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A new faunistic component of the Lower Triassic Panchet Formation of India increases the continental non-archosauromorph neodiapsid record in the aftermath of the end-Permian mass extinction

Published online by Cambridge University Press:  22 November 2021

Martín D. Ezcurra Open the ORCID record for Martín D. Ezcurra [Opens in a new window] ,
Saswati Bandyopadhyay  and
Kasturi Sen
Martín D. Ezcurra*
Affiliation:
Sección Paleontología de Vertebrados, CONICET−Museo Argentino de Ciencias Naturales ‘Bernardino Rivadavia’, Ciudad Autónoma de Buenos Aires, Argentina. School of Geography, Earth and Environmental Sciences, University of Birmingham, Edgbaston, Birmingham, UK.
Saswati Bandyopadhyay
Affiliation:
Geological Studies Unit, Indian Statistical Institute, Kolkata, India.
Kasturi Sen
Affiliation:
Geological Studies Unit, Indian Statistical Institute, Kolkata, India. Present Address: C/o Pratap Sen, Trags Electrical Engineering &Air Conditioning Company, B Ring Road, Mezzanine Floor, P O Box 470, Doha, Qatar.
*
*Corresponding author
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Abstract

The fossil record of Early Triassic diapsids is very important to understand how the end-Permian mass extinction affected ecosystems and the patterns and processes involved in the subsequent biotic recovery. Vertebrate fossil assemblages of continental deposits in current-day South Africa, China, and Russia are the best source of information of this clade during the aftermath of the extinction event. Although considerably less sampled, the Induan continental rocks of the Panchet Formation of the Damodar Basin (eastern India) have also yielded a relatively diverse vertebrate assemblage composed of fishes, temnospondyls, synapsids, and a single proterosuchid taxon. Here, we report on a small isolated diapsid partial ilium (ISIR 1132) from the upper Panchet Formation. This specimen has a distinct morphology compared to other tetrapods that we know, including a shallow emargination on the dorsal margin of the anterior portion of the iliac blade, and ratio between height of iliac blade versus maximum height of iliac acetabulum at level of the dorsalmost extension of supraacetabular crest ≤0.45. Comparisons and a quantitative phylogenetic analysis found ISIR 1132 as a non-archosauromorph neodiapsid. This new specimen expands the reptile diversity in the Panchet Formation as well as for the rest of Gondwana, where Early Triassic non-archosauromorph neodiapsid species are extremely scarce.

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Journal of Paleontology , Volume 96 , Issue 2 , March 2022 , pp. 428 - 438
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Copyright © The Author(s), 2021. Published by Cambridge University Press on behalf of The Paleontological Society

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References

Agassiz, L., 1833, Recherches sur les Poissons Fossils: Neuchatel, Petitpierre, v. 2, 336 p.Google Scholar
Ahl, E., 1930, Beitrage zur Lurch und Kriechtierfauna Kwangsi: Section 5, Eidechsen: Sitzungsberichte der Gesellschaft der Naturforschenden Freunde zu Berlin, v. 1930, p. 326–331.Google Scholar
Bakker, R.T., 1977, Tetrapod mass extinctions—a model of the regulation of speciation rates and immigration by cycles of topographic diversity, in Hallan, A., ed. Patterns of Evolution as Illustrated by the Fossil Record: Amsterdam, Elsevier, p. 439–468.CrossRefGoogle Scholar
Bandyopadhyay, S., 1999, Gondwana vertebrate faunas of India: Proceedings of the Indian National Science Academy, v. 65, p. 285–313.Google Scholar
Bandyopadhyay, S., and Ray, S., 2020, Gondwana vertebrate faunas of India: their diversity and intercontinental relationships: Episodes Journal of International Geoscience, v. 43, p. 438–460.Google Scholar
Bartholomai, A., 1979, New lizard-like reptiles from the Early Triassic of Queensland: Alcheringa, v. 3, p. 225–234.CrossRefGoogle Scholar
