Hostname: page-component-848d4c4894-ndmmz Total loading time: 0 Render date: 2024-05-24T08:33:25.264Z Has data issue: false hasContentIssue false

Silurian calymenid trilobite Alcymene n. gen., with new species from the Ludlow of Gotland, Sweden

Published online by Cambridge University Press:  20 May 2016

Lars Ramsköld
Museum of Palaeontology, University of Uppsala, Norbyvägen, S-752 36 Uppsala, Sweden
Jonathan M. Adrain
Department of Geology, University of Alberta, Edmonton, Canada T6G 2E3
Gregory D. Edgecombe
Earth and Environmental Sciences, Australian Museum, P.O. Box A285 Sydney South, N.S.W. 2000, Australia
Derek J. Siveter
The University Museum, Parks Road, Oxford OX1 3PW, United Kingdom


Alcymene n. gen. comprises a monophyletic group of calymenids known from the Ludlow Series of Sweden and the British Isles. The sister species Alcymene alveus and A. lindstroemi n. gen. and spp. (Hemse Beds, Gotland, Sweden) are most closely related to the type species A. neointermedia (Richter and Richter), which occurs in overlying and underlying strata at nearby localities. Structural modification of A. alveus and A. lindstroemi, especially the development of a greatly broadened and troughed librigenal border, can be interpreted as “snowshoe” adaptations for increased stability on an offshore mud substrate.

Research Article
Copyright © The Paleontological Society 

Access options

Get access to the full version of this content by using one of the access options below. (Log in options will check for institutional or personal access. Content may require purchase if you do not have access.)


