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Early Miocene land mammals from eastern North America are exceedingly rare. Over the past several decades a small, but significant, vertebrate fauna has been recovered by paleontologists and citizen scientists from the Belgrade Formation at the Martin Marietta Belgrade Quarry in eastern North Carolina. This assemblage has 12 land mammal taxa, including beaver (Castoridae), stem lagomorph, carnivorans (Mustelidae, Ailuridae), horses (Equidae), rhinoceros (Rhinocerotidae), tapir (Tapiridae), peccary (Tayassuidae), anthracothere (Anthracotheriidae), entelodont (Entelodontidae), and protoceratid (Protoceratidae). Taken together, the biochronology of this Maysville Local Fauna indicates a late Arikareean (Ar3/Ar4) to early Hemingfordian (He1) North American Land Mammal Age (NALMA). This interval, which includes the Runningwater Chronofauna, documents numerous important Holarctic immigrants, including Amphictis, Craterogale, and cf. Menoceras found at this locality. Strontium isotope stratigraphy (SIS) of shark teeth collected in situ from the Belgrade Formation yield an age of 21.4 ± 0.13 Ma, which validates the age of interbedded land mammals within this unit. It also is consistent with the late Arikareean (Ar3/Ar4) biochronology and Aquitanian Neogene marine stage. New SIS analyses of oysters (Striostrea gigantissima) and clams (Chione) from this mine, previously assigned to late Oligocene or Late Miocene, are significantly older (28.0 ± 0.22 Ma and 27.6 ± 0.26 Ma, respectively) than the land mammals. Depending upon stratigraphic interpretations, these may confirm an older marine facies within the Belgrade Formation. This locality is important because of its marine and terrestrial tie-ins that facilitate intercalibration of both NALMAs and Cenozoic marine stages.
We present new unconstrained simulations and constrained experiments of a pair of pitching hydrofoils in a leader–follower in-line arrangement. Free-swimming simulations with matched pitching amplitudes show self-organisation into stable formations at a constant gap distance without any control. Over a wide range of phase synchronisation, amplitude and Lighthill number typical of biology, we discover that the stable gap distance scales with the actual wake wavelength of an isolated foil rather than the nominal wake wavelength. A scaling law for the actual wake wavelength is derived and shown to collapse data across a wide Reynolds number range of $200 \leqslant Re \leqslant 59\,000$. Additionally, vortex analysis uncovers that the leader’s wake wavelength-to-chord ratio, $\lambda /c$, is the key dimensionless variable to maximise the follower’s/collective efficiency. When $\lambda /c \approx 2$ it ensures that the follower’s leading edge suction force and the net force from a nearby vortex pair act in the direction with the foil’s motion thereby reducing the follower’s power. Moreover, in both simulations and experiments mismatched foil amplitudes are discovered to increase the efficiency of hydrofoil schools by 70 % while maintaining a stable formation without closed-loop control. This occurs by (i) increasing the stable gap distance between foils to push them into a high-efficiency zone and (ii) raising the level of efficiency in these zones. This study bridges the gap between constrained and unconstrained studies of in-line schooling by showing that constrained-foil measurements can map out the potential efficiency benefits of schooling. These findings can aid in the design of high-efficiency biorobot schools.
Over the interval 2008–2023 a large number of studies have been published testing various aspects of punctuated equilibria, including the prevalence of stasis, and also the extent to which most evolutionary change is concentrated at cladogenesis. In the vast majority of studies, punctuated equilibria continued to be strongly validated, as widespread evidence for stasis accumulated, with only some rare incidences of gradual change found. Support for the importance of cladogenetic change has increased, and new analytical approaches to study punctuated equilibria have been developed. Over this time period, there has also been an increase in the number of studies that have concentrated on extant taxa to test for punctuated equilibria, and these have also corroborated its widespread presence. In this respect, punctuated equilibria has served as an important bridge between neontological and paleontological approaches to evolutionary biology. From 2008 to 2023, there has also been some drift in how stasis is defined, such that, in certain studies, the definition diverged from the original 1972 definition in important respects. Notably, it is the few studies that have most changed the definition of what stasis constitutes that have most challenged the validity of punctuated equilibria, indicating it is morphing interpretations and definitions rather than the discovery of data compatible with phyletic gradualism that are most responsible for divergent results.