Bassani, F., 1886, Sui fossili e sull'eta degli schisti bituminosi triasici di Besano in Lombardia: Atti Società Italiana di Scienze Naturali, v. 29, p. 15–72.Google Scholar
Baur, G., 1887, Ueber den Ursprung der Extremitaiten der Ichthyopterygia: Jahresberichte und Mitteilungen des Oberrheinischen Geologischen Vereins, v. 20, p. 17–20.Google Scholar
Benton, M.J., 1985, Classification and phylogeny of the diapsid reptiles: Zoological Journal of the Linnean Society, v. 84, p. 97–164.CrossRefGoogle Scholar
Benton, M.J., Zhang, Q., Hu, S., Chen, Z.Q., Wen, W., Liu, J., Huang, J., Zhou, C., Xie, T., Tong, J., and Choo, B., 2013, Exceptional vertebrate biotas from the Triassic of China, and the expansion of marine ecosystems after the Permo-Triassic mass extinction: Earth-Science Reviews, v. 125, p. 199–243.CrossRefGoogle Scholar
Bonaparte, J.F., 1963, Descripción del esqueleto postcraneano de Exaeretodon (Cynodontia—Traversodontidae): Acta Geológica Lilloana, v. 4, p. 5–52.Google Scholar
Broili, F., and Fischer, E., 1918, Trachelosaurus fischeri nov. gen. nov. sp. ein neuer Saurier aus dem Buntsandstein von Bernburg: Jahrbuch der Königlich Preussischen Geologischen Landesanstalt zu Berlin, v. 37, p. 359–414.Google Scholar
Broom, R., 1903a, On a new reptile (Proterosuchus fergusi) from the Karroo beds of Tarkastad, South Africa: Annals of the South African Museum, v. 4, 159–164.Google Scholar
Broom, R., 1903b, On the skull of a true lizard (Paliguana whitei) from the Triassic beds of South Africa: Records of the Albany Museum, v. 1, p. 1–3.Google Scholar
Broom, R., 1905, Notice of some new reptiles from the Karoo Beds of South Africa: Records of the Albany Museum, v. 1, p. 331–337.Google Scholar
Broom, R., 1914, A new thecodont reptile: Proceedings of the Zoological Society of London, v. 1914, p. 1072–1077.Google Scholar
Butler, R.J., Ezcurra, M.D., Liu, J., Sookias, R.B., and Sullivan, C., 2019, The anatomy and phylogenetic position of the erythrosuchid archosauriform Guchengosuchus shiguaiensis from the earliest Middle Triassic of China: PeerJ, v. 7, e6435. https://doi.org/10.7717/peerj.6435.CrossRefGoogle ScholarPubMed
Carroll, R.L., 1975, Permo-Triassic ‘lizards’ from the Karroo: Palaeontologia Africana, v. 18, p. 71–87.Google Scholar
Carroll, R.L., 1981, Plesiosaur ancestors from the upper Permian of Madagascar: Philosophical Transactions of the Royal Society of London. B, Biological Sciences, v. 293, p. 315–383.Google Scholar
Carroll, R.L., and Gaskill, P., 1985, The nothosaur Pachypleurosaurus and the origin of plesiosaurs: Philosophical Transactions of the Royal Society of London. B, Biological Sciences, v. 309, p. 343–393.Google Scholar
Case, E.C., 1928, A cotylosaur from the Upper Triassic of western Texas: Journal of Washington Academy of Science, v. 18, p. 177–178.Google Scholar
Charig, A.J., and Sues, H.-D., 1976, Proterosuchia, in Kuhn, O., ed. Handbuch der Paläoherpetologie, volume 13. Stuttgart, Gustav Fischer Verlag, p. 11–39.Google Scholar
Chen, Z.Q., and Benton, M.J., 2012, The timing and pattern of biotic recovery following the end-Permian mass extinction: Nature Geoscience, v. 5, p. 375–383.CrossRefGoogle Scholar
Colbert, E.H., 1966, A gliding reptile from the Triassic of New Jersey: American Museum Novitates, v. 3282, p. 1–23.Google Scholar
Colbert, E.H., 1970, The Triassic gliding reptile Icarosauru: Bulletin of the American Museum of Natural History, v. 143, p. 1–142.Google Scholar
Colbert, E.H., 1987, The Triassic reptile Prolacerta in Antarctica: American Museum Novitates, v. 2882, p. 1–19.Google Scholar
Conrad, J.L., 2006, Postcranial skeleton of Shinisaurus crocodilurus (Squamata: Anguimorpha): Journal of Morphology, v. 267, p. 759–775.CrossRefGoogle Scholar
Currie, P.J., 1980, A new younginid (Reptilia: Eosuchia) from the upper Permian of Madagascar: Canadian Journal of Earth Sciences, v. 17, p. 500–511.CrossRefGoogle Scholar
