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4 - The New Taphonomy and How It Can Boost the Role of the Asian Pleistocene Archaeological Record in Human Evolution

Published online by Cambridge University Press:  13 December 2025

Manuel Dominguez-Rodrigo
Affiliation:
Rice University, Houston
Enrique Baquedano
Affiliation:
University of Madrid
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Summary

Debates on human behavioral evolution have largely focused on African and European records, while Asia’s contribution remains underrepresented. Despite the significance of the Asian Pleistocene fossil record, its behavioral insights have been hindered by limited taphonomic research, restricted dissemination, and shifting academic trends. Many key Chinese archaeofaunal sites, particularly in karstic contexts, contain complex palimpsests that challenge traditional taphonomic methods prone to equifinality.

Advancements in artificial intelligence and computational archaeology now offer new ways to address these challenges. Machine learning classifiers, computer vision through convolutional neural networks, and 3D deep learning architectures enable precise discrimination of bone surface modifications. These techniques refine carnivore agency identification down to the taxon level and provide mathematical certainty in agency attribution, aiding in disentangling complex palimpsests.

This study highlights key Chinese archaeofaunal records, particularly Zhoukoudian, and proposes methodological approaches to improve their resolution. By integrating these cutting-edge techniques, the Asian Pleistocene record can take a more central role in discussions on early human behavioral variability. This research aims to establish a model for applying the “new taphonomy” globally, enhancing our understanding of hominin activities and their ecological contexts.

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References

Abellán, N, Jiménez-García, B, Aznarte, J, et al. 2021. Deep learning classification of tooth scores made by different carnivores: Achieving high accuracy when comparing African carnivore taxa and testing the hominin shift in the balance of power. Archaeol Anthropol Sci 13: 31.CrossRefGoogle Scholar
Ao, H, Dekkers, MJ, Wei, Q, et al. 2013. New evidence for early presence of hominids in north China. Sci Rep 3: 2403.CrossRefGoogle ScholarPubMed
Arriaza, MC and Domínguez-Rodrigo, M. 2016. When felids and hominins ruled at Olduvai Gorge: A machine learning analysis of the skeletal profiles of the non-anthropogenic Bed I sites. Quat Sci Rev 139: 4352.CrossRefGoogle Scholar
Arriaza, MC, Organista, E, Yravedra, J, et al. 2019. Striped hyenas as bone modifiers in dual human-to-carnivore experimental models. Archaeol Anthropol Sci 11: 3187–99.CrossRefGoogle Scholar
Bartram, EL Jr, and Marean, CW. 1999. Explaining the “Klasies pattern”: Kua ethnoarchaeology, the Die Kelders Middle Stone Age archaeofauna, long bone fragmentation and carnivore ravaging. J Archaeol Sci 26: 929.CrossRefGoogle Scholar
Bartram, LE, Kroll, EM, and Bunn, HT. 1991. Variability in camp structure and bone food refuse patterning at Kua San Hunter-Gatherer camps. In: The Interpretation of Archaeological Spatial Patterning. Springer US.Google Scholar
Bello, SM, Parfitt, SA, and Stringer, C. 2009. Quantitative micromorphological analyses of cut marks produced by ancient and modern handaxes. J Archaeol Sci 36: 1869–80.CrossRefGoogle Scholar