Angelin, N. P. 1854. Palaeontologia Scandinavica. I. Crustacea formationis transitionis. Fascicule 2:2192. Published by the author, Lund.Google Scholar
Bassett, M. G., Kaljo, D., and Teller, L. 1989. The Baltic region, p. 158170. In Holland, C. H. and Bassett, M. G. (eds.), A Global Standard for the Silurian System. National Museum of Wales, Geological Series No. 9.Google Scholar
Brongniart, A., and Desmarest, A. G. 1822. Histoire naturelle des Crustacés fossiles, sous les rapports zoologique et géologique. Savoir: les Trilobites par A. Brongniart. Les Crustacés proprement dits par A. G. Desmarest. Paris, 154 p.Google Scholar
Campbell, K. S. W. 1967. Trilobites of the Henryhouse Formation (Silurian) in Oklahoma. Oklahoma Geological Survey, Bulletin, 115:168.Google Scholar
Chatterton, B. D. E., Johnson, B. D., and Campbell, K. S. W. 1979. Silicified Lower Devonian trilobites from New South Wales. Palaeontology, 22:799837.Google Scholar
Chatterton, B. D. E., Siveter, D. J., Edgecombe, G. D., and Hunt, A. S. 1990. Larvae and relationships of the Calymenina (Trilobita). Journal of Paleontology, 64:255277.Google Scholar
Chlupáč, I. 1987. Ecostratigraphy of Silurian trilobite assemblages of the Barrandian area, Czechoslovakia. Newsletters on Stratigraphy, 17:169186.Google Scholar
Clarkson, E. N. K. 1966. The life attitude of the Silurian trilobite Phacops musheni Salter 1864. Scottish Journal of Geology, 2:7683.Google Scholar
Dalman, J. W. 1827. Om palaeaderna, eller de så kallade trilobiterna. Kungliga Svenska Vetenskaps-Akademiens Handlingar, 1826:113152, 226–294.Google Scholar
Fortey, R. A., and Owens, R. M. 1990. Trilobites, p. 121142. In McNamara, K. J. (ed.), Evolutionary Trends. Belhaven Press, London.Google Scholar
Harrington, H. J., Moore, R. C., and Stubblefield, C. J. 1959. Morphological terms applied to Trilobita, p. 117126. In Moore, R. C. (ed.), Treatise on Invertebrate Palaeontology. Part O, Arthropoda 1. Geological Society of America and University of Kansas Press, Lawrence.Google Scholar
Hede, J. E. 1960. The Silurian of Gotland, p. 4487. In Regnéll, G. and Hede, J. E., The Lower Palaeozoic of Scania. The Silurian of Gotland. Guide to excursions nos A 22 and C 17. International Geological Congress, XXI Session, Norden 1960, Guidebook Sweden d.Google Scholar
Jaanusson, V. 1986. Locality designations in old collections from the Silurian of Gotland. Department of Palaeozoology, Swedish Museum of Natural History, Stockholm, 19 p.Google Scholar
Jaeger, H. 1981. Comments on the graptolite chronology of Gotland, p. 12. In Laufeld, S. (ed.), Proceedings of Project Ecostratigraphy Plenary Meeting, Gotland, 1981. Sveriges Geologiska Undersökning, Rapporter och Meddelanden, 25.Google Scholar
Jell, P. A. 1978. Trilobite respiration and genal caeca. Alcheringa, 2:251260.Google Scholar
Kegel, W. 1928. Über obersilurische Trilobiten aus dem Harz und dem Rheinischen Schiefergebirge. Jahrbuch der preussischen geologischen Landesanstalt, 48 (for 1927):616647.Google Scholar
Kummerov, E. 1928. Beiträge zur Kenntnis der Fauna und der Herkunft der Diluvialgeschiebe. Jahrbuch der preussischen geologischen Landesanstalt, 48 (for 1927):159.Google Scholar
Larsson, K. 1979. Silurian tentaculitids from Gotland and Scania. Fossils and Strata, 11:1180.Google Scholar
Laufeld, S. 1974. Reference localities for palaeontology and geology in the Silurian of Gotland. Sveriges Geologiska Undersökning, Serie C, 705:1172.Google Scholar
Lindström, G. 1861. Bidrag till kännedomen om Gotlands brachiopoder. Öfversigt af Kongliga Vetenskaps-Akademiens Förhandlingar, 17 (for 1860):337382.Google Scholar
Lindström, G. 1885. Förteckning på Gotlands siluriska crustacéer. Öfversigt af Kongliga Vetenskaps-Akademiens Förhandlingar, 42(6):3799.Google Scholar
Männil, R. 1982. Wenlock and Late Silurian trilobite associations of the East Baltic area and their stratigraphical value, p. 6369. In Kaljo, D. and Klaamann, E. (eds.), Ecostratigraphy of the East Baltic Silurian. Academy of Sciences of the Estonian SSR, Institute of Geology, Valgus, Tallinn.Google Scholar
Mikulic, D. G., and Watkins, R. 1981. Trilobite ecology in the Ludlow Series of the Welsh Borderland, p. 101117. In Gray, J., Boucot, A. J., and Berry, W. B. N. (eds.), Communities of the Past. Hutchinson Ross Publishing Company, Stroudsburg, Pennsylvania.Google Scholar
