In ordinary speech a system of any sort is said to be stable when it cannot be upset easily, but the meaning attached to the word is usually somewhat vague. It is hardly surprising that this should be the case, when it is only within the last thirty years, and principally through the investigations of M. Poincaré, that the conception of stability has, even for physicists, assumed a definiteness and clearness in which it was previously lacking. The laws which govern stability hold good in regions of the greatest diversity ; they apply to the motion of planets round the sun, to the internal arrangement of those minute corpuscles of which each chemical atom is constructed, and to the forms of celestial bodies. In the present essay I shall attempt to consider the laws of stability as relating to the last case, and shall discuss the succession of shapes which may be assumed by celestial bodies in the course of their evolution. I believe further that homologous conceptions are applicable in the consideration of the transmutations of the various forms of animal and of vegetable life and in other regions of thought. Even if some of my readers should think that what I shall say on this head is fanciful, yet at least the exposition will serve to illustrate the meaning to be attached to the laws of stability in the physical universe.

The publication of The Origin of Species ushered in a new era in the study of Embryology. Whereas, before the year 1859 the facts of anatomy and development were loosely held together by the theory of types, which owed its origin to the great anatomists of the preceding generation, to Cuvier, L. Agassiz, J. Müller, and R. Owen, they were now combined together into one organic whole by the theory of descent and by the hypothesis of recapitulation which was deduced from that theory. The view that a knowledge of embryonic and larval histories would lay bare the secrets of race-history and enable the course of evolution to be traced, and so lead to the discovery of the natural system of classification, gave a powerful stimulus to morphological study in general and to embryological investigation in particular. In Darwin's words: “Embryology rises greatly in interest, when we look at the embryo as a picture, more or less obscured, of the progenitor, either in its adult or larval state, of all the members of the same great class.” In the period under consideration the output of embryological work has been enormous. No group of the animal kingdom has escaped exhaustive examination and no effort has been spared to obtain the embryos of isolated and out of the way forms, the development of which might have an important bearing upon questions of phylogeny and classification.

There are several points of view from which the subject of the present essay may be regarded. We may consider the fossil record of plants in its bearing: I. on the truth of the doctrine of Evolution; II. on Phylogeny, or the course of Evolution; III. on the theory of Natural Selection. The remarks which follow, illustrating certain aspects only of an extensive subject, may conveniently be grouped under these three headings.

The Truth of Evolution.

When The Origin of Species was written, it was necessary to show that the Geological Record was favourable to, or at least consistent with, the Theory of Descent. The point is argued, closely and fully, in Chapter x. “On the Imperfection of the Geological Record,” and Chapter xi. “On the Geological Succession of Organic Beings”; there is, however, little about plants in these chapters. At the present time the truth of Evolution is no longer seriously disputed, though there are writers, like Reinke, who insist, and rightly so, that the doctrine is still only a belief, rather than an established fact of science. Evidently, then, however little the Theory of Descent may be questioned in our own day, it is desirable to assure ourselves how the case stands, and in particular how far the evidence from fossil plants has grown stronger with time.