We describe the findings of cyclidans from the unpublished collection of the famous paleontologist B.I. Chernyshev (1888–1950) in the storage of the Academician F.N. Chernyshev Central Scientific Research Geological Survey Museum (CNIGR museum, St. Petersburg, Russia). These cyclidans were discovered by various researchers in the Carboniferous and Permian of the Urals. They are represented by the following taxa: Oonocarcinus uralicus new species, Uralocyclus feldmanni new species, Ambocyclus capidulum (Chernyshev, 1933), and Magnitocyclus (?) sp. indet. The discovery of the new species Oonocarcinus uralicus n. sp. greatly expands the geographic and stratigraphic interval of the genus Oonocarcinus Gemmellaro, 1890, previously known from the Middle Permian and Triassic. The discovery of Uralocyclus feldmanni n. sp. in the Mississippian deposits of the Chelyabinsk Oblast indicates a wide distribution of the genus Uralocyclus Mychko and Alekseev, 2018 in the Early Carboniferous, because Carboniferous representatives of this genus were previously known only from Ireland and England. The paper provides an up-to-date list of all known cyclidan occurrences and taxa in Russia.
The Paragaricocrinidae is an enigmatic late Paleozoic family of camerate crinoids that retained a robustly constructed calyx more typical of Devonian to Early Mississippian crinoids. The discovery of the oldest member of this family, Tuscumbiacrinus madisonensis n. gen. n. sp., initiated a phylogenetic investigation of the Paragaricocrinidae and consideration of its diversification and paleobiogeographic distribution. Phylogenetic analyses demonstrate the need to describe Tuscumbiacrinus n. gen and conduct revisions to preexisting taxa, resulting in the description of Palenciacrinus mudaensis n. gen. n. sp.; Pulcheracrinus n. gen.; Nipponicrinus hashimotoi n. gen. n. sp.; and Nipponicrinus akiyoshiensis n. gen. n. sp. Megaliocrinus exotericus Strimple is reassigned to Pulcheracrinus n. gen. In addition to having an anachronistic morphology, relatively few specimens are known through the ca. 76-million-year duration of this family. This pattern is unlikely to have resulted from low fossil sampling alone, and instead likely reflects low abundance and/or taxonomic richness of a long-lived waning clade. From its apparent origination in Laurussia during the Mississippian, the Paragaricocrinidae diversified into a cosmopolitan clade. Following a diversity drop during the Pennsylvanian, the Paragaricocrinidae persisted but exemplified characteristics of a dead clade walking until its eventual extinction during the middle Permian (Wordian).
Aerodynamic loads play a central role in many fluid dynamics applications, and we present a method for identifying the structures (or modes) in a flow that make dominant contributions to the time-varying aerodynamic loads in a flow. The method results from the combination of the force partitioning method (Menon & Mittal, 2021, J. Fluid Mech., vol. 907, A37) and modal decomposition techniques such as Reynolds decomposition, triple decomposition and proper orthogonal decomposition, and is applied here to three distinct flows – two-dimensional flows past a circular cylinder and an aerofoil, and the three-dimensional flow over a revolving rectangular wing. We show that the force partitioning method applied to modal decomposition of velocity fields results in complex, and difficult to interpret inter-modal interactions. We therefore propose and apply modal decomposition directly to the $Q$-field associated with these flows. The variable $Q$ is a nonlinear observable that is typically used to identify vortices in a flow, and we find that the direct decomposition of $Q$ leads to results that are more amenable to interpretation. We also demonstrate that this modal force partitioning can be extended to provide insights into the far-field aeroacoustic loading noise of these flows.