Currie, P.J., 1981, Hovasaurus bolei, an aquatic eosuchian from the Upper Permian of Madagascar: Palaeontologica Africana, v. 24, p. 99–163.Google Scholar
Currie, P.J., 1982, The osteology and relationships of Tangasaurus mennelli Haughton (Reptilia, Eosuchia): Annals of the South African Museum, v. 86, p. 247–265.Google Scholar
Das, D.P., and Gupta, A., 2012, New cynodont record from the lower Triassic Panchet Formation, Damodar valley: Journal of the Geological Society of India, v. 79, p. 175–180.CrossRefGoogle Scholar
De-Oliveira, T.M., Pinheiro, F.L., Da-Rosa, A.A.S., Dias-Da-Silva, S., and Kerber, L., 2020, A new archosauromorph from South America provides insights on the early diversification of tanystropheids: PloS One, v. 15, e0233216. https://doi.org/10.1371/journal.pone.0230890.CrossRefGoogle ScholarPubMed
Druckenmiller, P.S., Kelley, N.P., Metz, E.T., and Baichtal, J., 2020, An articulated Late Triassic (Norian) thalattosauroid from Alaska and ecomorphology and extinction of Thalattosauria: Scientific Reports, v. 10, 1–14.CrossRefGoogle ScholarPubMed
Evans, S.E., 1980, The skull of a new eosuchian reptile from the Lower Jurassic of South Wales: Zoological Journal of the Linnean Society, v. 70, p. 203–264.CrossRefGoogle Scholar
Evans, S.E., 1981, The postcranial skeleton of the Lower Jurassic eosuchian Gephyrosaurus bridensis: Zoological Journal of the Linnean Society, v. 73, p. 81–116.CrossRefGoogle Scholar
Evans, S.E., 1991, The postcranial skeleton of the choristodere Cteniogenys (Reptilia: Diapsida) from the Middle Jurassic of England: Geobios, v. 24, p. 187–199.CrossRefGoogle Scholar
Evans, S.E., 2009, An early kuehneosaurid reptile from the Early Triassic of Poland: Palaeontologia Polonica, v. 65, p. 145–178.Google Scholar
Evans, S.E., and Borsuk-Białynicka, M., 2009, A small lepidosauromorph reptile from the Early Triassic of Poland: Palaeontologia Polonica, v. 65, p. 179–202.Google Scholar
Evans, S.E., and Jones, M.E., 2010, The origin, early history and diversification of lepidosauromorph reptiles, in Bandyopadhyay, S., S., ed. New Aspects of Mesozoic Biodiversity: Berlin, Heidelberg Springer, p. 27–44.CrossRefGoogle Scholar
Ezcurra, M.D., 2016, The phylogenetic relationships of basal archosauromorphs, with an emphasis on the systematics of preterosuchian archosauriforms: PeerJ, v. 4, e1778. https://doi.org/10.7717/peerj.1778.CrossRefGoogle ScholarPubMed
Ezcurra, M.D., and Butler, R.J., 2018, The rise of the ruling reptiles and ecosystem recovery from the Permian-Triassic mass extinction: Proceedings of the Royal Society of London B, v. 285, 20180361. https://doi.org/10.1098/rspb.2018.0361.Google Scholar
Ezcurra, M.D., Scheyer, T., and Butler, R.J., 2014, The origin and early evolution of Sauria, reassessing the Permian saurian fossil record and the timing of the crocodile-lizard divergence: PLoS One, v. 9, e89165. https://doi.org/10.1371/journal.pone.0089165.CrossRefGoogle ScholarPubMed
Ezcurra, M.D., Velozo, P., Meneghel, M., and Piñeiro, G., 2015, Early archosauromorph remains from the Permo-Triassic Buena Vista Formation of north-eastern Uruguay: PeerJ, v. 3, e776. https://doi.org/10.7717/peerj.776.CrossRefGoogle ScholarPubMed
Ezcurra, M.D., Montefeltro, F.C., and Butler, R.J., 2016. The early evolution of rhynchosaurs: Frontiers in Ecology and Evolution, v. 3, 142. https://doi.org/10.3389/fevo.2015.00142.CrossRefGoogle Scholar
Ezcurra, M.D., Fiorelli, L.E., Martinelli, A.G., Rocher, S., von Baczko, M.B., Ezpeleta, M., Taborda, J.R.A., Hechenleitner, E.M., Trotteyn, M.J., and Desojo, J.B., 2017, Deep faunistic turnovers preceded the rise of dinosaurs in southwestern Pangaea: Nature Ecology and Evolution, v. 1, p. 1477–1483.CrossRefGoogle ScholarPubMed
Ezcurra, M.D., Bandyopadhyay, S., Butler, R.J., Sengupta, D., Sen, K., Sookias, R.B., Nesbitt, S.J., 2020, A new proterosuchid archosauriform from the Lower Triassic Panchet Formation of India. 1° Reunión Virtual de Comunicaciones de la Asociación Paleontológica Argentina, Libro de Resúmenes, p. 30–31.Google Scholar
