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Cambrian trilobites and biostratigraphy of the Grand Canyon and vicinity, USA

Published online by Cambridge University Press:  02 December 2025

Frederick Sundberg*
Affiliation:
Department of Earth and Planetary Sciences, University of New Mexico, Albuquerque, NM 87131, USA
John R. Foster
Affiliation:
Utah Field House of Natural History State Park Museum, 496 E. Main St., Vernal, UT 84078, USA
Mark Webster
Affiliation:
Department of the Geophysical Sciences, University of Chicago, 5734 South Ellis Avenue, Chicago, IL 60637, USA
Anne Miller
Affiliation:
Geosciences Program, Science and Resource Management, Grand Canyon National Park, 1824 S Thompson St. Ste. 200, Flagstaff, AZ 86001, USA
James Hagadorn
Affiliation:
Denver Museum of Nature and Science, 2001 Colorado Blvd., Denver, CO 80205, USA
*
Corresponding author: Frederick Sundberg; Email: sundbergfred@gmail.com

Abstract

Cambrian faunas of the Grand Canyon are neither diverse nor abundant, and several historical collections seem to have mixed taxa from several biozones. As a result, fine-scale reinterpretation of Grand Canyon biostratigraphy and broader-scale correlation of the Tonto Group and its fossils have been hampered by the lack of definitive and refined biostratigraphic indicators. Here we report new fossil occurrences from the Tonto Group, including from units that previously lacked biomineralized fossils, such as the Frenchman Mountain Dolostone. When integrated with historic collections, our data indicate that the Tonto Group ranges from the upper Dyeran to lowermost Steptoean Laurentian stages (global upper Series 2, Stage 4 to basal Furongian Series, Paibian Stage). Trilobite assemblages span the upper Dyeran (Peachella iddingsi Biozone) to lower Marjuman (Bolaspidella Biozone) stages and are distributed from the uppermost Tapeats Sandstone to lowermost Frenchman Mountain Dolostone.

Based on lithologic and paleontological data, three disconformities occur within the Cambrian of the Grand Canyon: (1) between the Dyeran and Delamaran stages in the uppermost Tapeats to lowermost Bright Angel Formation, (2) within the Delamaran Stage within the Bright Angel Formation, and (3) between the Delamaran and Topazan stages, higher in the Bright Angel Formation.

Cambrian trilobites from the Tonto Group consist of seven olenelloid, 17 corynexochid, and 33 ptychoparioid species, as well as 10 taxa that are not identifiable at generic or higher levels. New species include Kootenia barensis and Albertella nebeli. Some taxa are new to the Grand Canyon region, although they are known from the Great Basin (Nevada, California), southern Arizona, Montana, and Canada. These taxa include olenellids: Mesonacis fremonti (Walcott), Olenellus gilberti Meek in White, Olenellus terminatus Palmer, and Olenellus cf. O. fowleri Palmer; corynexochids: Anoria? bessus (Walcott), Caspimexis cf. C. radiatus (Palmer in Palmer and Halley), Kootenia aff. K. granulospinosa Palmer, 1968, and Zacanthoides cf. Z. typicalis (Walcott); and ptychopariids: Amecephalus cf. Am. normale (Resser), Am. piochensis (Walcott), Am. scapegoatensis (Deiss), Crepicephalus? upis Walcott, Ehmania weedi Resser, Eldoradia cf. E. prospectensis (Walcott), Glyphaspis tetonensis Resser, Kochina vestita Resser, Mexicella cf. Me. robusta Sundberg and McCollum, Mexicella mexicana Lochman, Modocia sp. indet., Solenopleurella? quadrata Rasetti, Trachycheilus granulosus Sundberg, and Tr. whirlwindensis Sundberg. Type specimens of Solenopleurella erosa Resser was divided into two species: the pygidia are assigned to Kootenia erosa (Resser) new combination, and the much smaller cranidia are assigned to Spencella? sp. indet.

UUID: http://zoobank.org/f3034b33-00d1-4d00-a9d5-0a73f7a330d2

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Memoir
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© The Author(s), 2025. Published by Cambridge University Press on behalf of Paleontological Society

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References

Adrain, J.M., 2011, Class Trilobita Walch, 1771, in Zhang, Z.Q., ed., Animal biodiversity: an outline of higher-level classification and survey of taxonomic richness: Zootaxa, v. 3148, p. 104109.10.11646/zootaxa.3148.1.15CrossRefGoogle Scholar
Angelin, N.P., 1854, Palaeontologia Scandinavica. Pars II. Crustacea Formationis Transitionis: Holmiae Stockholm, Norstedt and Söner, p. 2192.Google Scholar
Babcock, L.E., Robison, R.A., Rees, M.N., Peng, S.C., and Saltzman, M.R., 2007, The Global boundary Stratotype Section and Point (GSSP) of the Drumian Stage (Cambrian) in the Drum Mountains, Utah, USA: Episodes, v. 30, p. 8494.Google Scholar
Bordonaro, O.L., Pratt, B.R., and Robledo, V., 2013, Systematic, morphometric and palaeobiogeographic study of Blainia gregaria Walcott, 1916 (Trilobita, Ptychopariida) Middle Cambrian of the Precordillera of western Argentina: Geological Journal, v. 48, p. 126141.10.1002/gj.1344CrossRefGoogle Scholar
