2010). A taxonomic reappraisal of the South American genus Holocalyx (Leguminosae, Papilionoideae). Brittonia 62 (2): 110115.
McKee M. E. (1972). Exploration de l'intérieur de la Nouvelle-Calédonie. J. Soc. Océanistes 28 (36): 199229.
McNeill J., Barrie F. R., Burdet H. M., Demoulin V., Hawksworth D. L., Marhold K., Nicolson D. H., Prado J., Silva P. C., Skog J. E., Wiersema J. H. & Turland N. J. (2006). International Code of Botanical Nomenclature (Vienna Code) adopted by the Seventeenth International Botanical Congress, Vienna, Austria, July 2005. Regnum Veg. 146, 568 pp. Ruggell: A.R.G. Gantner Verlag KG.
McNeill J., Barrie F. R., Buck W. R., Demoulin V., Greuter W., Hawksworth D. L., Herendeen P. S., Knapp S., Marhold K., Prado J., Prud'homme van Reine W. F., Smith G. F., Wiersema J. H. & Turland N. J. (2012). International Code of Nomenclature for Algae, Fungi, and Plants (Melbourne Code) adopted by the Eighteenth International Botanical Congress, Melbourne, Australia, July 2011. Regnum Veg., 154. Ruggell: A.R.G. Gantner Verlag KG.
Mead J. P. (1928). The forests of the Fiji Islands. Empire Forest. J. 7: 4754.
Melikian A. P. & Bobrov A. V. (2000). Morfologiya shchenskikh reproduktivi'kh obganov i op't rostroenniya filogeneticheskoj sistemy poryadkov Podocarpales, Cephalotaxales i Taxales [Morphology of female reproductive structures and an attempt of the construction of phylogenetic system of orders Podocarpales, Cephalotaxales, and Taxales]. Bot. Zhurn. (Moscow & Leningrad) 85 (7): 5068.
Merkhofer L., Wilf P., Haas M. T., Kooyman R. M., Sack L., Scoffoni C. & Cúneo N. R. (2015). Resolving Australian analogs for an Eocene Patagonian paleorainforest using leaf size and floristics. Amer. J. Bot. 102 (7): 11601173.
Mill R. R. (2010). (1938) Proposal to reject the name Torreya bogotensis (Gymnospermae, ? Podocarpaceae). Taxon 59 (3): 975976.
Mill R. R. & Hill R. S. (2004). Validations of the names of seven Podocarpaceae macrofossils. Taxon 53 (4): 10431046.
Mill R. R. & Weston P. (2004). Proposals to reject the names Polypodiopsis and Polypodiopsis muelleri (Plantae vasculares, incertae sedis). Taxon 53 (1): 203205.
Morat P. (2010). Les botanistes récolteurs en Nouvelle-Calédonie de 1774 à 2005. Adansonia Sér. 3 32: 159216.
Müller E. & Dennis R. W. G. (1965). Fungi venezuelani: VIII: Plectascales, Sphaeriales, Loculoascomycetes. Kew Bull. 19 (3): 357386.
Muséum national d'histoire naturelle (no date). Paris herbarium database. Online. Available: (accessed 3 February 2015).
Nieto V. M. & Rodriguez J. (2003). Decussocarpus rospigliosii. In: Vozzo J. A. (ed.) Tropical Tree Seed Manual, p. 432. Washington, District of Columbia: United States Department of Agriculture Forest Service.
Niño J., Espinal C., Mosquera O. M. & Correa Y. M. (2007). Antimycotic activity of 20 plants from Colombian flora. Pharm. Biol. 41 (7): 491496.
Osborne L. D. (1967). Comparative decay resistance of twenty-five Fijian timber species in accelerated laboratory tests. Pacific Sci. 21: 539549.
Page C. N. (1989 [‘1988’]). New and maintained genera in the conifer families Podocarpaceae and Pinaceae. Notes Roy. Bot. Gard. Edinb. 45 (2): 377395.
Page C. N. (1990). Podocarpaceae. In: Kubitzki K. (ed.) The Families and Genera of Vascular Plants, Volume 1, Pteridophytes and Gymnosperms (ed. Kramer K. U. & Green P. S.), pp. 332346. Heidelberg: Springer-Verlag.
Parham J. W. (1964). Plants of the Fiji Islands. Gymnospermae. Suva: Government Press.