Ezcurra, M.D., Fiorelli, L.E., Trotteyn, M.J., Martinelli, A.G., and Desojo, J.B., 2021a, The rhynchosaur record, including a new stenaulorhynchine taxon, from the Chañares Formation (upper Ladinian–?lowermost Carnian levels) of La Rioja Province, north-western Argentina: Journal of Systematic Palaeontology, v. 18, p. 1907–1938.CrossRefGoogle Scholar
Ezcurra, M.D., Bandyopadhyay, S., and Gower, D.J., 2021b, A new erythrosuchid archosauriform from the Middle Triassic Yerrapalli Formation of south-central India: Ameghiniana, v. 58, p. 132–168.CrossRefGoogle Scholar
Ford, D.P., and Benson, R.B., 2020, The phylogeny of early amniotes and the affinities of Parareptilia and Varanopidae: Nature Ecology & Evolution, v. 4, p. 57–65.CrossRefGoogle ScholarPubMed
Fraser, N.C., 1982, A new rhynchocephalian from the British Upper Trias: Palaeontology, v. 25, p. 709–725.Google Scholar
Fraser, N.C., 1988, The osteology and relationships of Clevosaurus (Reptilia: Sphenodontida): Philosophical Transactions of the Royal Society of London. B, Biological Sciences, v. 321, p. 125–178.Google Scholar
Fraser, N.C., and Walkden, G.M., 1984, The postcranial skeleton of the Upper Triassic sphenodontid Planocephalosaurus robinsonae: Palaeontology, v. 27, 575–595.Google Scholar
Gauthier, J.A., and de Queiroz, K., 2020, Diapsida: in de Queiroz, K., Cantino, P.D., and Gauthier, J.A., eds. Phylonyms A Companion to the PhyloCode: Boca Ratón, London, New York, CRC Press, p. 1033–1040.Google Scholar
Goloboff, P.A., and Catalano, S.A., 2016, TNT version 1.5, including a full implementation of phylogenetic morphometrics: Cladistics, v. 32, p. 221–238.CrossRefGoogle ScholarPubMed
Goloboff, P.A., Farris, J.S., Källersjö, M., Oxelman, B., Ramírez, M.J., and Szumik, C.A., 2003, Improvements to resampling measures of group support: Cladistics, v. 19, p. 324–332.CrossRefGoogle Scholar
Goloboff, P.A., Farris, J.S., and Nixon, K.C., 2008, TNT, a free program for phylogenetic analysis: Cladistics, v. 24, p. 774–786.CrossRefGoogle Scholar
Gottmann-Quesada, A., and Sander, P.M., 2009, A redescription of the early archosauromorph Protorosaurus speneri Meyer, 1832, and its phylogenetic relationships: Palaeontographica Abteilung A, v. 287, 123–220.CrossRefGoogle Scholar
Gow, C.E., 1975, The morphology and relationships of Youngina capensis Broom and Prolacerta broomi Parrington: Palaeontologia Africana, v. 18, p. 89–131.Google Scholar
Gray, J.E., 1831, Note on a particular structure in the head of an Agama: Zoological Miscellany, v. 1, p. 13–14.Google Scholar
Gupta, A., 2009, Ichthyofauna of the Lower Triassic Panchet Formation, Damodar valley basin, West Bengal, and its implications: Indian Journal of Geosciences, v. 63, p. 275–286.Google Scholar
Gupta, A., and Das, D.P., 2011, Report of Lystrosaurus cf. curvatus and L. cf. declivis from the Early Triassic Panchet Formation, Damodar Valley: Indian Journal of Geosciences, v. 65, p. 119–130.Google Scholar
Hamley, T., Cisneros, J.C., and Damiani, R., 2021, A procolophonid reptile from the Lower Triassic of Australia: Zoological Journal of the Linnean Society, v. 192, p. 554–609.CrossRefGoogle Scholar
Harris, J.M., and Carroll, R.L., 1977. Kenyasaurus, a new eosuchian reptile from the Early Triassic of Kenya: Journal of Paleontology, v. 51, p. 139–149.Google Scholar
Huxley, T.H., 1865, On a collection of vertebrate fossils from the Panchet Rocks, Ranigunj, Bengal: Memoir of the Geological Survey of India, v. 3, p. 1–24.Google Scholar
Jalil, N.E., and Janvier, P., 2005, Les pareiasaures (Amniota, Parareptilia) du Permien supérieur du Bassin d'Argana, Maroc: Geodiversitas, v. 27, p. 35–132.Google Scholar
