In Palæolithic times when each family was self-contained, before social laws and customs had bound such units together to form tribes and clans, the hunters, for such they all were, followed their quarry through the forests and along the valleys, keeping to no set trackways. They needed none, for trackways imply thoroughfares to and from fixed sites or habitations—and the Palaeolithic hunter was a nomad.
When the amalgamation of families took place in early Neolithic or Epipalaeolithic times, no doubt the marsh-dwellers had recognized paths across the swamps to the mainland, and those who formed the shell-mounds of Ertebolle may have used beaten paths along the shore from estuary to estuary.