Brathovde, J.E., 1986, Stratigraphy of the Grand Wash Dolomite (Upper? Cambrian), Western Grand Canyon, Mohave Count, Arizona [MS thesis]: Flagstaff, Northern Arizona University, 140 p.Google Scholar
Campbell, D.P., 1974, Biostratigraphy of the Albertella and Glossopleura zones (Lower Middle Cambrian) of northern Utah and southern Idaho [M.S. thesis]: Salt Lake City, University of Utah, 295 p.Google Scholar
Clark, C.W., 1921, Lower and middle Cambrian formations of the Mohave Desert: University of California Publications Bulletin of the Department of Geological Sciences, v. 13, p. 17.Google Scholar
Cooper, G.A., Arellano, A.R.V., Johnson, J.H., Okulitch, V.J., Stoyanow, A., and Lochman, C., 1952, Cambrian stratigraphy and paleontology near Caborca, northwestern Sonora, Mexico: Smithsonian Miscellaneous Collections, v. 119, p. 1184.Google Scholar
Dehler, C., Sundberg, F., Karlstrom, K., Crossey, L., Schmitz, M., Rowland, S., and Hagadorn, J., 2024, The Cambrian of the Grand Canyon: refinement of a classic stratigraphic model: GSA Today, v. 34, no. 11, p. 411.10.1130/GSATG604A.1CrossRefGoogle Scholar
Deiss, C., 1939, Cambrian stratigraphy and trilobites of northwestern Montana: Geological Society of America, Special Paper, no. 18, p. 1135.10.1130/SPE18-p1CrossRefGoogle Scholar
Derby, J.R., 1965, Paleontology and stratigraphy of the Nolichucky Formation in southwest Virginia and northeast Tennessee [Ph.D. dissertation]: Virginia Polytechnic Institute and State University, Blacksburg, 468 p.Google Scholar
Eby, R.G., 1981, Early late Cambrian trilobite faunas of the Big Horse Limestone and correlative units in central Utah and Nevada [Ph.D. dissertation]: Stony Brook, State University of New York, 613 p.Google Scholar
Eddy, J.D., and McCollum, L.B., 1998, Early middle Cambrian Albertella Biozone trilobites of the Pioche Shale, southeastern Nevada: Journal of Paleontology, v. 72, p. 864887.10.1017/S0022336000027207CrossRefGoogle Scholar
Esteve, J., Sundberg, F.A., Zamora, S., and Gozalo, R., 2012, A new Alokistocaridae Resser, 1939 (Trilobita) from the middle Cambrian of Spain: Geobios, v. 45, p. 275283.Google Scholar
Foster, J.R., 2011, Article 6. Trilobites and other fauna from two quarries in the Bright Angel Shale (middle Cambrian, Series 3; Delamaran), Grand Canyon National Park, Arizona, in Hollingsworth, J.S., Sundberg, F.A., and Foster, J.R., eds., Cambrian Stratigraphy and Paleontology of Northern Arizona and Southern Nevada, The 16th Field Conference of the Cambrian Stage Subdivision Working Group, International Subcommission On Cambrian Stratigraphy Flagstaff, Arizona, And Southern Nevada, United States: Museum of Northern Arizona Bulletin, no. 67, p. 99120.Google Scholar
Foster, J.R., 2022, Continuous character variation within the GlossopleuraAnoriaSonoraspis plexus: dolichometopid trilobites from the Cadiz Formation (Cambrian: Miaolingian, Wuliuan), California: Palaeontologica Electronica, 25.2.a18, https://doi.org/10.26879/1173CrossRefGoogle Scholar
Foster, J.R., Sundberg, F.A., and Hagadorn, J.H., 2024, Cambrian trilobites and associated fossils from the Uinta Mountains of Utah (USA): Journal of Paleontology, v. 97, p. 452459.10.1017/jpa.2023.82CrossRefGoogle Scholar
Fritz, W.H., 1968, Lower and early middle Cambrian trilobites from the Pioche Shale, east-central Nevada, U.S.A: Palaeontology, v. 11, p. 183235.Google Scholar
Fryxell, J.E., and Duebendorfer, E.M., 2005, Origin and trajectory of the Frenchman Mountain Block, an extensional allochthon in the Basin and Range Province, southern Nevada: Journal of Geology, v. 113, p. 355371.10.1086/428810CrossRefGoogle Scholar
Gunther, L.F., and Gunther, V.G., 1981, Some middle Cambrian fossils of Utah: Brigham Young University Geology Studies, v. 28, 87 p.Google Scholar
Hagadorn, J.W., Kirschvink, J. L., Raub, T.D., and Rose, E.C., 2011, Above the Great Unconformity: a fresh look at the Tapeats Sandstone, Arizona-Nevada, USA., in Hollingsworth, J.S., Sundberg, F.A., and Foster, J.R., eds., Cambrian Stratigraphy and Paleontology of Northern Arizona and Southern Nevada, The 16th Field Conference of the Cambrian Stage Subdivision Working Group, International Subcommission On Cambrian Stratigraphy Flagstaff, Arizona, And Southern Nevada, United States: Museum of Northern Arizona Bulletin, no. 67, p. 6377.Google Scholar
Hall, J., 1859, Trilobites of the shales of the Hudson River Group, in Twelfth Annual Report of the New York State Cabinet for Natural History: Albany, C. Van Benthuysen, p. 5962.Google Scholar
Hall, J., 1862, Supplementary note to the thirteenth report of the Regents on the State Cabinet, in Fifteenth Annual Report of the Regents of the University of the State of New-York, on the condition of the State Cabinet of Natural History, and the Historical and Antiquarian Collection annexed thereto: Albany, Charles Van Benthuysen, p. 114.Google Scholar
Hall, J., and Whitfield, R.P., 1877, Paleontology: United States Geologic Exploration of the 48th Parallel, v. 4, p. 197302.Google Scholar
Handkamer, N.M., Pratt, B.R., and MacNaughton, R.B., 2022, Biostratigraphy and paleoecology of the trilobite faunas from the Mount Clark and Mount Cap formations (early and middle Cambrian), eastern Mackenzie Mountains, northwestern Canada: Journal of Paleontology, v. 96, S89, p. 147, https://doi.org/10.1017/jpa.2022.13Google Scholar