Bello, SM and Soligo, C. 2008. A new method for the quantitative analysis of cutmark micromorphology. J Archaeol Sci 35: 1542.CrossRefGoogle Scholar
Bello, SM, Verveniotou, E, Cornish, L, and Parfitt, SA. 2011. 3-Dimensional microscope analysis of bone and tooth surface modifications: Comparisons of fossil specimens and replicas. Scanning 33: 316–24.CrossRefGoogle Scholar
Binford, LR. 1981. Bones: Ancient men and modern myths. New York: Aca-demicBinfordBones: Ancient Men and Modern Myths 1981.Google Scholar
Binford, LR. 1984. Faunal Remains from Klasies River Mouth. Academic Press.Google Scholar
Binford, LR, Ho, CK, 1985. Taphonomy at a distance: Zhoukoudian, “the cave home of Beijing man”? [and comments and reply]. Curr Anthropol 26: 413–42.CrossRefGoogle Scholar
Binford, LR, Stone, NM, 1986. Zhoukoudian: A closer look [and comments and reply]. Curr Anthropol 27: 453–75.CrossRefGoogle Scholar
Black, D. 2009. Evidences of the use of fire by Sinanthropus*. Bull Geol Soc China 11(2): 107108.CrossRefGoogle Scholar
Blumenschine, RJ. 1986. Carcass consumption sequences and the archaeological distinction of scavenging and hunting. J Hum Evolut 15: 639–59.Google Scholar
Blumenschine, RJ. 1991. Hominid carnivory and foraging strategies, and the socio-economic function of early archaeological sites. Philos Trans R Soc Lond B Biol Sci 334: 211–9; discussion 219–21.Google ScholarPubMed
Boaz, NT and Ciochon, RL. 2004a. The adaptive behavior of the not-quite-human. Dragon Bone Hill 90–107.CrossRefGoogle Scholar
Boaz, NT and Ciochon, RL. 2004b. Dragon Bone Hill: An Ice-Age Saga of Homo erectus. Oxford University Press.CrossRefGoogle Scholar
Boaz, NT, Ciochon, RL, Xu, Q, and Liu, J. 2000. Large mammalian carnivores as a taphonomic factor in the bone accumulation at Zhoukoudian. Acta Anthropologica Sinica 19: 224–34.Google Scholar
Boaz, NT, Ciochon, RL, Xu, Q, and Liu, J. 2004. Mapping and taphonomic analysis of the Homo erectus loci at Locality 1 Zhoukoudian, China. J Hum Evol 46: 519–49.CrossRefGoogle ScholarPubMed
Brain, CK. 1981. The Hunters or the Hunted?: An Introduction to African Cave Taphonomy. University of Chicago Press.Google Scholar
Braun, DR, Pante, M, and Archer, W. 2016. Cut marks on bone surfaces: Influences on variation in the form of traces of ancient behaviour. Interface Focus 6: 20160006.CrossRefGoogle Scholar
Brophy, JK. 2011. Reconstructing the habitat mosaic associated with Australopithecus robustus: Evidence from quantitative morphological analysis of bovid teeth. Ph.D. dissertation, University of Louisiana.Google Scholar
Bunn, HT. 1986. Patterns of skeletal representation and hominid subsistence activities at Olduvai Gorge, Tanzania, and Koobi Fora, Kenya. J Hum Evol 15: 673–90.CrossRefGoogle Scholar
Bunn, HT, Bartram, LE, and Kroll, EM. 1988. Variability in bone assemblage formation from Hadza hunting, scavenging, and carcass processing. J Anthropol Archaeol 7: 412–57.CrossRefGoogle Scholar
Bunn, HT, Kroll, EM, Ambrose, SH, et al. 1986. Systematic butchery by Plio/Pleistocene hominids at Olduvai Gorge, Tanzania [and comments and reply]. Curr Anthropol 27: 431–52.CrossRefGoogle Scholar
Byeon, W, Domínguez-Rodrigo, M, Arampatzis, G, et al. 2019. Automated identification and deep classification of cut marks on bones and its paleoanthropological implications. J Comput Sci 32: 3643.CrossRefGoogle Scholar
Cifuentes-Alcobendas, G and Domínguez-Rodrigo, M. 2019. Deep learning and taphonomy: High accuracy in the classification of cut marks made on fleshed and defleshed bones using convolutional neural networks. Sci Rep 9: 18933.CrossRefGoogle ScholarPubMed
