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Earliest Wuchiapingian (Lopingian, late Permian) brachiopods in southern Hunan, South China: implications for the pre-Lopingian crisis and onset of Lopingian recovery/radiation

Published online by Cambridge University Press:  14 July 2015

Shu-Zhong Shen  and
Yi-Chun Zhang
Shu-Zhong Shen
Affiliation:
State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, P. R. China,
Yi-Chun Zhang
Affiliation:
State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, P. R. China,
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Abstract

The uppermost 5–15 m of the Douling Formation in the southern Hunan area, South China, yields a diverse fauna comprised of ammonoids, bivalves, and brachiopods. The brachiopods reported in this paper consist of 51 species in 34 genera and are dominated by the Lopingian (Late Permian) species associated with a few species persisting from the underlying Maokouan (Late Guadalupian). This fauna is of earliest Wuchiapingian in age as precisely constrained by the associated conodont Clarkina postbitteri postbitteri and the Guadalupian-type ammonoid fauna of the Roadoceras-Doulingoceras Zone in the brachiopod horizon. The discovery of the Lopingian species-dominated brachiopod fauna in the earliest Wuchiapingian in southern Hunan suggests a much less pronounced effect of the pre-Lopingian crisis (end-Guadalupian mass extinction) than the end-Changhsingian mass extinction in terms of brachiopods, a contemporaneous onset of the Lopingian recovery/radiation during the pre-Lopingian crisis period, and taxonomic selectivity of the pre-Lopingian crisis in terms of different fossil groups. New taxa are Echinauris doulingensis n. sp., Pararigbyella quadrilobata n. gen. and n. sp. and P. doulingensis n. gen. and n. sp.

Type
Research Article
Information
Journal of Paleontology , Volume 82 , Issue 5 , September 2008 , pp. 924 - 937
Copyright
Copyright © The Paleontological Society 

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References

Abich, H. 1878. Geologische Forschungen in den kaukasischen Landern I: Eine Bergkalkfauna aus der Araxesenge bei Djoulfa in Armenia. Wien, 128p.Google Scholar
Archbold, N. W. 1981. Quinquenella magnifica sp. nov. (Chonetida, Brachiopoda) from the Permian of Irian Jaya, Indonesia; a study of the ontogeny of a chonetid brachiopod. Publication of the Geological Research Development Center, Paleontology Series, 2:27–34.Google Scholar
Beyrich, H. E. 1865. Ueber eine Kohlenkalk-Fauna von Timor. Kaiserliche Akademie der Wissenschaften zu Berlin, Abhandlungen, 1864:61–98.Google Scholar
Broili, F. 1916. Die Permischen Brachiopoden von Timor, p. 1–104. In Wanner, J. (ed.), Paläontologie von Timor, Volume 7. Stuttgart.Google Scholar
Bronn, H. G. 1862. Die Klassen und Ordnungen der Weichthiere. Leipzig and Heidelberg, Malacozoa, 518 p.Google Scholar
Bruguière, J. G. 1797. Vers, Coquilles, Mollusques et Polypiers. Tableau encyclopédique et méthodique des trois règnes de la nature. Agasse, Paris, 2,96–314.Google Scholar
Chao, K. K. 1965. The Permian ammonoid-bearing formations of South China. Scientia Sinica, 14(2):1813–1825.Google Scholar
Chao, K. K. 1966. Permian ammonoid-bearing formations of South China. Journal of Stratigraphy, 1(2):170–187.Google Scholar
Chao, Y. T. 1927. Productidae of China, Pt. 1, Producti. Palaeontologia Sinica, Series B, 5(2):1–206.Google Scholar