Handkamer, N.M., Ichaso, A., Pratt, B.R., Mángano, M.G., and Buatois, L.A., 2023, Systematics and biostratigraphy of a new trilobite fauna collected from the subsurface Earlie Formation (Wuliuan Stage, Miaolingian Series, Cambrian) in southwestern Saskatchewan: Canadian Journal of Earth Sciences, v. 60, p. 13071326, https://doi.org/10.1139/cjes-2023-0003Google Scholar
Hardy, J.K., 1986, Stratigraphy and depositional environments of Lower and Middle Cambrian strata in the Lake Mead region, southern Nevada and northwestern Arizona [M.S. thesis]: Las Vegas, University of Nevada Las Vegas, 309 p.Google Scholar
Harrington, H.J., 1956, Olenellidae with advanced cephalic spines: Journal of Paleontology, v. 30, p. 5661.Google Scholar
Harrington, H.J., Henningsmoen, G., Howell, B.F., Jaanusson, V., Lochman-Balk, C., et al., 1959, Arthropoda 1. Part, O, Moore, R.C., ed., Treatise on Invertebrate Paleontology: Boulder, Colorado and Lawrence, Kansas, Geological Society of America and University of Kansas Press, 560 p.Google Scholar
Hupé, P., 1955, Classification des trilobites: Annales de Paléontologie, v. 41, p. 91325.Google Scholar
Jell, P.A., and Adrain, J.M., 2003, Available generic names for trilobites: Memoirs of the Queensland Museum, v. 48, p. 331553.Google Scholar
Karlstrom, K.E., Hagadorn, J., Gehrels, G.G., Mathews, W., Schmitz, M., Madronich, L., Mulder, J., Pecha, M., Geisler, D., and Crossey, L.J., 2018, Cambrian Sauk transgression in the Grand Canyon region redefined by detrital zircons: Nature Geoscience v. 11, p. 438443.10.1038/s41561-018-0131-7CrossRefGoogle Scholar
Karlstrom, K.E., Mohr, M.T., Schmitz, M.D., Sundberg, F.A., Rowland, S., Hagadorn, J., Foster, J.R., Crossey, L.J., Dehler, C., and Blakey, R., 2020, Redefining the Tonto Group of Grand Canyon and recalibrating the Series 2–Miaolingian Epoch boundary of the Cambrian timescale: Geology, v. 48, p. 425430.10.1130/G46755.1CrossRefGoogle Scholar
Kobayashi, T., 1935, The Cambro-Ordovician formations and faunas of South Chosen, Palaeontology, Part III, Cambrian faunas of South Chosen with special study on the Cambrian trilobite genera and families: Journal of the Faculty of Science, Imperial University of Tokyo, v. 4, no. 2, p. 49344.Google Scholar
Korolev, V.S., 1997, Sequence stratigraphy, sedimentology, and correlation of the undifferentiated Cambrian dolomites of the Grand Canyon and Lake Mead area [M.S. thesis]: Las Vegas, University of Nevada Las Vegas, 178 p.Google Scholar
Łabaj, M., 2015, Depositional dynamics in a mixed carbonate–siliciclastic system, trilobite fauna, biostratigraphy and biofacies: middle–upper Cambrian Abrigo Formation, southeastern Arizona [Ph.D. dissertation]: Saskatoon, University of Saskatchewan, 259 p.Google Scholar
Landing, E., Schmitz, M.D., Westrop, S.R., and Geyer, G., 2023, U–Pb zircon dates from North American and British Avalonia bracket the lower–middle Cambrian boundary interval, with evaluation of the Miaolingian Series as a global unit: Geological Magazine, v. 16, p. 17901816.10.1017/S0016756823000729CrossRefGoogle Scholar
Lerosey-Aubril, R., and Peel, J.S., 2018, Gut evolution in early Cambrian trilobites and the origin of predation on infaunal macroinvertebrates: evidence from muscle scars in Mesolenellus: Palaeontology, v. 61, p. 747760.Google Scholar
Levi-Setti, R., 1993, Trilobites, 2nd edition: Chicago, The University of Chicago Press, 342 p.Google Scholar
Lieberman, B.S., 1999 , Systematic revision of the Olenelloidea (Trilobita, Cambrian): Bulletin of the Peabody Museum of Natural History, v. 35, p. 1150.Google Scholar
Lin, J.P., Sundberg, F.A., Jiang, G., Montañez, I.P., and Wotte, T., 2019, Chemostratigraphic correlations across the first major trilobite extinction and faunal turnovers between Laurentia and South China: Scientific Reports, v. 9, 17392, https://doi.org/10.1038/s41598-019-53685-2CrossRefGoogle ScholarPubMed
Lochman, C., 1948, New Cambrian trilobite genera from northwest Sonora, Mexico: Journal of Paleontology, v. 22, p. 451467.Google Scholar
Lochman, C., and Hu, C.-H., 1960, Upper Cambrian faunas from the northwest Wind River Mountains, Wyoming, part 1: Journal of Paleontology, v. 34, p. 793834.Google Scholar
McCollum, L.B., and Sundberg, F.A., 2007, Cambrian trilobite biozonation of the Laurentian Delamaran Stage in the southern Great Basin, U.S.A.: implications for global correlations and defining a Series 3 global boundary stratotype: Memoirs of the Association of Australasian Palaeontologists, v. 34, p. 147156.Google Scholar
McCollum, M.B., and McCollum, L.B., 2011, Depositional sequences in the Laurentian Delamaran Stage, southern Great Basin, USA, in Hollingsworth, J.S., Sundberg, F.A., and Foster, J.R., eds., Cambrian Stratigraphy and Paleontology of Northern Arizona and Southern Nevada, The 16th Field Conference of the Cambrian Stage Subdivision Working Group, International Subcommission on Cambrian Stratigraphy Flagstaff, Arizona, And Southern Nevada, United States: Museum of Northern Arizona Bulletin, no. 67, p. 154175.Google Scholar