Parham J. W. (1972). Plants of the Fiji Islands, revised edition. Suva: Government Printer.
Parlatore P. (1868). Podocarpus. In: De Candolle A., Prodromus Systematis Vegetabilium, 16 (2): 507521.
Pilger R. (1903). Taxaceae. In: Engler A. (ed.) Das Pflanzenreich IV.5 (Heft 18). Leipzig: Wilhelm Engelmann.
Pilger R. (1923). Eine neue Podocarpus-Art aus Peru. Notizbl. Bot. Gart. Berlin-Dahlem 8 (74): 273274.
Pole M. (1992). Early Miocene flora of the Manuherikia Group, New Zealand. 2. Conifers. J. Roy. Soc. New Zealand 22 (4): 287302.
Pole M. (1997). Miocene conifers from the Manuherikia Group, New Zealand. J. Roy. Soc. New Zealand 27 (3): 355370.
Pole M. (2007). Conifer and cycad distribution in the Miocene of southern New Zealand. Austral. J. Bot. 55 (2): 143164.
Province Sud (2012). Les Chutes de la Madeleine. Online. Available: (downloaded 20 August 2014).
Quijano-Celis C. E., Gaviria M., Vanegas-López C., Ontiveros I., Echeverri L., Morales G. & Pino J. A. (2010). Chemical composition and antibacterial activity of the essential oil of Retrophyllum rospigliosii fruits from Colombia. Nat. Prod. Commun. 5 (7): 11331134.
Quijano-Celis C. E., Gaviria M., Vanegas-López C., Ontiveros I., Echeverri L., Pino J. A. & Morales G. (2013). Chemical composition and antibacterial activity of the essential oil of Retrophyllum rospigliosii (Pilger) C. N. Page leaves from Colombia. J. Essent. Oil Bear. Pl. 16 (5): 595599.
Rahimi R., Ghiasi S., Azimi H., Fakhari S. & Abdollahi M. (2010). A review of the herbal phosphodiesterase inhibitors: future perspective of new drugs. Cytokine 49 (2): 123129.
Rainforest Information Centre (no date). The NSW Good Wood Guide: Imported Timbers Whose Use Should be Avoided. Online. Available: (accessed 16 September 2015).
Rasband W. S. (2014). ImageJ, version 1.49d. Bethesda, Maryland: US National Institutes of Health. Online. Available: (accessed 10 July 2014 onwards).
Reveal J. L. (2012). A divulgation of ignored or forgotten binomials. Phytoneuron 2012-28: 1–64.
Ross J. (2007). Colour Analysis Tools in ImageJ. Auckland: Biomedical Imaging Research Unit, University of Auckland. Online. Available: (downloaded 12 February 2015).
Salter J. (2004). Comparative morphological, anatomical and embryological studies of Prumnopitys taxifolia and P. ferruginea (Podocarpaceae), and the hydrodynamics of their saccate pollen grains. Ph.D. thesis, University of Auckland.
Schedl K. E. (1965). Ein neuer Platypus aus Venezuela. Anz. Schädlingskunde 38: 87.
Schnee L. (1944). El género Podocarpus en Venezuela. Bol. Soc. Venez. Ci. Nat. 9 (59): 181188, pl. 1–6.
Sebert H. (1874). Notice sur les bois de la Nouvelle Calédonie suivie de considérations générales sur les propriétés mécaniques des bois et sur les procédés employés pour les mesurer. Paris: Arthus Bertrand.
Secco R. de S., Silva do Rosário A. & Rodrigues W. A. (2009). Podocarpaceae. In: Giulietti A. M., Rapini A., Gomes de Andrade M. J., de Queiroz L. P. & Cardano da Silva J. M. (orgs) Plantas Raras do Brasil, p. 341. Belo Horizonte: Conservação Internacional/Universidade Estadual de Feira de Santana.
Seemann B. (1862). Viti. An Account of a Government Mission to the Vitian or Fijian Islands in the Years 1860–1861. Cambridge: Macmillan.
Seemann B. (1863). Podocarpus vitiensis, a new coniferous tree, from the Viti Islands. J. Bot. 1: 3336, t. 2.
Seemann B. (1868). Flora Vitiensis: a Description of the Plants of the Viti or Fiji Islands with an Account of their History, Uses, and Properties, part 9. London: L. Reeve & Co.