Chao, Y. T. 1928. Productidae of China, Pt. 2, Chonetinae, Productinae, and Richthofeninae. Palaeontologia Sinica, Series B, 5(3):1–81.Google Scholar
Chao, Y. T. 1929. Carboniferous and Permian spiriferids of China. Palaeontologica Sinica, Ser. B, 11(1):1–101.Google Scholar
Chen, C. Z., Zhang, Z. M., and Xu, J. T. 1974. Bivalves (Permian), p. 302–303. In Nanjing Institute of Geology and Palaeontology (ed.), Handbook of the Stratigraphy and Palaeontology in Southwest China. Science Press, Beijing.Google Scholar
Chen, Z. Q., Campi, M. J., Shi, G. R., and Kaiho, K. 2005. Post-extinction brachiopod faunas from the Late Permian Wuchiapingian coal series of South China. Acta Palaeontologica Polonica, 50:343–363.Google Scholar
Erwin, D. H. 1996. Understanding biotic recoveries: extinction, survival and preservation during the end-Permian mass extinction, p. 398–418. In Jablonski, D., Erwin, D. H., and Lipps, J. H. (eds.), Evolutionary Paleobiology. The University of Chicago Press, Chicago and London.Google Scholar
Fang, Z. J. 1987. Bivalves from upper part of Permian in southern Hunan, China, p. 349–411. In Nanjing Institute of Geology and Palaeontology (ed.), Nanjing Institute of Geology and Palaeontology, Academia Sinica, Collection of postgraduate theses. Jiangsu Science and Technology Publishing House, Nanjing.Google Scholar
Feng, R. L. and Jiang, Z. L. 1978. Brachiopoda, p. 231–305. In Working Group of Stratigraphy and Palaeontology of Guizhou Province (ed.), Paleontological atlas of southwest China. Guizhou. Volume 2. Geological Publishing House, Beijing.Google Scholar
Frech, F. 1911. Die Dyas. In Richthofen, F. Von (ed.), China 5. Dietrich Reimer, Berlin, 289 p.Google Scholar
Fredericks, G. N. 1916. Paleontologicheskiia zametki. 2. O nekotorykh' verkhnepaleozoiskikh' brakhiopodakh' Evrazii (Paleontological notes. 2. On some Upper Paleozoic Brachiopoda of Eurasia). Trudy Geologicheskogo Komiteta, 156:1–87.Google Scholar
Gemmellaro, G. G. 1887. La fauna dei calcari con Fusulina della valle del Fiume Sosio nella provincia di Palermo. Giornale di Scienze Naturali ed Economiche, Palermo, 19:1–106.Google Scholar
Gemmellaro, G. G. 1899. La fauna dei calcari con Fusulina della valle del fiume Sosio nella provincia di Palermo. Samem, 22:95–214.Google Scholar
Grabau, A. W. 1924. Stratigraphy of China, Pt. 1, Palaeozoic and Older. Geological Survey of China, Beijing, 528 p.Google Scholar
Grabau, A. W. 1931. The Brachiopoda, Pt. 2, Studies for students, I, Palaeontology. Science Quarterly of the National University of Peking, 2(3):397–422.Google Scholar
Grant, R. E. 1968. Structural adaptation in two Permian brachiopod genera, Salt Range, West Pakistan. Journal of Paleontology, 42:1–32.Google Scholar
Grant, R. E. 1976. Permian brachiopods from southern Thailand. Journal of Paleontology, Memoir 9, 50(Supplement no. 3):1–269.Google Scholar
Gray, J. E. 1840. Synopsis of the contents of the British Museum, 42nd edition, London, 370 p.Google Scholar
Grunt, T. A. and Dmitriev, V. Y. 1973. Permskie brakhiopody Pamira (Permian Brachiopoda of the Pamir). Akademiia Nauk SSSR, Paleontologicheskii Institut, Trudy, 136:1–212.Google Scholar
Hall, J. 1852. Notes upon some of the fossils collected on the route from the Missouri River to the Great Salt Lake, and in the vicinity of the Latter Place, by the expedition under the command of caption Howard Stansbury, exploration and survey of the valley of the Great Salt Lake of Utah. Appendix E:401–414.Google Scholar