McKee, E.D., 1945, Cambrian history of the Grand Canyon region, part I. Stratigraphy and ecology of the Grand Canyon Cambrian, in McKee, E.D., and Resser, C.E., Cambrian history of the Grand Canyon region: Carnegie Institution of Washington Publication, v. 563, p. 1170.Google Scholar
McKee, E.D., and Resser, C.E., 1945, Cambrian history of the Grand Canyon region: Carnegie Institution of Washington Publication 563, 232 p.Google Scholar
Meek, F.B., 1870, Descriptions of fossils collected by the U. S. Geological Survey, under charge of Clarence King: Proceedings of the Academy of Natural Sciences of Philadelphia, ser. 2, v. 14, p. 5664.Google Scholar
Meek, F.B., 1873, Preliminary palaeontology report, consisting of list and descriptions of fossils, with remarks on the age of the rocks in which they are found: Annual Report of the United States Geological Survey of Territories no. 6, p. 429518.Google Scholar
Meek, F.B., 1877, Palaeontology: United States Geological Exploration of the Fortieth Parallel. Clarence King, Geologist-in-charge, v. 4, p. 1197.Google Scholar
Meek, F.B., and Hayden, F.V., 1861, Descriptions of new Lower Silurian (Primordial), Jurassic, Cretaceous and Tertiary fossils collected in Nebraska by the exploring expedition under the command of Capt. Wm. F. Reynolds, U.S. Top. Engrs; with remarks on the rocks from which they were obtained: Proceedings of the Academy of Natural Science, Philadelphia, ser. 2, v. 5, p. 415447.Google Scholar
Melzak, A., and Westrop, S.R., 1994, Mid-Cambrian (Marjuman) trilobites from the Pika Formation, southern Canadian Rocky Mountains, Alberta: Canadian Journal of Earth Sciences, v. 31, p. 969985.10.1139/e94-086CrossRefGoogle Scholar
Miller, B.M., 1936, Cambrian trilobites from northwestern Wyoming: Journal of Paleontology, v. 10, p. 2334.Google Scholar
Miller, J.F., Evans, K.R., and Dattilo, B.F., 2012, The Great Carbonate Bank in the miogeocline of western central Utah: tectonic influences on sedimentation, in Derby, J.R., Fritz, R.D., Longacre, S.A., Morgan, W.A., and Sternbach, C.A., eds., The great American carbonate bank: the geology and economic resources of the Cambrian–Ordovician Sauk megasequence of Laurentia: American Association of Petroleum Geologists, Memoir 98, p. 769854.Google Scholar
Morgan, C.A., 2021, Quantitative trilobite biostratigraphy of the middle Cambrian (Wuliuan Stage; Miaolingian Series) and microfacies analysis of the middle Cambrian Stephen formation, Southern Canadian Rocky Mountains [Ph.D. dissertation]: Calgary, University of Calgary, 445 p.Google Scholar
Owen, D.D., 1852, Report of a Geological Survey of Wisconsin, Iowa, and Minnesota, and Incidentally a Portion of Nebraska Territory: Philadelphia, Lippincot, Grambo and Co., 638 p.Google Scholar
Özdikmen, H., 2005, Homonymous generic names in trilobites based on Nomenclator Zoologicus, vols. 1–9, A to M: Paleontological Journal, v. 39, p. 563564.Google Scholar
Pack, F.J., 1906, Cambrian fossils from the Pioche Mountains, Nevada: Journal of Geology, v. 14, p. 290302.Google Scholar
Pack, P.D., and Gayle, H.B., 1971, A new olenellid trilobite, Biceratops nevadensis, from the lower Cambrian near Las Vegas, Nevada: Journal of Paleontology, v. 45, p. 893898.Google Scholar
Palmer, A.R., 1954, An appraisal of the Great Basin middle Cambrian trilobites described before 1900: United States Geological Survey, Professional Paper 264D, p. 5385.Google Scholar
Palmer, A.R., 1968, Cambrian trilobites of east-central Alaska: United States Geological Survey, Professional Paper 559-B, 115 p.Google Scholar
Palmer, A.R., 1998, Terminal early Cambrian extinction of the Olenellina: documentation from the Pioche Formation, Nevada: Journal of Paleontology, v. 72, p. 650672.10.1017/S0022336000040373CrossRefGoogle Scholar
Palmer, A.R., and Halley, R.B., 1979, Physical stratigraphy and trilobite biostratigraphy of the Carrara Formation (lower and middle Cambrian) in the southern Great Basin: United States Geological Survey Professional Paper 1047, 131 p.Google Scholar
Palmer, A.R., and Repina, L.N., 1993, Through a glass darkly: taxonomy, phylogeny and biostratigraphy of the Olenellina: The University of Kansas Paleontological Contributions, no. 3, 35 p.Google Scholar
Pari, G., Briggs, D.E.G., Gaines, R.R., Roach, B.T., and Webster, M., 2023, Exceptional lower Cambrian fossils from a long-lost locality in Vermont, USA: Geology Today, v. 39, p. 152157.10.1111/gto.12444CrossRefGoogle Scholar
Pates, S., and Bicknell, R.D.C., 2019, Elongated thoracic spines as potential predatory deterrents in olenelline trilobites from the lower Cambrian of Nevada: Palaeogeography, Palaeoclimatology, Palaeoecology, v. 516, p. 295306.10.1016/j.palaeo.2018.12.013CrossRefGoogle Scholar
Peel, J.S., 2020, Middle Cambrian trilobites (Miaolingian, Ehmaniella Biozone) from the Telt Bugt Formation of Daugaard-Jensen Land, western North Greenland: Bulletin of the Geological Society of Denmark, v. 68, p. 114.Google Scholar