In this study, we explore the evolution of instabilities in magneto-quasi-geostrophic (MQG) modons on the $f$-plane using a magnetohydrodynamic rotating shallow water model. The numerical experiments have been conducted using a recently proposed second-order flux-globalisation-based path-conservative central-upwind scheme. Our focus is on the evolution and interactions of three key configurations: singular, regular and hollow MQG modons, which represent cases where the magnetic field is confined within the separatrix, evenly distributed inside and outside the separatrix and localised outside the separatrix, respectively. The singular MQG modon emerges as the most stable configuration, demonstrating the greatest resilience to destabilising forces. A notable observation is its transition from a quadrupolar to a tripolar magnetic field structure before reverting to a quadrupole adjusted magnetic modon, accompanied by a clockwise rotation of the system. In terms of stability, singular modons are the most stable ones, while hollow modons are the least stable. As instabilities develop, southward or northward displacements become significantly more pronounced than eastward or westward movements, primarily due to the Coriolis force. Among the configurations, the hollow (singular) modons experience the biggest (smallest) displacements. Additionally, we investigate modon collisions and highlight three scenarios: interactions between cyclonic and anticyclonic components that form a composite modon with meridional bifurcation; collisions of cyclonic vortices that produce a tripolar structure with counterclockwise rotation; and collisions between anticyclonic components that result in a stable, quasi-stationary tripolar configuration. The resulting magnetic poles exhibit a checkered pattern, with their amplitude decreasing with increasing distance from the central vortex.
A primary tenet of punctuated equilibria (PE) is that stasis, that is, little to no net morphological change, characterizes the histories of species. In the past ~50 years since PE was proposed, stasis has been recognized in the evolutionary histories of many species, but consensus has not been reached concerning its causes.
One unresolved issue is whether viable ecological mechanisms for stasis exist. We argue that a promising potential ecological explanation for stasis is coevolutionary alternation, which addresses how antagonists (e.g., predators or parasites and their groups of victims) coevolve over eco-evolutionary time across broad spatial scales. Coevolutionary alternation predicts different patterns of predator preferences and prey defenses within different populations and alternation of high and low levels of prey defenses as predator preferences evolve. The geographic structure of populations experiencing different environmental pressures and coevolutionary dynamics can yield stasis in such traits on the scale of entire species. We suggest that predator–prey coevolutionary alternation could be modeled using coupled stochastic differential equations (SDEs), which have been used to study correlative and causal connections among time series. SDEs can handle irregular sampling intervals, measurement uncertainty, and feedback loops between time series and can incorporate environmental proxies and time series from multiple geographic locations. We advocate developing this approach further to test the role of coevolutionary alternation in stasis and make recommendations for how SDEs might be used to model the coevolutionary feedback of predator(s) and multiple prey populations evolving in response to one another across space in a constantly changing environment.
Rare earth elements (REEs) preserved in speleothems have garnered increasing attention as ideal proxies for the paleoenvironmental reconstruction. However, due to their typically low contents in stalagmites, the availability of stalagmite-based REE records remains limited. Here we present high-resolution REEs alongside oxygen isotope (δ18O) records in stalagmite SX15a from Sanxing Cave, southwestern China (110.1–103.3 ka). This study demonstrates that REE records could provide useful information for the provenance and formation process of the stalagmite, due to consistent distribution pattern across different periods indicating stable provenance. More interestingly, the total REE (ΣREE) record could serve as an effective indicator to reflect local hydrological processes associated with monsoonal precipitation. During Marine Isotopic Stage (MIS) 5d, a relatively low ΣREE content is consistent with the positive SX15a δ18O and negative NGRIP δ18O, reflecting a dry-cold environment; while during MIS 5c, a generally high ΣREE content suggests a humid-warm circumstance. Furthermore, the ΣREE record captured four prominent sub-millennial fluctuations within the Greenland interstadial 24 event, implying a combined influence by the regional climate and local soil redox conditions. Our findings indicate that the stalagmite-based REE records would be a useful proxy for better understanding of past climate and environment changes.
Viscous flow through high-permeability channels occurs in many environmental and industrial applications, including carbon sequestration, groundwater flow and enhanced oil recovery. In this work, we study the displacement of a less-viscous fluid by a more-viscous fluid in a layered porous medium in a rectilinear configuration, where two low-permeability layers sandwich a higher-permeability layer. We derive a theoretical model that is validated using corroborative laboratory experiments, when the influence of the density difference is negligible. We find that the location of the propagating front increases with time according to a power-law form $x_f \propto t^{1/2}$, while the fluid–fluid interface exhibits a self-similar shape, when the motion of the displaced fluid is negligible in an unconfined porous medium. In the experimental set-up, distinct permeability layers were constructed using various sizes of spherical glass beads. The working fluids comprised fresh water as the less-viscous ambient fluid, and a glycerine–water mixture as the more-viscous injecting fluid. Our experimental measurement show a better match with the theory for the experiments performed at low Reynolds numbers and with permeable boundaries in the far field.
