Hostname: page-component-8448b6f56d-t5pn6 Total loading time: 0 Render date: 2024-04-20T03:03:05.999Z Has data issue: false hasContentIssue false

Geometrical constraints in the construction of graptolite stipes

Published online by Cambridge University Press:  08 April 2016

Richard A. Fortey*
Affiliation:
Department of Palaeontology, British Museum (Natural History), Cromwell Road, London SW7 5BD, England

Abstract

The geometry of the simpler types of graptolite stipes can be quantified by assuming them to consist of stacked cylinders representing the thecal tubes. It can be shown that specification of any three characters, for example, thecal inclination, thecal spacing, and stipe width, effectively defines the others (thecal overlap, thecal length and width, thecal “density”); hence the parameters commonly used in the definition of species are not independent. Variation within and between species can be represented by three-character plots. The effects of altering the apertural angle from 90° on such characters as thecal spacing can also be represented geometrically. If thecae are considered as cones rather than cylinders, it follows that as stipe width increases, thecal curvature will describe a sine curve if the thecae are to remain in contact; this observation is matched on real graptolites. Thecal curvature beyond 90° to the dorsal wall is generally impossible without thecae detaching from their neighbors to produce thecal isolation. Growth of thecae along a stipe can be represented graphically. Computer fits to growth curves for five dichograptids show that they can be closely described by an exponential decrease in growth increments along the stipe, and that this model is more accurate than two possible alternatives.

Type
Research Article
Copyright
Copyright © The Paleontological Society 

Access options

Get access to the full version of this content by using one of the access options below. (Log in options will check for institutional or personal access. Content may require purchase if you do not have access.)

References

Literature Cited

Bouček, B. 1973. Lower Ordovician Graptolites of Bohemia. 164 pp., 24 pls.Akademia, Prague.Google Scholar
Briggs, D. E. G. and Williams, S. H. 1981. The restoration of flattened fossils. Lethaia 14:157164.CrossRefGoogle Scholar
Bulman, O. M. B. 1970. Treatise on Invertebrate Palaeontology. Pt. 5, rev., ed. Teichert, C.163 pp. University of Kansas and Geological Society of America.Google Scholar
Cooper, R. A. and Fortey, R. A. 1982. The Ordovician graptolites of Spitzbergen. Bull. Br. Mus. Nat. Hist. (Geol.) 36:157302.Google Scholar
Erdtmann, B. D. 1970. Morphometrie und Computerstatistik am Beispid der Graptolithengattung Dicellograptus. N. Jahrb. Geol. Paläont. 134, 1969–1970:235266.Google Scholar
Hutt, J. 1975. The Llandovery graptolites of the English Lake District, Pt. 2. Paleontogr. Soc. Monogr. Pp. 57137, pls. 11–26.Google Scholar
Mu, A-T., Ge, M. Y., Chen, X., et al. 1979. Lower Ordovician graptolites of Southwest China Palaeont. Sinica, 156B(13):1192. [Chinese, English summary]Google Scholar
Rickards, R. B., Hutt, J., and Berry, W. B. N. 1977. Evolution of the Silurian and Devonian graptoloids. Bull. Br. Mus. Nat. Hist. (Geol.) 28:1120, pls. 1–6.Google Scholar
Rickards, R. B., Hutt, J., and Berry, W. B. N. 1970. The Llandovery (Silurian) graptolites of the Howgill Fells, Northern England. Palaeontogr. Soc. Monogr. 108 pp., 8 pls.Google Scholar
Ruedemann, R. 1947. Graptolites of North America. Mem. Geol. Soc. Am. 19:1652.Google Scholar
Urbanek, A. 1973. Organisation and evolution of graptolite colonies. In: Boardman, R. S., et al., eds. Animal Colonies: Their Development and Function through Time. Dowden, Hutchinson & Ross, New York.Google Scholar