Hamlet, B. 1928. Permische Brachiopoden, Lamellibranchiaten und Gastropoden von Timor. Jaarboek van het Mijnwezen in Nederlandsch Oost-Indie. Gravenhage, 56(2):1–115.Google Scholar
He, X. L. and Zhu, M. L. 1979. A new form of brachiopods and its systematical classification. Journal of China University of Mining and Technology, (4):131–140.Google Scholar
Hou, H. F., Zhan, L. P., and Chen, B. W. 1979. The coal-bearing strata and fossils of the Late Permian from Guangdong. Geological Publishing House, Beijing, 166 p.Google Scholar
Hu, S. Z. 1983. Brachiopods from the Hsiaochiangpien Limestone, southern Jiangsi. Acta Palaeontogica Sinica, 22:338–345.Google Scholar
Hu, S. Z. 1994. On the event of Dongwu Movement and its relation with Permian subdivision. Journal of Stratigraphy, 18:309–315.Google Scholar
Huang, T. K. 1932. Late Permian brachiopods of southwest China. Palaeontologia Sinica, Series B, 9(1):1–138.Google Scholar
Huang, T. K. 1933. Late Permian Brachiopoda of southwestern China, Pt. II, Palaeontologia Sinica, Series B, 9(2):1–172.Google Scholar
Isozaki, Y., Shimizu, N., Yao, J. X., Ji, Z. S., and Matsuda, T. 2007a. End-Permian extinction and volcanism-induced environmental stress: The Permian-Triassic boundary interval of lower-slope facies at Chaotian, South China. Palaeogeography, Palaeoclimatology, Palaeoecology, 252:218–238.Google Scholar
Isozaki, Y., Kawahata, H., and Ota, A. 2007b. A unique carbon isotope record across the Guadalupian–Lopingian (Middle–Upper Permian) boundary in mid-oceanic paleo-atoll carbonates: The high-productivity “Kamura event” and its collapse in Panthalassa. Global and Planetary Change, 55:21–38.Google Scholar
Jin, Y. G. 1993. Pre-Lopingian benthos crisis, p. 269–278, C. R. XII ICC-P, Volume 2, Buenos Aires.Google Scholar
Jin, Y. G. and Hu, S. Z. 1978. Brachiopods of the Kuhfeng Formation in South Anhui and Nanking Hills. Acta Palaeontologica Sinica 17:101–127.Google Scholar
Jin, Y. G., Liao, Z. T., and Fang, B. X. 1974. Brachiopoda (Permian). p. 308–313, In Nanjing Institute of Geology and Palaeontology (ed.), Handbook of the Stratigraphy and Palaeontology in Southwest China. Science Press, Beijing.Google Scholar
Jin, Y. G., Mei, S. L., Wang, W., Wang, X. D., Shen, S. Z., Shang, Q. H., and Chen, Z. Q. 1998. On the Lopingian Series of the Permian System, p. 1–18. In Jin, Y. G., Wardlaw, B. R., and Wang, Y. (eds.), Permian stratigraphy, environments and resources, Palaeoworld 9. Volume 2. China University of Science and Technology Press, Hefei.Google Scholar
Jin, Y. G., Shen, S. Z., Henderson, C. M., Wang, X. D., Wang, W., Wang, Y., Cao, C. Q., and Shang, Q. H. 2006. The Global Stratotype Section and Point (GSSP) for the boundary between the Capitanian and Wuchiapingian stage (Permian). Episodes, 29(4):253–262.Google Scholar
Jin, Y. G., Wang, Y., Wang, W., Shang, Q. H., Cao, C. Q., and Erwin, D. H. 2000. Pattern of marine mass extinction near the Permian-Triassic boundary in South China. Science, 289:432–436.Google Scholar
Jin, Y. G., Zhang, J., and Shang, Q. H. 1994. Two phases of the end-Permian mass extinction, p. 813–822. In Embry, A. F., Beauchamp, B., and Glass, D. J. (eds.), Pangea: Global Environments and Resources. Canadian Society of Petroleum Geologists, Memoir 17.Google Scholar