Antarctic glaciers have been considered classically uninhabited. However, they constitute an authentic biome and are populated by microorganisms that not only survive in them but also maintain an active metabolism. The South Shetland archipelago is a good study model to observe the diversity and evolution of the microbial populations that inhabit its glaciers. From a geological point of view, this archipelago is of considerable interest due to the intense and relatively recent volcanic eruptions on Deception Island. Additionally, it has been a place of transit for human and animal populations over time. All of these factors have influenced the composition and diversity of the microbial communities inhabiting the glacial ice. Among these microorganisms, a great diversity of bacteria, archaea, viruses and microeukaryotes such as algae and unicellular fungi have been identified thanks to high-throughput technologies. These cold-adapted microorganisms develop molecular mechanisms of adaptation to the extreme environment they inhabit and contribute to global energy cycles through the processing of organic and inorganic compounds. This review summarizes current knowledge on the biodiversity, ecology and molecular mechanisms of adaptation of cold-adapted microorganisms, and it details the specific characteristics of the microbial populations housed in the Antarctic glaciers in the South Shetland archipelago.
Balancing theoretical and practice-oriented elements, this book introduces researchers, teachers, and students in international sustainable development law to the IFIs' safeguard policies. It also scrutinizes the case law of independent accountability mechanisms that interpret those policies and afford recourse to individuals and communities adversely affected by development projects. The book's focus on the procedural and substantive features of IFIs' safeguard systems contributes to a more concrete understanding of these organizations' participation in the international lawmaking process on sustainable development. It puts IFIs in the spotlight and provides an international legal critique of their activities to match their notoriety in popular consciousness and to enhance their accountability to those they harm. By approaching international (economic) law and sustainable development through the lens of economic, environmental, and social issues arising in development projects primarily in the Global South, the book presents a needed counterbalance to existing literature on the topic.
We study the homogeneous isotropic turbulence of a shear-thinning fluid modelled by the Carreau model, and show how the variable viscosity affects the multiscale behaviour of the turbulent flow. We show that Kolmogorov theory can be extended to such non-Newtonian fluids, provided that the correct choice of average is taken when defining the mean Kolmogorov scale and dissipation rate, to properly capture the effect of the variable viscosity. Thus the classical phenomenology à la Kolmogorov can be observed in the inertial range of scale, with the energy spectra decaying as $k^{-5/3}$, with $k$ being the wavenumber, and the third-order structure function obeying the $4/5$ law. The changing viscosity instead strongly alters the small scale of turbulence, leading to an enhanced intermittent behaviour of the velocity field.
A nonlinear Schrödinger equation for pure capillary waves propagating at the free surface of a vertically sheared current has been used to study the stability and bifurcation of capillary Stokes waves on arbitrary depth. A linear stability analysis of weakly nonlinear capillary Stokes waves on arbitrary depth has shown that (i) the growth rate of modulational instability increases as the vorticity decreases whatever the dispersive parameter $kh$, where $k$ is the carrier wavenumber and $h$ the depth; (ii) the growth rate is significantly amplified for shallow water depths; and (iii) the instability bandwidth widens as the vorticity decreases. Particular attention has been paid to damping due to viscosity and forcing effects on modulational instability. In addition, a linear stability analysis to transverse perturbations in deep water has been carried out, demonstrating that the dominant modulational instability is two-dimensional whatever the vorticity. Near the minimum of linear phase velocity in deep water, we have shown that generalised capillary solitary waves bifurcate from linear capillary Stokes waves when the vorticity is positive. Moreover, we have shown that the envelope of pure capillary waves in deep water is unstable to transverse perturbations. Consequently, deep-water generalised capillary solitary waves are expected to be unstable to transverse perturbations.