Poulsen, C., 1927, The Cambrian, Ozarkian and Canadian faunas of northwest Greenland: Meddelelser om Grønland, v. 70, p. 233343.Google Scholar
Poulsen, C., 1958, Contribution to the palaeontology of the lower Cambrian Wulff River Formation: Meddelelser om Grønland, v. 162, no. 2, p. 125.Google Scholar
Pratt, B.R., 1992, Trilobites of the Marjuman and Steptoean stages (upper Cambrian), Rabbitkettle Formation, southern Mackenzie Mountains, northwest Canada: Palaeontographica Canadiana, no. 9, p. 1179.Google Scholar
Qiu, H., Lu, Y.H., Zhu, Z.L., Bi, D.C., Lin, T.R., et al., 1983: Trilobita, in Palaeontological Atlas of East China, v. 1: Beijing, Geological Publishing House, p. 28–254; 574610. [in Chinese]Google Scholar
Rasetti, F., 1951, Middle Cambrian stratigraphy and faunas of the Canadian Rocky Mountains: Smithsonian Miscellaneous Collection, v. 116, no. 5, p. 1277.Google Scholar
Rasetti, F., 1963, Middle Cambrian ptychoparioid trilobites from the conglomerates of Quebec: Journal of Paleontology, v. 37, p. 575594.Google Scholar
Rasetti, F., 1965, Upper Cambrian trilobite faunas of northeastern Tennessee: Smithsonian Miscellaneous Collections v. 148, no. 3, p. 1127.Google Scholar
Raw, F., 1936, Mesonacidae of Comley in Shropshire, with a discussion of classification within the family: Quarterly Journal of the Geological Society of London, v. 92, p. 236293.Google Scholar
Raymond, P.E., 1924, New upper Cambrian and Lower Ordovician trilobites from Vermont: Proceedings of the Boston Society of Natural History, v. 37, p. 389466.Google Scholar
Raymond, P.E., 1928, Two new Cambrian trilobites: American Journal of Science, v. 5, p. 309313.Google Scholar
Reed, F.R.C., 1910, The Cambrian Fossils of Spiti: Palaeontologia Indica: The Geological Survey of India, ser. 15, v. 7, Memoir 1, 70 p.Google Scholar
Resser, C.E., 1935, Nomenclature of some Cambrian trilobites: Smithsonian Miscellaneous Collections, v. 93, no. 5, 46 p.Google Scholar
Resser, C.E., 1936, Second contribution to nomenclature on Cambrian trilobites: Smithsonian Miscellaneous Collections, v. 95, no. 4, 29 p.Google Scholar
Resser, C.E., 1937, Third contribution to nomenclature of Cambrian trilobites: Smithsonian Miscellaneous Collections, v. 95, no. 22, 29 p.Google Scholar
Resser, C.E., 1938a. Cambrian System (restricted) of the southern Appalachians: Geological Society of America Special Papers 15, p. 1140.Google Scholar
Resser, C.E., 1938b, Middle Cambrian fossils from Pend Oreille Lake, Idaho: Smithsonian Miscellaneous Collections, v. 97, no. 3, 12 p.Google Scholar
Resser, C.E., 1939a, The Spence Shale and its fauna: Smithsonian Miscellaneous Collections, v. 97, 29 p.Google Scholar
Resser, C.E., 1939b, The Ptarmigania strata of the Northern Wasatch Mountains: Smithsonian Miscellaneous Collections, v. 98, 72 p.Google Scholar
Resser, C.E., 1945, Part II. Cambrian fossils of the Grand Canyon, in McKee, E.D., and Resser, C.E., Cambrian history of the Grand Canyon region: Carnegie Institution of Washington , Publication 563, p. 171232.Google Scholar
Resser, C.E., and Howell, B.F., 1938, Lower Cambrian Olenellus Zone of the Appalachians: Bulletin of the Geological Society of America, v. 49, p. 195248.10.1130/GSAB-49-195CrossRefGoogle Scholar
Richter, R., 1932, Crustacea (Paläontologie), in Dittler, R., Joos, G., Korschelt, E., Linek, G., Oltmanns, F., and Schaum, K. eds., Handwörterbuch der Naturwissenschaften, 2nd ed.: Jena, Gustav Fisher, p. 840846.Google Scholar
Robison, R.A., 1964, Late middle Cambrian faunas from western Utah: Journal of Paleontology v. 38, p. 510566.Google Scholar
Robison, R.A., and Babcock, L.E., 2011, Systematics, paleobiology, and taphonomy of some exceptionally preserved trilobites from Cambrian Lagerstätten of Utah: Paleontological Contributions, Paleontological Institute, University of Kansas, no. 5, 47 p.Google Scholar
Robison, R.A., Babcock, L.E., and Gunther, V.G., 2015, Exceptional Cambrian fossils from Utah: a window into the age of trilobites: Utah Geological Survey Miscellaneous Publication 15-1, 97 p., https://doi.org/10.34191/mp-15-1Google Scholar
Rominger, C., 1887, Description of primordial fossils from Mt. Stephens, N.W. Territory of Canada: Proceedings Academy of Natural Sciences Philadelphia, 1887, p. 1219.Google Scholar
Rose, E.C., 2003, Depositional environment and history of the Cambrian Tonto Group, Grand Canyon, Arizona [MS thesis]: Flagstaff, Northern Arizona University, 349 p.Google Scholar
Rose, E.C., 2006, Nonmarine aspects of the Tonto Group of the Grand Canyon, USA, and broader implications: Palaeoworld, v. 15, p. 223241.Google Scholar
Rose, E.C., 2011, Modification of the nomenclature and a revised depositional model for the Cambrian Tonto Group of the Grand Canyon, Arizona, in Hollingsworth, J.S., Sundberg, F.A., and Foster, J.R., eds., Cambrian Stratigraphy and Paleontology of Northern Arizona and Southern Nevada, The 16th Field Conference of the Cambrian Stage Subdivision Working Group, International Subcommission on Cambrian Stratigraphy Flagstaff, Arizona, and Southern Nevada, United States: Museum of Northern Arizona Bulletin, no. 67, p. 7798.Google Scholar