Cifuentes-Alcobendas, G and Domínguez-Rodrigo, M. 2021. More than meets the eye: Use of computer vision algorithms to identify stone tool material through the analysis of cut mark micro-morphology. Archaeol Anthropol Sci 13: 167.CrossRefGoogle Scholar
Cleghorn, N, Marean, CW, and Pickering, TR. 2007. The destruction of skeletal elements by carnivores: The growth of a general model for skeletal element destruction and survival in zooarchaeological assemblages. In: Breathing Life into Fossils: Taphonomic Studies in Honor of CK (Bob) Brain Stone Age. Institute Press, pp. 3766.Google Scholar
Courtenay, LA and González-Aguilera, D. 2020. Geometric morphometric data augmentation using generative computational learning algorithms. Appl Sci 10: 9133.CrossRefGoogle Scholar
Courtenay, LA, Huguet, R, González-Aguilera, D, and Yravedra, J. 2019a. A hybrid geometric morphometric deep learning approach for cut and trampling mark classification. NATO Adv Sci Inst Ser E Appl Sci 10: 150.Google Scholar
Courtenay, LA, Maté-González, , Aramendi, J, et al. 2018. Testing accuracy in 2D and 3D geometric morphometric methods for cut mark identification and classification. PeerJ 6: e5133.CrossRefGoogle ScholarPubMed
Courtenay, LA, Yravedra, J, and Huguet, R. 2019b. Combining machine learning algorithms and geometric morphometrics: A study of carnivore tooth marks. Palaeogeogr Palaeoclimatol Palaeoecol 522: 2839.CrossRefGoogle Scholar
Courtenay, LA, Yravedra, J, Huguet, R, et al. 2019c. New taphonomic advances in 3D digital microscopy: A morphological characterisation of trampling marks. Quat Int 517: 5566.CrossRefGoogle Scholar
Cruz-Uribe, K. 1991. Distinguishing hyena from hominid bone accumulations. J Field Archaeol 18: 467–86.CrossRefGoogle Scholar
David Mech, L. 2003. The Wolves of Denali. University of Minnesota Press.CrossRefGoogle Scholar
David Mech, L and Boitani, L. 2010. Wolves: Behavior, Ecology, and Conservation. University of Chicago Press.Google Scholar
David Mech, L, Smith, DW, and MacNulty, DR. 2015. Wolves on the Hunt: The Behavior of Wolves Hunting Wild Prey. University of Chicago Press.CrossRefGoogle Scholar
Domínguez-Rodrigo, M. 2011. Critical review of the MNI (minimum number of individuals) as a zooarchaeological unit of quantification. Archaeol Anthropol Sci 4: 4759.CrossRefGoogle Scholar
Domínguez-Rodrigo, M. 2015. Taphonomy in early African archaeological sites: Questioning some bone surface modification models for inferring fossil hominin and carnivore feeding interactions. J Afr Earth Sci 108: 42–6.CrossRefGoogle Scholar
Domínguez-Rodrigo, M. 2019. Successful classification of experimental bone surface modifications (BSM) through machine learning algorithms: A solution to the controversial use of BSM in paleoanthropology? Archaeol Anthropol Sci 11: 2711–25.CrossRefGoogle Scholar
Domínguez-Rodrigo, M and Baquedano, E. 2018. Distinguishing butchery cut marks from crocodile bite marks through machine learning methods. Sci Rep 8: 5786.CrossRefGoogle ScholarPubMed
Domínguez-Rodrigo, M, Baquedano, E, Organista, E, et al. 2021a. Early Pleistocene faunivorous hominins were not kleptoparasitic, and this impacted the evolution of human anatomy and socio-ecology. Sci Rep 11: 16135.CrossRefGoogle Scholar
Domínguez-Rodrigo, M, Barba, R, and Egeland, CP. 2007. Deconstructing Olduvai: A taphonomic study of the Bed I sites. Springer Science & Business Media.CrossRefGoogle Scholar