Kayser, E. 1883. Obercarbonische fauna von Loping, p. 1–269. In Richthofen, von (ed.), China, Volume 4, D. Reimer, Berlin.Google Scholar
Kozur, H. 1975. Beitrage zur conodontenfauna des Perm. Geologisch Palaeontologische Mitteilungen Innsbruck, 5(4):1–44.Google Scholar
Lazarev, S. S. 1986. Osnovnye napravleniia evoliutsii i sistema brakhiopod podotriada Productidina, (Author's abstract of doctoral dissertation in biology), Akademiia Nauk SSSR, Paleontologicheskii Institut, Moscow, 41 p.Google Scholar
Liao, Z. T. 1979. Brachiopod assemblage zone of Changhsing Stage and brachiopods from Permo-Triassic boundary beds in China. Journal of Stratigraphy, 3(3):200–213.Google Scholar
Liao, Z. T. 1980. Late Permian brachiopods from western Guizhou, p. 241–277. In Nanjing Institute of Geology and Palaeontology (ed.), Late Permian coal-bearing strata and biota from western Guizhou and eastern Yunnan provinces. Science Press, Beijing.Google Scholar
Liao, Z. T. 1987. Palaeoecological characters and stratigraphic significance of silicified brachiopods of the Upper Permian from Heshan, Laibin, Guangxi, p. 81–126. In Nanjing Institute of Geology and Palaeontology (ed.), Stratigraphy and paleontology of systematic boundaries in China: Permian-Triassic boundary 1. Nanjing University Press, Nanjing.Google Scholar
Licharew, B. K. 1937. Permskie Brachiopoda Severnogo Kavkaza. Semeistva Chonetidae Hall et Clarke i Productidae Gray (Brachiopoda of the Permian System of U.S.S.R. Fasc. I. Permian Brachiopoda of North Caucasus: Families: Chonetidae Hall and Clarke and Productidae Gray). Monografii po Paleontologii SSSR, 39(1):1–152.Google Scholar
McCoy, F. 1851. Aviculopecten. Annals Magazine of Natural History, 2:171.Google Scholar
Mei, S. L. and Wardlaw, B. R. 1996. On the Permian “liangshanensisbitteri” zone and the related problems, p. 130–140. In Wang, H. Z. and Wang, X. L. (eds.), Centennial Memorial Volume of Professor Sun Yunzhu: Stratigraphy and Palaeontology. China University of Geosciences Press, Wuhan.Google Scholar
Mei, S. L., Henderson, C. M., and Wardlaw, B. R. 2002. Evolution and distribution of the conodont Sweetognathus and Iranognathus and related genera during the Permian, and their implications for climate changes. Palaeogeography Palaeoclimatology Palaeoecology, 180:57–91.Google Scholar
Mei, S. L., Jin, Y. G., and Wardlaw, B. R. 1994. Zonation of conodonts from the Maokouan-Wuchiapingian boundary strata, South China, p. 225–233. In Jin, Y. G., Utting, J., and Wardlaw, B. R. (eds.), Permian stratigraphy, environments and resources, Palaeoworld 4. Volume 1. Nanjing University Press, Nanjing.Google Scholar
Mei, S. L., Jin, Y. G., and Wardlaw, B. R. 1998. Conodont succession of the Guadalupian-Lopingian boundary strata in Laibin of Guangxi, China and West Texas, USA, p. 53–76. In Jin, Y. G., Wardlaw, B. R., and Wang, Y. (eds.), Permian stratigraphy, environments and resources, Palaeoworld 9. Volume 2. Nanjing University Press, Nanjing.Google Scholar
Meng, F. Y. 1980. Subdivision of the Permian System in southern Hunan and a discussion on the age of the Douling Coal-Series. Geological Review, 26:233–238.Google Scholar
Muir-Wood, H. M. 1955. A history of the classification of the Phylum Brachiopoda. British Museum Natural History, London, 124 p.Google Scholar