Rowland, S.M., 2011, Stop 4A The Cambrian section at Sheep Mountain, Nevada, in Hollingsworth, J.S., Sundberg, F.A., and Foster, J.R., eds., Cambrian Stratigraphy and Paleontology of Northern Arizona and Southern Nevada, The 16th Field Conference of the Cambrian Stage Subdivision Working Group, International Subcommission on Cambrian Stratigraphy Flagstaff, Arizona, And Southern Nevada, United States: Museum of Northern Arizona Bulletin, no. 67, p. 191195.Google Scholar
Rowland, S.M., Korolev, S., Hagadorn, J.H., and Ghosh, K., 2023, Frenchman Mountain Dolostone: a new formation of the Cambrian Tonto Group, Grand Canyon and Basin and Range, USA: Geosphere, v. 19, p. 719747.Google Scholar
Rusconi, C., 1945, Trilobites Silúricos de Mendoza: Anales de la Sociedad Científica Argentina, v. 139, p. 216219.Google Scholar
Schenk, E.T., and Wheeler, H.E., 1942, Cambrian sequence in western Grand Canyon, Arizona: The Journal of Geology, v. 50, p. 882899.10.1086/625089CrossRefGoogle Scholar
Schwimmer, D.R., 1973, The Middle Cambrian Biostratigraphy of Montana and Wyoming [Ph.D. dissertation]: Stony Brook, State University of New York, 452 p.Google Scholar
Schwimmer, D.R., 1975, Quantitative taxonomy and biostratigraphy of middle Cambrian trilobites from Montana and Wyoming: Mathematical Geology, v. 7, p. 149166.10.1007/BF02080490CrossRefGoogle Scholar
Schwimmer, D.R., 1989, Taxonomy and biostratigraphic significance of some middle Cambrian trilobites from the Conasauga Formation in western Georgia: Journal of Paleontology, v. 63, p. 484494.10.1017/S0022336000019703CrossRefGoogle Scholar
Shimer, H.W., and Shrock, R.R., 1944, Index Fossils of North America: Cambridge, Massachusetts, The MIT Press, 837 p.Google Scholar
Spaeth, F., 1938, Über die von Herrn R. Malaise 1934 in Burma gesammelten Cassidinen: Entomologiska Tidskrift, v. 59, p. 228236.Google Scholar
Stoyanow, A., 1958, Sonoraspis and Albertella in the Inyo Mountains, California: Bulletin of the Geological Society of America v. 69, p. 347352.Google Scholar
Stoyanow, A., and Susuki, T., 1955, Discovery of Sonoraspis in southern California: Geological Society of America Bulletin, v. 66, p. 467470.10.1130/0016-7606(1955)66[467:DOSISC]2.0.CO;2CrossRefGoogle Scholar
Sundberg, F.A., 1991, Paleogeography of western Utah and eastern Nevada during the Ehmaniella Biochron (Middle Cambrian), in Cooper, J.D., and Stevens, C., eds., Paleozoic Paleogeography of the Western United States, II: Pacific Section, Society of Economic Paleontologists and Mineralogists, p. 387399.Google Scholar
Sundberg, F.A., 1994, Corynexochida and Ptychopariida (Trilobita, Arthropoda) of the Ehmaniella Biozone (middle Cambrian), Utah and Nevada: Los Angeles County Museum of Natural History, Contributions in Science 446, 137 p.Google Scholar
Sundberg, F.A., 1999, Redescription of Alokistocare subcoronatum (Hall and Whitfield, 1877), the type species of Alokistocare, and the status of Alokistocaridae, Resser, 1938b (Ptychopariida: Trilobita, Middle Cambrian): Journal of Paleontology, v. 73, p. 11261143.Google Scholar
Sundberg, F.A., 2005, The Topazan Stage, a new Laurentian stage (Lincolnian Series—“Middle” Cambrian): Journal of Paleontology, v. 79, p. 6371.10.1666/0022-3360(2005)079<0063:TTSANL>2.0.CO;22.0.CO;2>CrossRefGoogle Scholar
Sundberg, F.A., 2007, Nightmare on Resser Street, in Landing, E., Milukic, D.G., and Kluessendorf, J., eds., Fabulous fossils—300 years of worldwide research on trilobites: New York State Museum Bulletin 507, p. 213224.Google Scholar
Sundberg, F.A., 2011, Delamaran biostratigraphy and lithostratigraphy of southern Nevada, in Hollingsworth, J.S., Sundberg, F.A., and Foster, J.R., eds., Cambrian Stratigraphy and Paleontology of Northern Arizona and Southern Nevada, The 16th Field Conference of the Cambrian Stage Subdivision Working Group, International Subcommission on Cambrian Stratigraphy Flagstaff, Arizona, and Southern Nevada, United States: Museum of Northern Arizona Bulletin, no. 67, p. 174185.Google Scholar
Sundberg, F.A., 2018, Trilobite biostratigraphy of the Cambrian 5 and Drumian stages, Series 3 (Laurentian Delamaran, Topazan, and Marjuman stages, Lincolnian Series) of the lower Emigrant Formation at Clayton Ridge, Esmeralda County, Nevada: Journal of Paleontology, Memoir 76, 44 p.Google Scholar
Sundberg, F.A., 2020, Trilobite fauna (Wuliuan Stage, Miaolingian Series, Cambrian) of the lower Lakeview Limestone, Pend Oreille Lake, Idaho: Journal of Paleontology, v. 94, S79, 49 p.10.1017/jpa.2020.38CrossRefGoogle Scholar
Sundberg, F.A., and McCollum, L.B., 2000, Ptychopariid trilobites of the lower–middle Cambrian boundary interval, Pioche Shale, southeastern Nevada: Journal of Paleontology, v. 74, p. 604630.10.1666/0022-3360(2000)074<0604:PTOTLM>2.0.CO;22.0.CO;2>CrossRefGoogle Scholar
Sundberg, F.A., and McCollum, L.B., 2002, Kochiella Poulsen, 1927, and Hadrocephalites new genus (Trilobita: Ptychopariida) from the lower middle Cambrian of North America: Journal of Paleontology, v. 76, p. 7694.Google Scholar