Milne Edwards, H. 1840. Histoire Naturelle des Crustacés, Comprenant l'Anatomie, la Physiologie et la Classification de ces Animaux, 3. Roret, Paris, 638 p.Google Scholar
Münster, G. Graf Zu. 1840. Beiträge zur Petrefaktenkunde III. 7. Die Versteinerungen des Übergangskalkes mit Clymenien und Orthoceratiten von Oberfranken, p. 33121.Google Scholar
Ramsköld, L. 1985a. Silurian phacopid and dalmanitid trilobites from Gotland. Stockholm Contributions in Geology, 40:162.Google Scholar
Ramsköld, L. 1985b. Studies on Silurian trilobites from Gotland, Sweden. University of Stockholm and Swedish Museum of Natural History, Stockholm, 24 p.Google Scholar
Ramsköld, L. 1986. Silurian encrinurid trilobites from Gotland and Dalarna, Sweden. Palaeontology, 29:527575.Google Scholar
Reed, F. R. C. 1920. Description of two trilobites, p. 219222. In Gardiner, C. I. (ed.), The Silurian Rocks of May Hill. Cotteswold Naturalists Field Club, Proceedings, 20:185–222.Google Scholar
Richter, R., and Richter, E. 1954. Die Trilobiten des Ebbe-Sattels und zu vergleichende Arten (Ordovizium, Gotlandium/Devon). Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 488:176.Google Scholar
Rickards, R. B. 1976. The sequence of Silurian graptolite zones in the British Isles. Geological Journal, 11:153188.Google Scholar
Schrank, E. 1970. Calymeniden (Trilobita) aus silurischen Geschieben. Berichte der deutschen Gesellschaft der geologischen Wissenschaften. A. Geologie, Paläontologie, 15:109146.Google Scholar
Shirley, J. 1933. A redescription of the known British Silurian species of Calymene (s.l.). Manchester Literary and Philosophical Society, Memoirs and Proceedings, 77:5167.Google Scholar
Shirley, J. 1936. Some British trilobites of the family Calymenidae. Geological Society of London, Quarterly Journal, 92:384421.Google Scholar
Shirley, J. 1962. Review of the correlation of the supposed Silurian strata of Artois, Westphalia, the Taunas and Polish Podolia, p. 234242. In Symposiums-Band der 2 internationalen Arbeitstagung über die Silur/Devon-Grenze und die Stratigraphie von Silur und Devon.Google Scholar
Siveter, D. J. 1977. The Middle Ordovician of the Oslo region, Norway. 27. Trilobites of the family Calymenidae. Norsk Geologisk Tidsskrift, 56:335396.Google Scholar
Siveter, D. J. 1979. Metacalymene Kegel, 1927, a calymenid trilobite from the Kopanina Formation (Silurian) of Bohemia. Journal of Paleontology, 53:367379.Google Scholar
Siveter, D. J. 1980. Evolution of the Silurian trilobite Tapinocalymene from the Wenlock of the Welsh Borderlands. Palaeontology, 23:783802.Google Scholar
Siveter, D. J. 1983. Calymene lawsoni and allied species from the Silurian of Britain and their stratigraphic significance. Special Papers in Palaeontology, 30:6988.Google Scholar
Swinnerton, H. H. 1915. Suggestions for a revised classification of trilobites. Geological Magazine, New Series, 2:407496, 538–545.Google Scholar
Thayer, C. W. 1975. Morphological adaptations of benthic invertebrates to soft substrata. Journal of Marine Research, 33:177189.Google Scholar
Thomas, A. T. 1980. Trilobite associations in the British Wenlock, p. 447451. In Harris, A. L., Holland, C. H., and Leake, B. E. (eds.), The Caledonides of the British Isles—reviewed. Geological Society of London, Special Publication No 8.Google Scholar
Thomas, A. T., and Fortey, R. A. 1985. Phylum Arthropoda. Subphylum Trilobita, p. 207229. In Murray, J. (ed.), Atlas of Invertebrate Macrofossils. Longman, The Palaeontological Association, London.Google Scholar
Tillman, C. G. 1960. Spathacalymene, an unusual new Silurian trilobite. Journal of Paleontology, 34:891895.Google Scholar
Tomczykowa, E. 1991. Upper Silurian and Lower Devonian trilobites of Poland. Prace Państwowego Institutu Geologicznego, 134:162.Google Scholar
Wahlenberg, G. 1818. Petrificata Telluris Svecanae. Nova Acta Regiae Scientiarum upsaliensis, 8 (for 1821):1116.Google Scholar
whittington, H. B. 1971. Silurian calymenid trilobites from United States, Norway, and Sweden. Palaeontology, 14:455477.Google Scholar
whittington, H. B., and Evitt, W. R. 1954. Silicified Middle Ordovician trilobites. Geological Society of America, Memoir 59:1137.Google Scholar