Muir-Wood, H. M. 1962. On the morphology and classification of the brachiopod Suborder Chonetoidea. British Museum Natural History, London, 132 p.Google Scholar
Muir-Wood, H. M. and Cooper, G. A. 1960. Morphology, classification and life habits of the Productoidea (Brachiopoda). Geological Society of America, Memoir 81:1–447.CrossRefGoogle Scholar
Nakazawa, K. and Newell, N. D. 1968. Permian bivalves of Japan. Memoir of the Faculty of Science, Kyoto University Series of Geology and Mineralogy, 35:1–108.Google Scholar
Pan, H. Z. and Erwin, D. H. 1994. Gastropod diversity patterns in South China during the Chihsia-Ladinian and their mass extinction, p. 249–262. In Jin, Y. G., Utting, J., and Wardlaw, B. R. (eds.), Permian stratigraphy, Environments and Resources, Volume 1, Palaeontology and Stratigraphy, Palaeoworld 4. Nanjing University Press, Nanjing.Google Scholar
Plummer, F. B. and Scott, G. 1937. Upper Paleozoic ammonites in Texas. University of Texas Bulletin 3701, 515 p.Google Scholar
Raup, D. M. 1976. Species diversity in the Phanerozoic: An interpretation. Paleobiology, 2:289–297.Google Scholar
Reed, F. R. C. 1944. Brachiopoda and Mollusca of the Productus Limestone of the Salt Range. Palaeontologia Indica, new series, 23(2):1–678.Google Scholar
Rong, J. Y. and Shen, S. Z. 2002. Comparative analysis of the end-Permian and end-Ordovician brachiopod mass extinctions and survivals in South China. Palaeogeography Palaeoclimatology Palaeoecology, 188:25–38.CrossRefGoogle Scholar
Rudwick, M. J. S. 1968. Some analytic methods in the study of ontogeny in fossils with accretionary skeletons. Paleobiological aspects of growth and development, a symposium. Memoir of the Paleontological Society 2, Journal of Paleontology, 42:35–49.Google Scholar
Ruzhentsev, V. E. 1940. Concerning the taxonomic position of some Upper Paleozoic ammonoids. Nauk, 28(3):284–288.Google Scholar
Ruzhentsev, V. E. 1959. Klassifikatsia nadsemeistva Otocerataceae. Paleontologicheskii Zhurnal, 1959(2):56–67.Google Scholar
Ruzhentsev, V. E. and Sarytcheva, T. G. 1965. The development and change of marine organisms at the Paleozoic and Mesozoic boundary. Nauka, Moscow, 108, 431 p.Google Scholar
Sarytcheva, T. G. and Sokolskaja, A. N. 1959. On the classification of the pseudopunctate brachiopods. Doklady Akademia Nauk SSSR, 125:181–184.Google Scholar
Schlotheim, E. F. von. 1816. Beiträge zur Naturgeschichte der Versteinerungen in geognostischer Hinsicht. Leonhard's Taschenbuch für die gesammte Mineralogie, 7(1):3–134.Google Scholar
Schellwien, E. 1903. Palaeozoische und triadische fossilien aus Ostasien. in Futterer's “Durch Asien,” Band 3, 1–150.Google Scholar
Schuchert, C. and LeVene, C. M. 1929. Fossilium catalogus. 1: Animalia, pt. 42: Brachiopoda. W. Junk, Berlin, 140 p.Google Scholar
Scotese, C. R. and Langford, R. P. 1995. Pangea and the paleogeography of the Permian, p. 3–19. In Scholle, P. A., Peryt, T. M., and Ulmer-Scholle, D. S. (eds.), The Permian of Northern Pangea, volume 1: Paleogeography, paleoclimates, stratigraphy. Springer-Verlag, Berlin.Google Scholar
Sepkoski, J. J. Jr. 1976. Species diversity in the Phanerozoic: species area effects. Paleobiology, 2:298–303.CrossRefGoogle Scholar
Sepkoski, J. J. Jr. 1979. A kinetic model of Phanerozoic taxonomic diversity II. Early Phanerozoic families and multiple equilibria. Paleobiology, 5:222–252.Google Scholar