Sundberg, F.A., and McCollum, L.B., 2003, Trilobites of the lower middle Cambrian Poliella denticulata Biozone (new) of southeastern Nevada: Journal of Paleontology, v. 77, p. 331359.10.1666/0022-3360(2003)077<0331:TOTLMC>2.0.CO;22.0.CO;2>CrossRefGoogle Scholar
Sundberg, F.A., and Webster, M., 2021, Corynexochine trilobites of the Harkless Formation and Mule Spring Limestone (Cambrian Series 2, Stage 4), Clayton Ridge, Nevada: Journal of Paleontology, v. 95, p. 12411258.10.1017/jpa.2021.41CrossRefGoogle Scholar
Sundberg, F.A., and Webster, M., 2022, “Ptychoparioid” trilobites of the Harkless Formation and Mule Spring Limestone (Cambrian Series 2, Stage 4), Clayton Ridge, Nevada: Journal of Paleontology, v. 96, p. 886920.10.1017/jpa.2021.124CrossRefGoogle Scholar
Sundberg, F.A., Karlstrom, K.E., Geyer, G., Foster, J.R., Hagadorn, J.W., Mohr, M.T., Schmitz, M.D., Dehler, C.M., and Crossey, L.J., 2020, Asynchronous trilobite extinctions at the early to middle Cambrian transition: Geology, v. 48, p. 441445.Google Scholar
Sundberg, F.A., Webster, M., and Geyer, G., 2022, Biostratigraphical significance of a new trilobite fauna from the Harkless Formation (upper Stage 4, Series 2, Cambrian), Nevada, USA: Lethaia, v. 55, no. 3, https://doi.org/10.18261/let.55.3.8.CrossRefGoogle Scholar
Swinnerton, H.H., 1915, Suggestions for a revised classification of trilobites: Geological Magazine (n. ser.), v. 6, p. 487496, 538–545.10.1017/S0016756800203634CrossRefGoogle Scholar
Tortello, M.F., 2022, Cambrian trilobites from the Glossopleura walcotti Zone (Miaolingian Series, Wuliuan Stage) of Mendoza, western Argentina: Journal of Paleontology, v. 96, p. 611630.Google Scholar
Tullberg, S.A., 1880, Om Agnostus-artema i de kambriska aflagringarma vid Andrarum: Sveriges Geologiska Undersökning, Avhandlingar och uppsatser, Series C, v. 42, p. 138.Google Scholar
Ulrich, E.O., and Bassler, R.S., 1931, Cambrian bivalved Crustacea of the order Conchostraca: United States National Museum Proceedings, v. 78, art. 4, p. 1130.10.5479/si.00963801.78-2847.1CrossRefGoogle Scholar
Walcott, C.D., 1884, Paleontology of the Eureka District: United States Geological Survey Monographs v. 8, p. 1298.Google Scholar
Walcott, C.D., 1885, Paleozoic notes: new genus of Cambrian trilobites, Mesonacis: American Journal of Science, ser. 3, v. 29, p. 328330.10.2475/ajs.s3-29.172.328CrossRefGoogle Scholar
Walcott, C.D., 1886, Second contribution to the studies on the Cambrian fauna of North America: Bulletin of the United States Geological Survey, v. 30, p. 1369.Google Scholar
Walcott, C.D., 1888, Cambrian fossils from Mount Stephens, Northwest Territory of Canada: American Journal of Science, ser. 3, v. 36, p. 163166.Google Scholar
Walcott, C.D., 1889, Description of new genera and species of fossils from the middle Cambrian: United States National Museum Proceedings, v. 11, p. 441446.10.5479/si.00963801.11-738.441CrossRefGoogle Scholar
Walcott, C.D., 1890, The fauna of the lower Cambrian or Olenellus Zone: 10th Annual Report of Director of the United State Geological Survey for 1888–1889, pt.1, p. 509774.Google Scholar
Walcott, C.D., 1898, Cambrian Brachiopoda: Obolus and Lingulella, with description of new species: Proceedings of the United States National Museum, v. 21, p. 385420.10.5479/si.00963801.21-1152.385CrossRefGoogle Scholar
Walcott, C.D., 1899, Cambrian fossils of the Yellowstone National Park: United States Geological Survey Monographs, 32, p. 440478.Google Scholar
Walcott, C.D., 1901, Cambrian Brachiopoda: Obolella, subgenus Glyptias; Bicia; Obolus, subgenus Westonia; with descriptions of new species: Proceedings of the United States National Museum, v. 23, p. 669695.10.5479/si.00963801.1229.669CrossRefGoogle Scholar
Walcott, C.D., 1903, Cambrian Brachiopoda: Acrotreta; Linnarssonella; Obolus; with descriptions of new species: Proceedings of the United States National Museum, v. 25, p. 577612.10.5479/si.00963801.1299.577CrossRefGoogle Scholar
Walcott, C.D., 1905, Cambrian Brachiopoda with description of new genera and species: United States National Museum, Proceedings, v. 28, p. 227337.Google Scholar
Walcott, C.D., 1908, Cambrian geology and paleontology, III. Cambrian Brachiopoda: descriptions of new genera and species: Smithsonian Miscellaneous Collections, v. 53, p. 52137.Google Scholar
Walcott, C.D., 1910, Olenellus and other genera of the Mesonacidae: Smithsonian Miscellaneous Collections, v. 53, no. 6, p. 231422.Google Scholar
Walcott, C.D., 1912, Cambrian geology and paleontology, 2, middle Cambrian Branchiopoda, Malacostraca, Trilobita and Merostromata: Smithsonian Miscellaneous Collections, v. 57, no. 6, p. 145228.Google Scholar
Walcott, C.D., 1913, Cambrian geology and paleontology, No. 11. New lower Cambrian subfauna: Smithsonian Miscellaneous Collections, v. 57, no. 11, p. 309326.Google Scholar
Walcott, C.D., 1916a, Cambrian geology and paleontology III, no. 3, Cambrian trilobites: Smithsonian Miscellaneous Collections, v. 64, no. 3, p. 157258.Google Scholar
Walcott, C.D., 1916b, Cambrian geology and paleontology II, no. 5, Cambrian trilobites: Smithsonian Miscellaneous Collections, v. 64, no. 5, p. 303456.Google Scholar