Silba J. (1983). A new species of Decussocarpus de Laub. (Podocarpaceae) from Brazil. Phytologia 54 (6): 460462.
Silba J. (1986). Encyclopaedia Coniferae. Phytologia Mem. 8. Corvallis, Oregon: Harold N. Moldenke & Alma L. Moldenke.
Sinclair W. T., Mill R. R., Gardner M. F., Woltz P., Jaffré T., Preston J., Hollingsworth M. L., Ponge A. & Möller M. (2002). Evolutionary relationships of the New Caledonian heterotrophic conifer, Parasitaxus usta (Podocarpaceae), inferred from chloroplast trnL-F intron/spacer and nuclear rDNA ITS2 sequences. Plant Syst. Evol. 233: 79104.
Smith A. C. (1979). Flora vitiensis nova. A new flora of Fiji (Spermatophytes only). Vol. 1. Introduction; Gymnospermae, Families 1–6; Angiospermae, Monocotyledones, Families 7–43 (except Family 32, Orchidaceae). Lawai: Pacific Tropical Botanical Garden (Podocarpaceae, pp. 92–108).
Stafleu F. A. & Cowan R. S. (1976). Entry for Emmanuel Drake del Castillo (1855–1904). In: Taxonomic Literature, ed. 2, p. 678. Utrecht: Bohn, Scheltema & Holkema.
Stevenson J. & Hope G. (2005). A comparison of late Quaternary forest changes in New Caledonia and northeastern Australia. Quatern. Res. 64: 372383.
Suprin B. (2011a). Podocarpacées. Le bois bouchon. In: Florilège des Plantes en Nouvelle-Calédonie, 1, pp. 169172. Nouméa: Éditions Photosynthèse.
Suprin B. (2011b). Podocarpacées. Le “palissandre”. In: Florilège des Plantes en Nouvelle-Calédonie, 1: 173. Nouméa: Éditions Photosynthèse.
Takhtajan A. (1986). Floristic Regions of the World. Berkeley, California: University of California Press.
Takeuchi W. (2010). A floristic reconnaissance of montane environments in the Foja Mts of Papua Province, Indonesia. Harvard Pap. Bot. 15 (1): 1125.
Thiers B. (continuously updated). Index Herbariorum: A Global Directory of Public Herbaria and Associated Staff. New York Botanical Garden's Virtual Herbarium. Online. Available: (accessed 23 July 2014 etc.).
Thomas P. (2010a). Retrophyllum comptonii. The IUCN Red List of Threatened Species 2010: e.T42542A10719849. Online. Available: (downloaded 29 October 2015).
Thomas P. (2010b). Retrophyllum minus. The IUCN Red List of Threatened Species 2010: e.T34108A9837487. Online. Available: (downloaded 29 October 2015).
Thomas P. (2013). Retrophyllum vitiense. The IUCN Red List of Threatened Species 2013: e.T42543A2986320. Online. Available: (downloaded 29 October 2015).
Thurber G. (1870). 26. Torreya Bogotensis. Bull. Torrey Bot. Club 1 (2): 7.
Tomlinson P. B. & Zacharias E. H. (2001). Phyllotaxis, phenology and architecture in Cephalotaxus, Torreya and Amentotaxus (Coniferales). Bot. J. Linn. Soc. 135 (3): 215228.
Torres-Romero J. H. (1988). Podocarpaceae. In: Pinto P. & Lozano G. (eds) Flora Colombia Monogr. 5, pp. 567. Bogotá: Instituto de Ciencias Naturales, Museo de Historia Natural, Faculdad de Ciencias, Universidad Nacional de Colombia.
van Steenis C. G. G. J. (1981). Rheophytes of the World. Alphen aan den Rijn, the Netherlands, and Rockville, Maryland: Sijthoff & Noordhoff.
Vargas Rincón C. A. (2011). Caracterización florística y fitogeográfica del sector sur de la Serranía de Perijá y áreas adyacentes de la Cordillera Oriental Colombiana. Master's thesis, Universidad Nacional de Colombia.
Vásquez Correa Á. M. (2010). Anatomía de la madera de las podocarpáceas colombianas. Ph.D. thesis, Universidad de Córdoba.