Jaquier, V.P., Fraser, N.C., Furrer, H., and Scheyer, T.M., 2017, Osteology of a new specimen of Macrocnemus aff. M. fuyuanensis (Archosauromorpha, Protorosauria) from the Middle Triassic of Europe: potential implications for species recognition and paleogeography of tanystropheid protorosaurs: Frontiers in Earth Science, v. 5, 91. https://doi.org/10.3389/feart.2017.00091.CrossRefGoogle Scholar
Jenkins, F.A. Jr., 1970, The Chañares (Argentina) Triassic reptile fauna VII. The postcranial skeleton of the traversodontid Massetognathus pascuali (Therapsida, Cynodontia): Breviora, v. 352, p. 1–28.Google Scholar
Jiang, D.Y., Motani, R., Huang, J.D., Tintori, A., Hu, Y. C., Rieppel, O., Fraset, N.C., Ji, C., Kelley, N. P., Fu, W.-L., and Zhang, R., 2016, A large aberrant stem ichthyosauriform indicating early rise and demise of ichthyosauromorphs in the wake of the end-Permian extinction: Scientific Reports, v. 6, 26232. https://doi.org/10.1038/srep26232.CrossRefGoogle ScholarPubMed
Jiang, D.-Y., Lin, W.-B., Rieppel, O., Motani, R., and Sun, Z.-Y., 2019, A new Anisian (Middle Triassic) eosauropterygian (Reptilia, Sauropterygia) from Panzhou, Guizhou Province, China: Journal of Vertebrate Paleontology, v. 38, e1480113. https://doi.org/10.1080/02724634.2018.1480113.Google Scholar
Ketchum, H.F., and Barrett, P.M., 2004, New reptile material from the Lower Triassic of Madagascar: implications for the Permian Triassic extinction event: Canadian Journal of Earth Sciences, v. 41, p. 1–8.Google Scholar
Klembara, J., and Bartík, I., 1999, The postcranial skeleton of Discosauriscus Kuhn, a seymouriamorph tetrapod from the lower Permian of the Boskovice Furrow (Czech Republic): Earth and Environmental Science Transactions of The Royal Society of Edinburgh, v. 90, p. 287–316.CrossRefGoogle Scholar
Kuhn, O., 1966, Die Reptilien, System und Stammesgeschichte: Krailling bei München, Oeben, 154 p.Google Scholar
Lane, H.H., 1945, New mid-Pennsylvanian reptiles from Kansas: Transactions of the Kansas Academy of Science, v. 47, p. 381–390.CrossRefGoogle Scholar
Laurin, M., and Reisz, R.R., 2020, Reptilia, in de Queiroz, K., Cantino, P.D., and Gauthier, J.A., eds., Phylonyms A Companion to the PhyloCode: Boca Ratón, London, New York, CRC Press, p. 1027–1031.Google Scholar
Li, C., Wu, X.-C., Rieppel, O., Wang, L.-T., and Zhao, L.-J., 2008, An ancestral turtle from the Late Triassic of southwestern China: Nature, v. 456, p. 497–501.CrossRefGoogle ScholarPubMed
Li, C., Jiang, D.Y., Cheng, L., Wu, X.C., and Rieppel, O., 2014, A new species of Largocephalosaurus (Diapsida: Saurosphargidae), with implications for the morphological diversity and phylogeny of the group: Geological Magazine, v. 151, 100–120.CrossRefGoogle Scholar
Linnaeus, C., 1758, Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis: Editio decima, reformata. Tomus 1: Stockholm, Laurentius Salvius, 824 p.Google Scholar
Long, R.A., and Murry, P.A., 1995, Late Triassic (Carnian and Norian) tetrapods from the southwestern United States: New Mexico Museum of Natural History and Science Bulletin, v. 4, p. 1–254.Google Scholar
Lydekker, R., 1885, The Reptilia and Amphibia of the Maleria and Denwa groups: Memoirs of the Geological Survey of India, Palaeontologia Indica, Series IV, Indian Pretertiary Vertebrata, v. 1, p. 1–38.Google Scholar
Maidment, S.C.R., Sennikov, A.G., Ezcurra, M.D., Dunne, E.M., Gower, D.J., Hedrick, B., Meade, L.E., Raven, T.J., Paschchenko, D.I., and Butler, R.J., 2020, The postcranial skeleton of the erythrosuchid archosauriform Garjainia prima from the Early Triassic of European Russia: Royal Society Open Science, v. 7, 201089. https://doi.org/10.1098/rsos.201089.CrossRefGoogle ScholarPubMed
Martinelli, A.G., Francischini, H., Dentzien-Dias, P.C., Soares, M.B., and Schultz, C.L., 2017, The oldest archosauromorph from South America: postcranial remains from the Guadalupian (mid-Permian) Rio do Rasto Formation (Paraná basin), southern Brazil: Historical Biology, v. 29, 76–84.CrossRefGoogle Scholar