Walcott, C.D., 1917, Cambrian Geology and Paleontology IV, No. 3—fauna of the Mount Whyte Formation: Smithsonian Miscellaneous Collections, v. 67, no. 3, p. 61114.Google Scholar
Walcott, C.D., 1924, Cambrian and lower Ozarkian trilobites. Cambrian geology and paleontology, pt. 5: Smithsonian Miscellaneous Collections, v. 75, n. 2, p. 5360.Google Scholar
Walcott, C.D., 1925, Cambrian geology and paleontology V, no. 3, Cambrian and Ozarkian trilobites: Smithsonian Miscellaneous Collections, v. 75, no. 3, p. 61146.Google Scholar
Wanless, H.R. Jr. 1973, Cambrian of the Grand Canyon–a reevaluation of the depositional environments [Ph.D. dissertation]: Baltimore, Johns Hopkins University, 114 p.Google Scholar
Wanless, H.R., 1975, Carbonate tidal flats of the Grand Canyon Cambrian, in Ginsburg, R.N., ed., Tidal Deposits: Berlin, Heidelberg, Springer,.Google Scholar
Webster, M., 2007, Ontogeny and evolution of the early Cambrian trilobite genus Nephrolenellus (Olenelloidea): Journal of Paleontology, v. 81, p. 11681193.Google Scholar
Webster, M., 2011a, Trilobite biostratigraphy and sequence stratigraphy of the upper Dyeran (traditional Laurentian “Lower Cambrian”) in the southern Great Basin, U.S.A., in Hollingsworth, J.S., Sundberg, F.A., and Foster, J.R., eds., Cambrian Stratigraphy and Paleontology of Northern Arizona and Southern Nevada, The 16th Field Conference of the Cambrian Stage Subdivision Working Group, International Subcommission on Cambrian Stratigraphy Flagstaff, Arizona, and Southern Nevada, United States: Museum of Northern Arizona Bulletin, no. 67, p. 121154.Google Scholar
Webster, M., 2011b, Litho- and biostratigraphy of the Dyeran–Delamaran boundary interval at Frenchman Mountain, Nevada, in Hollingsworth, J.S., Sundberg, F.A., and Foster, J.R., eds., Cambrian Stratigraphy and Paleontology of Northern Arizona and Southern Nevada, The 16th Field Conference of the Cambrian Stage Subdivision Working Group, International Subcommission on Cambrian Stratigraphy Flagstaff, Arizona, and Southern Nevada, United States: Museum of Northern Arizona Bulletin, no. 67, p. 195203.Google Scholar
Webster, M., 2011c, Litho- and biostratigraphy of the Dyeran-Delamaran boundary interval in the Pioche-Caliente region, Nevada, in Hollingsworth, J.S., Sundberg, F.A., and Foster, J.R., eds., Cambrian Stratigraphy and Paleontology of Northern Arizona and Southern Nevada, The 16th Field Conference of the Cambrian Stage Subdivision Working Group, International Subcommission on Cambrian Stratigraphy Flagstaff, Arizona, and Southern Nevada, United States: Museum of Northern Arizona Bulletin, no. 67, p. 203215.Google Scholar
Webster, M., 2015, Ontogeny and intraspecific variation of the early Cambrian trilobite Olenellus gilberti, with implications for olenelline phylogeny and macroevolutionary trends in phenotypic canalization: Journal of Systematic Palaeontology, v. 13, no. 1, p. 174.10.1080/14772019.2013.852903CrossRefGoogle Scholar
Webster, M., and Caron, J.-B., 2024, Trilobites of the Cranbrook Lagerstätte (Eager Formation, Cambrian Stage 4), British Columbia: Journal of Paleontology, v. 98, p. 460503.10.1017/jpa.2023.89CrossRefGoogle Scholar
Webster, M., and Landing, E., 2016, Geological context, biostratigraphy and systematic revision of late early Cambrian olenelloid trilobites from the Parker and Monkton formations, northwestern Vermont, U.S.A: Australasian Palaeontological Memoirs 49, p. 193240.Google Scholar
White, C.A., 1874, Preliminary report upon invertebrate fossils collected by the expeditions of 1871, 1872, and 1873, with descriptions of new species: Report of the United States Army Geographical and Geological Exploratory Surveys West of the 100th Meridian, Washington, Government Printing Office, 27 p.Google Scholar
White, W.W. III, 1973, Paleontology and depositional environments of the Cambrian Wheeler Formation, Drum Mountains, west-central Utah [M.S. thesis]: Salt Lake City, University of Utah, 135 p.Google Scholar
Whittington, H.B., 1989, Olenelloid trilobites: type species, functional morphology and higher classification: Philosophical Transactions of the Royal Society of London, B 324, p. 111147.Google Scholar
Whittington, H.B., 1992, Trilobites. Fossils Illustrated, Volume 2: Woodbridge, England, Boydell Press, 145 p.Google Scholar
Whittington, H.B., Chatterton, B.D.E., Speyer, S.E., Fortey, R.A., Owens, R.M., et al., 1997, Treatise on Invertebrate Paleontology. Part O. Arthropoda 1. Trilobita, Revised. Volume 1: Introduction, Order Agnostida, Order Redlichiida: Boulder, Colorado and Lawrence, Kansas, Geological Society of America and University of Kansas, 530 p.Google Scholar
Zhang, W.T., and Jell, P.A., 1987, Cambrian Trilobites of North China. Chinese Cambrian Trilobites Housed in the Smithsonian Institution: Beijing, Science Press, 322 p.Google Scholar
Zhao, Y.L., Yuan, J.L., Babcock, L.E., Guo, Q.J., Peng, J., et al., 2019, Global Standard Stratotype-Section and Point (GSSP) for the conterminous base of the Miaolingian Series and Wuliuan Stage (Cambrian) at Balang, Jianhe, Guizhou, China: Episodes, v. 42, p. 165184.Google Scholar