Vásquez Correa Á. M. & Alcantara Vara E. (2009). Anatomía y densidad de la madera de árboles de pino romerón [Retrophyllum rospigliosii (Pilger) C.N. Page] de ocho años de edad. Revista Fac. Nac. Agron. Medellín Univ. Antioquia 62 (1): 48694880.
Vásquez Correa Á. M., Alcántara Vara E. & Herrera Machuca M. Á. (2010). Wood anatomy of Colombian Podocarpaceae (Podocarpus, Prumnopitys and Retrophyllum). Bot. J. Linn. Soc. 164 (3): 293302.
Vicuña-Miñano E. E. (2005). Las Podocarpáceas de los bosques montanos del noroccidente peruano. Podocarpaceae of the montane forests from northwestern Peru. Revista Peruana Biol. 12: 283288.
Vicuña-Miñano E. & León J. M. (2003). Notas sobre Podocarpáceas de cuatro bosques montanos de la Provincia de San Ignacio – Cajamarca, Perú. Arnaldoa 10: 1943.
Vieillard E. & Deplanche É. (1863). Essai sur la Nouvelle-Calédonie. Paris: Challamel aîné.
Vieira L. do N., Rogalski M., Faoro H., Fraga H. P. de F., dos Anjos K. G., Picchi G. F. A., Nodari R. O., Pedrosa F. de O., de Souza E. M., & Guerra M. P. (2016). The plastome sequence of the endemic Amazonian conifer, Retrophyllum piresii (Silba) C.N. Page, reveals different recombination events and plastome isoforms. Tree Genet. Genomes 12: 10 (11 pp). Online. Available: (accessed 27 January 2016).
Wasscher J. (1941). The genus Podocarpus in the Netherlands Indies. Blumea 4 (3): 359481.
Wikramanayake E., Dinerstein E., Loucks C. J., Olson D. M., Morriaon J., Lamoreux J., McKnight M. & Hedao P. (2002). Terrestrial Regions of the Indo Pacific. A Conservation Assessment. Washington, District of Columbia: Island Press.
Wildfinder (no date). Online. Available: (accessed 5 August 2014 etc.).
Wilf P. (2012). Rainforest conifers of Eocene Patagonia: attached cones and foliage of the extant Southeast Asian and Australasian genus Dacrycarpus (Podocarpaceae). Amer. J. Bot. 99 (3): 562584.
Wilf P., Escapa I. H. & Cúneo N. R. (2012). Eocene rainforest conifers of the Patagonian fire lakes. Botany 2012, July 7–11 2012, Columbus, Ohio, abstract no. 175.
Wilf P., Escapa I. H., Cúneo N. R., Kooyman R. M., Johnson K. R. & Iglesias A. (2014). First South American Agathis (Araucariaceae), Eocene of Patagonia. Amer. J. Bot. 101 (1): 156179.
Williams R. L. (2004). An intellectual biography of Elie-Abel Carriere (1818–1896). Brittonia 56 (4): 365374.
Wood S. L. (1971). New records and species of American Platypodidae (Coleoptera). Great Basin Naturalist 31 (4): 243253.
Wood S. L. & Bright D.E. Jr. (1992). A catalog of Scolytidae and Platypodidae (Coleoptera), Part 2: Taxonomic Index. Great Basin Naturalist Mem, no. 13. [‘Hosts of Scolytidae and Platypodidae’, pp. 1241–1348.]
Yaguana C., Lozano D., Neill D. A. & Asanza M. (2012). Diversidad florística y estructura del bosque nublado del Río Numbala, Zamora-Chinchipe, Ecuador: El “bosque gigante” de Podocarpaceae adyacente al Parque Nacional Podocarpus. Revista Amazón.: Ci. Tecnol. 1 (3): 226247.
Zenteno-Ruiz F. S. (2007). Retrophyllum rospigliosii (Podocarpaceae), un nuevo registro de pino de monte, en el noroeste de Bolivia. Kempffiana 3 (2): 35. Online. Available: (downloaded 19 August 2009).
Zonneveld B. J. M. (2012). Conifer genome sizes of 172 species, covering 64 of 67 genera, range from 8 to 72 picogram. Nordic J. Bot. 30 (4): 490502.
Zürich Herbaria (no date). Database. Online. Available: (accessed July 2014).
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