McGowan, C., and Motani, R., 2003, Handbook of Paleoherpetology, Part 8 Ichthyopterygia: Munich, Verlag Dr. Friedrich Pfeil, 175 p.Google Scholar
Moss, J.L., 1972, The morphology and phylogenetic relationships of the lower Permian tetrapod Tseajaia campi Vaughn (Amphibia: Seymouriamorpha): University of California Publications in Geological Sciences, v. 98, p. 1–72.Google Scholar
Müller, J., 2005, The anatomy of Askeptosaurus italicus from the Middle Triassic of Monte San Giorgio and the interrelationships of thalattosaurs (Reptilia, Diapsida): Canadian Journal of Earth Sciences, v. 42, 1347–1367.CrossRefGoogle Scholar
Nesbitt, S.J., Stocker, M.R., Small, B.J., and Downs, A., 2009, The osteology and relationships of Vancleavea campi (Reptilia: Archosauriformes): Zoological Journal of the Linnean Society, v. 157, p. 814–864.CrossRefGoogle Scholar
Nesbitt, S.J., Flynn, J.J., Prichard, A.C., Parrish, J.M., Ranivoharimanana, L., and Wyss, A.R., 2015, Postcranial anatomy and relationships of Azendohsaurus madagaskarensis: Bulletin of the American Museum of the Natural History, v. 398, p. 1–126.CrossRefGoogle Scholar
Nopsca, F., 1925, Askeptosaurus, ein neues Reptil der Trias von Besano: Centralblatt für Mineralogie, Geologie, und Paläontologie B, v. 1925, p. 265–267.Google Scholar
Nopcsa, F., 1928, Paleontological notes on reptiles: Geologica Hungarica, Series Palaeontologica, v. 1, p. 1–84.Google Scholar
Nosotti, S., and Rieppel, O., 2003, Eusaurosphargis dalsassoi n. gen. n. sp., a new, unusual diapsid reptile from the Middle Triassic of Besano (Lombardy, N Italy): Memorie della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano Ile, v. 31, p. 1–33.Google Scholar
Osborn, H.F., 1903, The reptilian subclasses Diapsida and Synapsida and the early history of the Diaptosauria: Memoirs of the American Museum of Natural History, v. 1, p. 449–519.Google Scholar
Pal, S., 2021, Fossil remains of proterosuchian from the Panchet Formation, Damodar valley of India: Current Science, v. 120, p. 1749–1752.CrossRefGoogle Scholar
Parrington, F.R., 1935, XVI.—On Prolacerta broomi, gen. et sp. n., and the origin of lizards: Annals and Magazine of Natural History, v. 16, p. 197–205.CrossRefGoogle Scholar
Parrington, F.R., 1953, LXXII.—On Aenigmasaurus grallator, gen. et sp. nov., a problematic reptile from the L. Trias: Journal of Natural History, v. 6, p. 721–738.CrossRefGoogle Scholar
Peecook, B.R., Smith, R.M.H., and Sidor, C.A., 2019, A novel archosauromorph from Antarctica and an updated review of a high-latitude vertebrate assemblage in the wake of the end-Permian mass extinction: Journal of Vertebrate Paleontology, v. 38, e1536664. https://doi.org/10.1080/02724634.2018.1536664.CrossRefGoogle Scholar
Peyer, B.D.T., 1955, Die Triasfauna der Tessiner Kalkalpen. XVIII. Helveticosaurus zollingeri, n.g. n.sp.: Schweizerische Paläontologische Abhandlungen, v. 72, p. 3–50.Google Scholar
Pinheiro, F.L., França, M.A.G., Lacerda, M.B., Butler, R.J., and Schultz, C.L., 2016, An exceptional fossil skull from South America and the origins of the archosauriform radiation: Scientific Reports, v. 6, 22817. https://doi.org/10.1038/srep22817.CrossRefGoogle ScholarPubMed
Piveteau, J., 1926, Paleontologie de Madagascar, XIII. Amphibiens et reptiles permiens: Annales de Paleontologie, v. 15, p. 53–180.Google Scholar
Pol, D., and Escapa, I. H., 2009, Unstable taxa in cladistic analysis: identification and the assessment of relevant characters: Cladistics, v. 25, p. 515–527.Google ScholarPubMed
Pritchard, A.C., Turner, A.H., Nesbitt, S.J., Irmis, R.B., and Smith, N.D., 2015, Late Triassic tanystropheids (Reptilia, Archosauromorpha) from northern New Mexico (Petrified Forest Member, Chinle Formation) and the biogeography, functional morphology, and evolution of Tanystropheidae: Journal of Vertebrate Paleontology, v. 35, e911186. https://doi.org/10.1080/02724634.2014.911186.CrossRefGoogle Scholar