Domínguez-Rodrigo, M, Bunn, HT, and Yravedra, J. 2014. A critical re-evaluation of bone surface modification models for inferring fossil hominin and carnivore interactions through a multivariate approach: Application to the FLK Zinj archaeofaunal assemblage (Olduvai Gorge, Tanzania). Quat Int 322–323: 3243.CrossRefGoogle Scholar
Domínguez-Rodrigo, M, Cifuentes-Alcobendas, G, Jiménez-García, B, et al. 2020. Artificial intelligence provides greater accuracy in the classification of modern and ancient bone surface modifications. Sci Rep 10: 18862.CrossRefGoogle ScholarPubMed
Domínguez-Rodrigo, M, Cobo-Sánchez, L, Yravedra, J, et al. 2018. Fluvial spatial taphonomy: A new method for the study of post-depositional processes. Archaeol Anthropol Sci 10: 1769–89.CrossRefGoogle Scholar
Domínguez Rodrigo, M, Fernández López, SR, and Alcalá, L. 2011. How can taphonomy be defined in the XXI century? J Paleontol 9: 113.Google Scholar
Domínguez-Rodrigo, M, Gidna, A, Baquedano, E, et al. 2021b. A 3D taphonomic model of long bone modification by lions in medium-sized ungulate carcasses. Sci Rep 11: 4944.CrossRefGoogle ScholarPubMed
Domínguez-Rodrigo, M, Juana, S de, Galán, AB, and Rodríguez, M. 2009. A new protocol to differentiate trampling marks from butchery cut marks. J Archaeol Sci 36: 2643–54.CrossRefGoogle Scholar
Domínguez-Rodrigo, M, Saladié, P, Cáceres, I, et al. 2017. Use and abuse of cut mark analyses: The Rorschach effect. J Archaeol Sci 86: 1423.CrossRefGoogle Scholar
Domínguez-Rodrigo, M, Vegara-Riquelme, M, Palomeque-González, J, et al. 2025. Testing the reliability of geometric morphometric and computer vision methods to identify carnivore agency using bi-dimensional information. Quat Sci Adv 17: 100268.Google Scholar
Dongsheng, L, Senshui, Z, Xinzhi, W, et al. 1998. Comments on a Science paper regarding the evidence of fire use at Zhoukoudian Locality 1. Acta Anthropol Sin 17(4): 317–29.Google Scholar
Egeland, AG, Egeland, CP, and Bunn, HT. 2008. Taphonomic analysis of a modern spotted hyena (Crocuta crocuta) den from Nairobi, Kenya. J Taphon 6: 275–99.Google Scholar
Faith, JT. 2007. Sources of variation in carnivore tooth-mark frequencies in a modern spotted hyena (Crocuta crocuta) den assemblage, Amboseli Park, Kenya. J Archaeol Sci 34: 1601–9.CrossRefGoogle Scholar
Ferraro, JV, Plummer, TW, Pobiner, BL, et al. 2013. Earliest archaeological evidence of persistent hominin carnivory. PLoS One 8: e62174.CrossRefGoogle ScholarPubMed
Gaboardi, M, Deng, T, and Wang, Y. 2005. Middle Pleistocene climate and habitat change at Zhoukoudian, China, from the carbon and oxygen isotopic record from herbivore tooth enamel. Quater Res 63 (3): 329–38.CrossRefGoogle Scholar
Gao, X, Cote, P, Blais, J-P, et al. 2016. Geophysical investigations identify hidden deposits with great potential for discovering Peking Man fossils at Zhoukoudian, China. Quat Int 400: 30–5.CrossRefGoogle Scholar
Gao, X, Zhang, S, Zhang, Y, and Chen, F. 2017. Evidence of hominin use and maintenance of fire at Zhoukoudian. Curr Anthropol 58: S267–77.CrossRefGoogle Scholar
Goldberg, P, Weiner, S, Bar-Yosef, O, et al. 2001. Site formation processes at Zhoukoudian, China. J Hum Evol 41: 483530.CrossRefGoogle ScholarPubMed
Grayson, DK and Frey, CJ. 2004. Measuring skeletal part representation in archaeological faunas. J Taphon 2: 2742.Google Scholar