Sepkoski, J. J. Jr. 1981. A factor analytic description of the Phanerozoic marine record. Paleobiology, 7(1):35–53.Google Scholar
Shen, S. Z. and Archbold, N. W. 2001. Chonetoidea (Brachiopoda) from the Lopingian (Late Permian) of South China. Alcheringa, 25(3–4):327–349.Google Scholar
Shen, S. Z., Cao, C. Q., Henderson, C. M., Wang, X. D., Shi, G. R., Wang, W., and Wang, Y. 2006a. End-Permian mass extinction pattern in the northern peri-Gondwanan region. Palaeoworld, 15:3–30.Google Scholar
Shen, S. Z., Grunt, T. A., and Jin, Y. G. 2004. A comparative study of Comelicaniidae Merla, 1930 (Brachiopoda: Athyridida) from the Lopingian (Late Permian) of South China and Transcaucasia in Azerbaijan and Iran. Journal of Paleontology, 78:884–899.Google Scholar
Shen, S. Z., He, X. L., and Zhu, M. L. 1992. Changhsingian brachiopods from Zhongliang Hill of Chongqing, Sichuan, South China, p. 171–196. In Gu, D. Y. (ed.), the symposium on stratigraphy and paleontology of oil and gas bearing areas in China (3). The Petroleum Industry Press, Beijing.Google Scholar
Shen, S. Z. and Shi, G. R. 1996. Diversity and extinction patterns of Permian Brachiopoda of South China. Historical Biology, 12:93–110.CrossRefGoogle Scholar
Shen, S. Z. and Shi, G. R. 2002. Paleobiogeographical extinction patterns of Permian brachiopods in the Asian-western Pacific region. Paleobiology, 28:449–463.Google Scholar
Shen, S. Z. and Shi, G. R. 2007. Lopingian (Late Permian) brachiopods from South China, Pt. 1, Orthotetida, Orthida and Rhynchonellida. Bulletin of the Tohoku University Museum, no. 6:1–102.Google Scholar
Shen, S. Z., Zhang, H., Li, W. Z., Mu, L., and Xie, J. F. 2006b. Brachiopod diversity patterns from Carboniferous to Triassic in South China. Geological Journal, 41:345–361.Google Scholar
Shen, S. Z., Wang, Y., Henderson, C. M., Cao, C. Q., and Wang, W. 2007. Biostratigraphy and lithofacies of the Permian System in the Laibin-Heshan area of Guangxi, South China. Palaeoworld, 16:120–139 Google Scholar
Sheng, J. Z. 1962. Permian stratigraphy of China. Science Record, 4(4):190–195.Google Scholar
Sheng, J. Z. and Jin, Y. G. 1994. Correlation of Permian deposits in China, p. 14–113. In Jin, Y. G., Utting, J., and Wardlaw, B. R. (eds.), Permian stratigraphy, environments and resources, Palaeoworld 4, Volume 1, Nanjing University Press, Nanjing.Google Scholar
Shi, G. R., Shen, S. Z., and Tong, J. N. 1999. Two discrete, possibly unconnected, Permian marine mass extinctions, p. 148–150. In Yin, H. F. and Tong, J. N. (eds.), Proceedings of the International Conference on Pangea and the Paleozoic-Mesozoic Transition. China University of Geosciences Press, Wuhan.Google Scholar
Shi, G. R. and Shen, S. Z. 1997. A Late Permian brachiopod fauna from Selong, Southern Xizang (Tibet), China. Proceedings of the Royal Society of Victoria, 109(1):37–56.Google Scholar
Sowerby, J. 1812–1815. The Mineral Conchology of Great British, Vol. 1, Published by the author, 234 p.Google Scholar
Sowerby, J. 1818. The mineral conchology of Great Britain or coloured figures and descriptions of those remains of Testaceous animals or shells, which have been preserved at various times and depths in the Earth, Vol. 2, Arding and Merrett, London, 251p.Google Scholar
Spath, L. F. 1930. The Eo-Triassic invertebrate fauna of East Greenland. Meddelelser om Grønland, 83(1):1–90.Google Scholar