Ray, S., 2005, Lystrosaurus (Therapsida, Dicynodontia) from India: taxonomy, relative growth and cranial dimorphism: Journal of Systematic Palaeontology, v. 3, p. 203–221.CrossRefGoogle Scholar
Reisz, R.R., 1981, A diapsid reptile from the Pennsylvanian of Kansas: University of Kansas Publications of the Museum of Natural History, v. 7, p. 1–74.Google Scholar
Rieppel, O., 1995. The genus Placodus: systematics, morphology, paleobiogeography, and paleobiology: Fieldiana (Geology), n.s., v. 31, p. 1–44.Google Scholar
Robinson, P.L., 1958, Some new vertebrate fossils from the Panchet Series of West Bengal: Nature, v. 182, p. 1722–1723.CrossRefGoogle Scholar
Romer, A.S., and Price, L.W., 1940, Review of the Pelycosauria: Geological Society of America Special Papers, v. 28, p. 1–538.CrossRefGoogle Scholar
Sato, T., Cheng, Y.N., Wu, X.C., and Li, C., 2010, Osteology of Yunguisaurus Cheng et al., 2006 (Reptilia; Sauropterygia), a Triassic pistosauroid from China: Paleontological Research, v. 14, p. 179–195.CrossRefGoogle Scholar
Satsangi, P.P., 1964, A note on Chasmatosaurus from the Panchet Series of Raniganj Coalfield, India: Current Science, v. 33, p. 651–652.Google Scholar
Satsangi, P.P., 1987, The vertebrate faunas of the Permian and Lower Triassic sequence of India: Geological Survey of India Special Publication, v. 11, p. 165–178.Google Scholar
Scheyer, T.M., Romano, C., Jenks, J., and Bucher, H., 2014, Early Triassic marine biotic recovery: the predators' perspective: PLoS One, v. 9, e88987. https://doi.org/10.1371/journal.pone.0088987.CrossRefGoogle ScholarPubMed
Scheyer, T.M., Neenan, J.M., Bodogan, T., Furrer, H., Obrist, C., and Plamondon, M., 2017, A new, exceptionally preserved juvenile specimen of Eusaurosphargis dalsassoi (Diapsida) and implications for Mesozoic marine diapsid phylogeny: Scientific Reports, v. 7, 4406. https://doi.org/10.1038/s41598-017-04514-x.CrossRefGoogle ScholarPubMed
Scheyer, T.M., Spiekman, S.N.F., Sues, H.-D., Ezcurra, M.D., Butler, R.J., and Jones, M.E.H., 2020, Colobops: a juvenile rhynchocephalian reptile (Lepidosauromorpha), not a diminutive archosauromorph with an unusually strong bite: Royal Society Open Science, v. 7, 192179. https://doi.org/10.1098/rsos.192179.CrossRefGoogle Scholar
Schoch, R.R., 2006, A complete trematosaurid amphibian from the Middle Triassic of Germany: Journal of Vertebrate Paleontology, v. 26, p. 29–43.CrossRefGoogle Scholar
Schoch, R.R., and Sues, H.-D., 2015, A Middle Triassic stem-turtle and the evolution of the turtle body plan: Nature, v. 523, p. 584–587.CrossRefGoogle ScholarPubMed
Schoch, R.R., and Sues, H.-D., 2018a, A new lepidosauromorph reptile from the Middle Triassic (Ladinian) of Germany and its phylogenetic relationships: Journal of Vertebrate Paleontology, v. 38, e1444619. https://doi.org/10.1080/02724634.2018.1444619.CrossRefGoogle Scholar
Schoch, R.R., and Sues, H.-D., 2018b, Osteology of the Middle Triassic stem-turtle Pappochelys rosinae and the early evolution of the turtle skeleton: Journal of Systematic Palaeontology, v. 16, p. 927–965.CrossRefGoogle Scholar
Sennikov, A.G., 1996, Evolution of the Permian and Triassic tetrapod communities of Eastern Europe: Palaeogeography, Palaeoclimatology, Palaeoecology, v. 120, p. 331–351.CrossRefGoogle Scholar
Sennikov, A.G., 1997, An enigmatic reptile from the upper Permian of the Volga River Basin: Paleontological Journal, v. 31, p. 94–101.Google Scholar
Simões, T.R., Vernygora, O., Caldwell, M.W., and Pierce, S.E., 2020, Megaevolutionary dynamics and the timing of evolutionary innovation in reptiles: Nature Communications, v. 11, 3322. https://doi.org/10.1038/s41467-020-17190-9.CrossRefGoogle ScholarPubMed
Smith, R.M.H., and Botha-Brink, J., 2014, Anatomy of a mass extinction: sedimentological and taphonomic evidence for drought-induced die-offs at the Permo-Triassic boundary in the main Karoo Basin, South Africa: Palaeogeography, Palaeoclimatology, Palaeoecology, v. 396, p. 99–118.CrossRefGoogle Scholar