Gümrükçü, M and Pante, MC. 2018. Assessing the effects of fluvial abrasion on bone surface modifications using high-resolution 3-D scanning. J Archaeol Sci: Rep 21: 208–21.Google Scholar
Guo, SL., Liu, SH, Sun, SF, et al. 1980. Age determination of Peking Man by fission track dating. Chin Sci Bull 25:535–36.Google Scholar
Haynes, G., 1980. Prey bones and predators: Potential ecologic information from analysis of bone sites. Ossa 7: 7597.Google Scholar
Jiménez-García, B, Abellán, N, Baquedano, E, et al. 2020a. Corrigendum to “Deep learning improves taphonomic resolution: High accuracy in differentiating tooth marks made by lions and jaguars”. J R Soc Interface 17: 20200782.CrossRefGoogle Scholar
Jiménez-García, B, Aznarte, J, Abellán, N, et al. 2020b. Deep learning improves taphonomic resolution: High accuracy in differentiating tooth marks made by lions and jaguars. J R Soc Interface 17: 20200446.CrossRefGoogle Scholar
Juana, S de and Domínguez-Rodrigo, M. 2011. Testing analogical taphonomic signatures in bone breaking: A comparison between hammerstone-broken equid and bovid bones. Archaeometry 53: 9961011.CrossRefGoogle Scholar
Klein, RG and Cruz-Uribe, K. 1996. Exploitation of large bovids and seals at Middle and Later Stone Age sites in south Africa. J Hum Evol 31: 315–34.CrossRefGoogle Scholar
Kuhn, BF, Berger, LR, and Skinner, JD. 2010. Examining criteria for identifying and differentiating fossil faunal assemblages accumulated by hyenas and hominins using extant hyenid accumulations. Int J Osteoarchaeol 20: 1535.CrossRefGoogle Scholar
Lee Lyman, R. 1994. Vertebrate Taphonomy. Cambridge University Press.CrossRefGoogle Scholar
Li, F. 2016. An experimental study of bipolar reduction at Zhoukoudian locality 1, north China. Quat Int 400: 23–9.CrossRefGoogle Scholar
Li, F, Chen, F, Zhang, S, and Gao, X. 2018. Linking spatial grids of the old and new excavations at Zhoukoudian Locality 1, China. J Hum Evol 121 (August): 166–69.CrossRefGoogle ScholarPubMed
Lin, S., 1985. Large fossil mammals of Locality 1 of Zhoukoudian and the hunting behavior of Peking Man. In: Wu, R., Ren, ME, Zhang, X, et al (Eds.), Multi-disciplinary Study of the Peking Man Site at Zhoukoudian. Science Press, pp. 95101(in Chinese).Google Scholar
Lyman, RL and Others. 2010. What taphonomy is, what it isn’t, and why taphonomists should care about the difference. J Taphon 8: 116.Google Scholar
Marean, CW, Domínguez-Rodrigo, M, and Pickering, TR. 2004. Skeletal element equifinality in Zooarchaeology begins with method: The evolution and status of the “shaft critique.” J Taphon 2: 6998.Google Scholar
Marean, CW and Kim, SY. 1998. Mousterian large‐mammal remains from Kobeh cave behavioral implications for Neanderthals and early modern humans. Curr Anthropol 39: S79114.CrossRefGoogle Scholar
Merritt, SR. 2012. Factors affecting Early Stone Age cut mark cross-sectional size: Implications from actualistic butchery trials. J Archaeol Sci 39: 2984–94.CrossRefGoogle Scholar
Milo, RG. 1998. Evidence for hominid predation at Klasies River Mouth, South Africa, and its implications for the behaviour of early modern humans. J Archaeol Sci 25: 99133.CrossRefGoogle Scholar
Moclán, A and Domínguez-Rodrigo, M. 2018. An experimental study of the patterned nature of anthropogenic bone breakage and its impact on bone surface modification frequencies. J Archaeol Sci 96: 113.CrossRefGoogle Scholar