Stanley, S. M. and Yang, X. 1994. A double mass extinction at the end of the Paleozoic Era. Science, 266:1340–1344.CrossRefGoogle ScholarPubMed
Stehli, F. G. 1954. Lower Leonardian Brachiopoda of the Sierra Diablo. Bulletin of the American Museum of Natural History, 105(3):263–385.Google Scholar
Stehli, F. G. 1956. Notes on oldhaminid brachiopods. Journal of Paleontology, 30(2):305–313.Google Scholar
Sun, Y. C. 1939. The uppermost Permian ammonoids from Kwangsi and their stratigraphical significance. Contributions from the Geological Institute, National University of Peking, 28:35–49.Google Scholar
Termier, G., Termier, H., de Lapparent, A. F., and Marin, P. 1974. Monographie du Permo-Carbonifere de Wardak (Afghanistan central). Documents des Laboratoires de Geologie, Lyon, Hors Serie 2, 1–167.Google Scholar
Tien, C. C. 1929. Study on the stratigraphy of the Upper Palaeozoics in central Hunan. National Research Institute of China. Memoir of the Institute of Geology, (7):69–92.Google Scholar
Tong, Z. X. 1978. Brachiopoda, p. 210–267. In Geological Institute of Southwest China (ed.), Paleontological atlas of southwest China, Sichuan. Vol. 2. Geological Publishing House, Beijing.Google Scholar
Ustritsky, V. I., Chu, B., and Chan, A. Z. 1960. Stratigraphy and fauna of the Carboniferous and Permian deposits of the western part of the Kun-Lun Mountains. Monograph of the Institute of Geology, PRC Minister of Geology, Series Stratigraphy and Paleontology, 5(1):1–132.Google Scholar
Waagen, W. 1883. Salt Range fossils, Volume I, Productus Limestone fossils, Brachiopoda. Memoirs of the Geological Survey of India, Palaeontologia Indica, Series 13, 4(2):391–546.Google Scholar
Waagen, W. 1884. Salt Range fossils, Vol. I, Productus Limestone fossils, Brachiopoda. Memoirs of the Geological Survey of India, Palaeontologia Indica, Series 13, 4(3–4):547–728.Google Scholar
Wang, G. P., Liu, Q. Z., Jin, Y. G., Hu, S. Z., Liang, W. P., and Liao, Z. T. 1982. Brachiopoda, p. 186–256. In Nanjing Institute of Geology and Mineralogy (ed.), Paleontological atlas of southeast China, Late Paleozoic Vol. 2. Geological Publishing House, Beijing.Google Scholar
Wang, H. Y. and Jin, Y. G. 1991. On permianellids (Brachiopoda). Acta Palaeontologica Sinica, 30:481–501.Google Scholar
Wang, X. D. and Sugiyama, T. 2000. Diversity and extinction patterns of Permian coral faunas of China. Lethaia, 33:285–294.Google Scholar
Wang, X. D., Wang, X. J., Zhang, F., and Zhang, H. 2006. Diversity patterns of carboniferous and Permian rugose corals in South China. Geological Journal, 41:329–343.Google Scholar
Wang, Y., Jin, Y. G. and Fang, D. W. 1964. The brachiopod fossils in China. Science Press, Beijing, 777p.Google Scholar
Wanner, J. and Sieverts, D H. 1935. Zur Kenntnis Der Permischen Brachiopoden von Timor. Neues Jahrbuh fur Mineralogie. Geologie und Palaeontologie, Abteilung B, 74:201–281.Google Scholar
Waterhouse, J. B. 1963. Etheripecten, a new aviculopectinid genus from the Permian. New Zealand Journal of Geology and Geophysics, 6:193–196.Google Scholar
Waterhouse, J. B. 1966. Lower Carboniferous and upper Permian brachiopods from Nepal. Jahrbuch der Geologischen Bundesanstalt, 12:5–99.Google Scholar
Waterhouse, J. B. 1975. New Permian and Triassic brachiopod taxa. Papers of the Department of Geology, University of Queensland, 7(2):1–23.Google Scholar