Spiekman, S.N., Fraser, N.C., and Scheyer, T.M., 2021, A new phylogenetic hypothesis of Tanystropheidae (Diapsida, Archosauromorpha) and other “protorosaurs”, and its implications for the early evolution of stem archosaurs: PeerJ, v. 9, e11143. https://peerj.com/articles/11143.CrossRefGoogle Scholar
Stocker, M.R., Nesbitt, S.J., Kligman, B.T., Paluh, D.J., Marsh, A.D., Blackburn, D.C., and Parker, W.G., 2019, The earliest equatorial record of frogs from the Late Triassic of Arizona: Biology Letters, v. 15, p. 20180922. https://doi.org/10.1098/rsbl.2018.0922.CrossRefGoogle ScholarPubMed
Sulej, T., 2007, Osteology, variability, and evolution of Metoposaurus, a temnospondyl from the Late Triassic of Poland: Palaeontologia Polonica, v. 64, p. 29–139.Google Scholar
Sumida, S.S., 1989, The appendicular skeleton of the early Permian genus Labidosaurus (Reptilia, Captorhinomorpha, Captorhinidae) and the hind limb musculature of captorhinid reptiles: Journal of Vertebrate Paleontology, v. 9, p. 295–313.CrossRefGoogle Scholar
Sumida, S.S., 1997, Locomotor features of taxa spanning the origin of amniotes, in Sumida, S.S., and Martin, K.L.M., eds., Amniote Origins: Completing the Transition to Land: London, Academic Press, p. 353–398.CrossRefGoogle Scholar
Sumida, S.S., Pelletier, V., and Berman, D.S., 2014, New information on the basal pelycosaurian-grade synapsid Oedaleops, in Kammerer, C.F., Angielczyk, K.D., and Fröbisch, J., eds., Early Evolutionary History of the Synapsida: Dordrecht, Springer, p. 7–23.CrossRefGoogle Scholar
Surkov, M.V., Kalandadze, N.N., and Benton, M.J., 2005, Lystrosaurus georgi, a dicynodont from the Lower Triassic of Russia: Journal of Vertebrate Paleontology, v. 25, p. 402–413.CrossRefGoogle Scholar
Tatarinov, L.P., 1960, Discovery of pseudosuchians in the upper Permian of SSSR: Paleontological Journal, v. 1960, p. 74–80. [in Russian]Google Scholar
Thulborn, R.A., 1979, A proterosuchian thecodont from the Rewan Formation of Queensland: Memoirs of the Queensland Museum, v. 19, p. 331–355.Google Scholar
Tripathi, C., and Satsangi, P.P., 1963, Lystrosaurus fauna from the Panchet Series of the Indian Gondwana: Palaeontologica Indica New Series, v. 37, p. 1–53.Google Scholar
Vaughn, P.P., 1955, The Permian reptile Araeoscelis restudied: Bulletin of the Museum of Comparative Zoology, v. 113, p. 305–467.Google Scholar
Viglietti, P.A., Benson, R.B., Smith, R.M., Botha, J., Kammerer, C.F., et al. , 2021, Evidence from South Africa for a protracted end-Permian extinction on land: Proceedings of the National Academy of Sciences, v. 118, e2017045118. https://doi.org/10.1073/pnas.2017045118.CrossRefGoogle ScholarPubMed
von Meyer, H., 1832, Palaeologica zur Geschichte der Erde und ihrer Geschöpfe: Frankfurt am Main, Verlag von Siegmund Schmerber, 560 p.Google Scholar
Warren, A., Damiani, R., and Sengupta, D.P., 2009, Unique stereospondyl mandibles from the early Triassic Panchet formation of India and the Arcadia formation of Australia. Patterns and processes in early vertebrate evolution: Special Papers in Palaeontology, v. 81, p. 161–173.Google Scholar
Watson, D.M.S., 1912, Eosuchus colletti, gen. et spec. nov.: Records of the Albany Museum, v. 2, p. 298–299.Google Scholar
Watson, D.M.S., 1914, Procolophon trigoniceps, a cotylosaurian reptile from South Africa: Proceedings of the Royal Society of London, v. 1914, p. 735–747.Google Scholar
Watson, D.M.S., 1957, On Millerosaurus and the early history of the sauropsid reptiles: Philosophical Transactions of the Royal Society of London. Series B, Biological Sciences, v. 240, p. 325–398.Google Scholar
Williston, S.W., 1910, New Permian reptiles: rhachitomous vertebrae: The Journal of Geology, v. 18, p. 585–600.CrossRefGoogle Scholar
Witzmann, F., and Schoch, R.R., 2006, The postcranium of Archegosaurus decheni, and a phylogenetic analysis of temnospondyl postcrania: Palaeontology, v. 49, p. 1211–1235.CrossRefGoogle Scholar