Moclán, A, Domínguez-Rodrigo, M, and Yravedra, J. 2019. Classifying agency in bone breakage: An experimental analysis of fracture planes to differentiate between hominin and carnivore dynamic and static loading using machine learning (ML) algorithms. Archaeol Anthropol Sci 11: 4663–80.CrossRefGoogle Scholar
Moclán, A, Huguet, R, Márquez, B, et al. 2020. Identifying the bone-breaker at the Navalmaíllo Rock Shelter (Pinilla del Valle, Madrid) using machine learning algorithms. Archaeol Anthropol Sci 12: 46.CrossRefGoogle Scholar
Moclán, A, Huguet, R, Márquez, B, et al. 2021. A Neanderthal hunting camp in the central system of the Iberian Peninsula: A zooarchaeological and taphonomic analysis of the Navalmaíllo Rock Shelter (Pinilla del Valle, Spain). Quat Sci Rev 269: 107142.CrossRefGoogle Scholar
Monahan, CM. 1998. The Hadza carcass transport debate revisited and its archaeological implications. J Archaeol Sci 25: 405–24.CrossRefGoogle Scholar
Nunn, CL. 2011. The Comparative Approach in Evolutionary Anthropology and Biology. University of Chicago Press.Google Scholar
Oliver, JS, Plummer, TW, Hertel, F, and Bishop, LC. 2019. Bovid mortality patterns from Kanjera South, Homa Peninsula, Kenya and FLK-Zinj, Olduvai Gorge, Tanzania: Evidence for habitat mediated variability in Oldowan hominin hunting and scavenging behavior. J Hum Evol 131: 6175.CrossRefGoogle ScholarPubMed
Pante, MC, Muttart, MV, Keevil, TL, et al. 2017. A new high-resolution 3-D quantitative method for identifying bone surface modifications with implications for the Early Stone Age archaeological record. J Hum Evol 102: 111.CrossRefGoogle ScholarPubMed
Parkinson, JA. 2018. Revisiting the hunting-versus-scavenging debate at FLK Zinj: A GIS spatial analysis of bone surface modifications produced by hominins and carnivores in the FLK 22 assemblage, Olduvai Gorge, Tanzania. Palaeogeogr Palaeoclimatol Palaeoecol 511: 2951.CrossRefGoogle Scholar
Pei, W-C. 1933. Le rôle des animaux et des causes naturelles dans la cassure des os. Geological Survey of China 1938 Palæontologia Sinica New ser. D, no. 7, whole ser. no. 118.Google Scholar
Peterhans, JK. 1990. The Roles of Porcupines, Leopards and Hyenas in Ungulate Carcass Dispersal: Implications for Paleoanthropology. The University of Chicago.Google Scholar
Pickering, TR. 2002. Reconsideration of criteria for differentiating faunal assemblages accumulated by hyenas and hominids. Int J Osteoarchaeol 12: 127–41.CrossRefGoogle Scholar
Pickering, TR, Marean, CW, and Domínguez-Rodrigo, M. 2003. Importance of limb bone shaft fragments in zooarchaeology: A response to “On in situ attrition and vertebrate body part profiles” (2002), by M.C. Stiner. J Archaeol Sci 30: 1469–82.CrossRefGoogle Scholar
Pizarro-Monzo, M and Domínguez-Rodrigo, M. 2020. Dynamic modification of cut marks by trampling: Temporal assessment through the use of mixed-effect regressions and deep learning methods. Archaeol Anthropol Sci 12: 4.CrossRefGoogle Scholar
Pizarro-Monzo, M, Prendergast, ME, Gidna, AO, et al. 2021. Do human butchery patterns exist? A study of the interaction of randomness and channelling in the distribution of cut marks on long bones. J R Soc Interface 18: 20200958.CrossRefGoogle ScholarPubMed
Pizarro, M, Organista, E, Cobo-Sánchez, L, Baquedano, E, Domínguez-Rodrigo, M. 2022. Determining the diagenetic paths of archaeofaunal assemblages and their paleoecological implications through the computer vision analysis of bone surface modifications: An application to Oldowan sites from Olduvai Gorge (Tanzania). J Quat Sci 37: 543–57.Google Scholar