Waterhouse, J. B. 1978. Permian Brachiopoda and Mollusca from northwest Nepal. Palaeontographica Abteilung A, 160(1–6):1–175.Google Scholar
Williams, A. 1965. Brachiopoda. In Moore, R. C. (ed.), Treatise on invertebrate paleontology, Pt. H, Vol. 1–2. Geological Society of America and The University of Kansas Press, Lawrence, 927 p.Google Scholar
Williams, A., Brunton, C. H. C., Carlson, S. J., and Others, W. 2000. Treatise on invertebrate paleontology, Pt. H, Brachiopoda, Vol. 2, 3, Linguliformea, Craniiformea, and Rhynchonelliformea (Pt.). The Geological Society of America, Inc. and The University of Kansas, Lawrence, 919 p.Google Scholar
Yang, D. L. 1984. Brachiopoda, p. 27–102. In Feng, S. N., Xu, S. Y., Lin, J. X., and Yang, D. L. (eds.), Biostratigraphy of the Yangtze George Area (3). Geological Publishing House, Beijing.Google Scholar
Yang, D. L., Ni, S. Z., Chang, M. L., and Chao, R. X. 1977. Brachiopoda, p. 303–470. In Geological Institute of Hubei Province, Geological Bureau of Henan Province, Geological Bureau of Hubei Province, Geological Bureau of Hunan Province, Geological Bureau of Guangdong Province and Geological Bureau of Guangxi Autonomous Region (eds.), Paleontological atlas of central South China 2. Geological Publishing House, Beijing.Google Scholar
Yang, X. N., Zhou, J. P., Liu, J. R., and Shi, G. J. 1999. Evolutionary pattern of fusulinacean foraminifer in Maokouan, middle Permian. Science in China Series D-Earth Sciences, 42:456–464.Google Scholar
Yin, H. F., Wu, S. B., Ding, M. H., Zhang, K. X., Tong, J. N., Yang, F. Q., and Lai, X. L. 1996. The Meishan section, candidate of the global stratotype section and point of Permian-Triassic boundary, p. 31–45. In Yin, H. F. (ed.), The Palaeozoic-Mesozoic boundary candidates of global stratotype section and point of the Permian-Triassic boundary. China University of Geosciences Press, Wuhan.Google Scholar
Zeng, Y., He, X. L., and Zhu, M. L. 1995. Brachiopod communities and their succession and replacement in the Permian of Huayingshan area. China University of Mining and Technology Press, Xuzhou, 187 p.Google Scholar
Zhang, S. X. and Jin, Y. G. 1976. Late Paleozoic brachiopods from the Mount Jolmo Lungma region, p. 159–271. In Xizang scientific expedition team of Chinese Academy of Sciences (ed.), a report of scientific expedition in the Mount Jolmo Lungma Region (1966–1968). Volume 2. Science Press, Beijing.Google Scholar
Zhang, Y. X. 1981. Late Permian bivalves from Yuanjia of Jiahe, Hunan Province. Acta Palaeontogica Sinica, 20(3):260–265.Google Scholar
Zhao, R. X. and Tan, Z. X. 1984. Lower Permian brachiopods from Sangzhi, Hunan. Acta Palaeontologica Sinica 23:30–35.Google Scholar
Zhou, Z. R. 1985. Several problems of the early Permian ammonoids from South China. Palaeontologia Cathayana, 2:179–210.Google Scholar
Zhou, Z. R. 1987. Early Permian ammonite-fauna from southeastern Hunan, p. 285–348. In Nanjing Institute of Geology and Palaeontology (ed.), Nanjing Institute of Geology and Palaeontology, Academia Sinica, Collection of postgraduate theses, No. 1. Jiangsu Science and Technology Publishing House, Nanjing.Google Scholar
Zhou, Z. R. and Gong, Y. H. 1994. On the study of the boundary strata between Guadalupian and Lopingian Series. Permophiles, 24:24–26.Google Scholar
Zhu, T. 1990. The Permian coal-bearing strata and palaeobiocoenosis of Fujian. Geological Publishing House, Beijing, 127 pp.Google Scholar