Pokines, JT and Kerbis Peterhans, JC. 2007. Spotted hyena (Crocuta crocuta) den use and taphonomy in the Masai Mara National Reserve, Kenya. J Archaeol Sci 34: 1914–31.CrossRefGoogle Scholar
Prendergast, ME and Domínguez-Rodrigo, M. 2008. Taphonomic analyses of a hyena den and a natural-death assemblage near Lake Eyasi (Tanzania). J Taphon 6: 301–36.Google Scholar
Rodrigo, MD and Pickering, TR. 2010. A multivariate approach for discriminating bone accumulations created by spotted hyenas and leopards: Harnessing actualistic data from East and Southern Africa. J Taphon 8: 155–79.Google Scholar
Saladié, P, Rodríguez-Hidalgo, A, Domínguez-Rodrigo, M, et al. 2021. Dragged, lagged, or undisturbed: Reassessing the autochthony of the hominin-bearing assemblages at Gran Dolina (Atapuerca, Spain). Archaeol Anthropol Sci 13: 65.CrossRefGoogle Scholar
Shen, C, Zhang, X, and Gao, X. 2016. Zhoukoudian in transition: Research history, lithic technologies, and transformation of Chinese Palaeolithic archaeology. Quat Int 400: 413.CrossRefGoogle Scholar
Stiner, MC. 1994. Honor among thieves: A zooarchaeological study of Neandertal ecology. Princeton University Press.Google Scholar
Von Bertalanffy, L. 1956. General system theory. Gen Syst 1: 110.Google Scholar
Weiner, S, Xu, Q, Goldberg, P, Liu, J, and Bar-Yosef, O. 1998. Evidence for the Use of Fire at Zhoukoudian, China. Science 281(5374): 251-253. https://science.sciencemag.org/content/281/5374/251.abstract.CrossRefGoogle Scholar
Wu, X. 1999. Investigating the Possible Use of Fire at Zhoukoudian, China. Science 283 (5400): 299.CrossRefGoogle Scholar
Yravedra, J, Lagos, L, and Bárcena, F. 2011. A taphonomic study of wild wolf (Canis lupus) modification of horse bones in Northwestern Spain. J Taphon 9: 3765.Google Scholar
Yravedra, J, Maté-González, , Courtenay, LA, et al. 2019. The use of canid tooth marks on bone for the identification of livestock predation. Sci Rep 9: 16301.CrossRefGoogle Scholar
Zanolli, C, Pan, L, Dumoncel, J, et al. 2018. Inner Tooth Morphology of Homo Erectus from Zhoukoudian. New Evidence from an Old Collection Housed at Uppsala University, Sweden. J Hum Evol 116 (March): 113.CrossRefGoogle ScholarPubMed
Zhang, Y, Guo, Z, Deng, C, et al. 2014. The use of fire at Zhoukoudian: Evidence from magnetic susceptibility and color measurements. Chin Sci Bull 59: 1013–20.Google Scholar
Zhang, SQ., Chen, FY., Zhang, Y., Li, J., Gao, X., 2016. A taphonomic study on the skeletal remains of Cervus (Sika) grayi from layer 3 of the Peking Man site at Zhoukoudian during the 2009–2010 field seasons. Quat Int 400: 3646.CrossRefGoogle Scholar
Zhong, M, Shi, C, Gao, X, et al. 2014. On the possible use of fire by Homo erectus at Zhoukoudian, China. Chin. Sci Bull 59(3):335343.CrossRefGoogle Scholar
Zhou, C, Liu, Z, Wang, Y, and Huang, Q. 2000. Climatic cycles investigated by sediment analysis in Peking Man’s cave, Zhoukoudian, China. J Archaeol Sci 27 (2): 101–10.CrossRefGoogle Scholar
Zhu, Z, Dennell, R, Huang, W, et al. 2018. Hominin occupation of the Chinese Loess Plateau since about 2.1 million years ago. Nature 559: 608–12.CrossRefGoogle ScholarPubMed
Zhu, RX, Potts, R, Pan, YX, et al. 2008. Early evidence of the genus Homo in East Asia. J Hum Evol 55: 1075–85.CrossRefGoogle Scholar
Zhu, RX, Potts, R, Xie, F, et al. 2004. New evidence on the earliest human presence at high northern latitudes in northeast Asia. Nature 431: 559–62.CrossRefGoogle ScholarPubMed

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