Görtler vortices induced by concave curvature in supersonic turbulent flows are investigated using resolvent analysis and large-eddy simulations at Mach 2.95 and Reynolds number $ Re_{\delta }=63\,500$ based on the boundary-layer thickness $ \delta$. Resolvent analysis reveals that the most amplified coherent structures manifest as streamwise counter-rotating vortices with optimal spanwise wavelength $ 2.4\delta$ at cut-off frequency $f\delta /{u}_{\infty } =0.036$, where $ {u}_{\infty }$ is the freestream velocity. The leading spectral proper orthogonal decomposition modes with spanwise wavelength approximately $ 2\delta$ align well with the predicted coherent structures from resolvent analysis at $f\delta /{u}_{\infty } =0.036$. These predicted and extracted coherent structures are identified as Görtler vortices, driven by the Görtler instability. The preferential spanwise scale of the Görtler vortices is further examined under varying geometric and freestream parameters. The optimal spanwise wavelength is insensitive to the total turning angle beyond a critical value, but sensitive to the concave curvature $ K$ at the same turning angle. A limit spanwise wavelength $ 1.96\delta$, corresponding to an infinite concave curvature as $ K\rightarrow \infty$, is identified and validated. Increasing the freestream Mach number or decreasing the ratio of wall temperature to freestream temperature reduces the optimal wavelength normalised by $ \delta$, while variations in freestream Reynolds number have negligible impact. Additionally, a modified definition of the turbulent Görtler number $ G_{T}$ based on the peak eddy viscosity in boundary layers is proposed and employed to assess the occurrence of Görtler instability.
The supersonic wake of a circular cylinder in Mach 3 flow was studied through spectral proper orthogonal decomposition (SPOD) of high-speed focussing schlieren datasets. A wavenumber decomposition of the SPOD eigenvectors was found to be an effective tool for isolating imaging artefacts from the flow features, resulting in a clearer interpretation of the SPOD modes. The cylinder wake consists of both symmetric and antisymmetric instabilities, with the former being the dominant type. The free shear layers that form after the flow separates from the cylinder surface radiate strong Mach waves that interact with the recompression shocks to release significant disturbances into the wake. The wake shows a bimodal vortex shedding behaviour with a purely hydrodynamic instability mode around a Strouhal number of 0.2 and an aeroacoustic instability mode around Strouhal number of 0.42. The hydrodynamic mode, which is presumably the same as the incompressible case, is weaker and decays rapidly as the wake accelerates due to increasing compressibility. The aeroacoustic mode is the dominant shedding mode and persists farther into the wake because of an indirect energy input received through free-stream acoustic waves. A simple aeroacoustic feedback model based on an interaction between downstream propagating shear-layer instabilities and upstream propagating acoustic waves within the recirculation region is shown to accurately predict the shedding frequency. Based on this model, the vortex shedding in supersonic flows over a circular cylinder occurs at a universal Strouhal number (based on approach free-stream velocity and feedback path length) of approximately 0.3.
In this work we focus on expected flow in porous formations with highly conductive isolated fractures, which are of non-negligible length compared with the scales of interest. Accordingly, the definition of a representative elementary volume (REV) for flow and transport predictions may not be possible. Recently, a non-local kernel-based theory for flow in such formations has been proposed. There, fracture properties like their expected pressure are represented as field quantities. Unlike existing models, where fractures are assumed to be small compared with the scale of interest, a non-local kernel function is used to quantify the expected flow transfer between a point in the fracture domain and a potentially distant point in the matrix continuum. The transfer coefficient implied by the kernel is a function of the fracture characteristics that are in turn captured statistically. So far the model has successfully been applied for statistically homogeneous cases. In the present work we demonstrate the applicability for heterogeneous cases with spatially varying fracture statistics. Moreover, a scaling law is presented that relates the transfer coefficient to the fracture characteristics. Test cases involving discontinuously and continuously varying fracture statistics are presented, and the validity of the scaling law is demonstrated.
Fluids at supercritical pressures exhibit large variations in density near the pseudo-critical line, such that buoyancy plays a crucial role in their fluid dynamics. Here, we experimentally investigate heat transfer and turbulence in horizontal hydrodynamically developed channel flows of carbon dioxide at $88.5$ bar and $32.6\,^{\circ }\rm C$, heated at either the top or bottom surface to induce a strong vertical density gradient. In order to visualise the flow and evaluate its heat transfer, shadowgraphy is used concurrently with surface temperature measurements. With moderate heating, the flow is found to strongly stratify for both heating configurations, with bulk Richardson numbers $Ri$ reaching up to 100. When the carbon dioxide is heated from the bottom upwards, the resulting unstably stratified flow is found to be dominated by the increasingly prevalent secondary motion of thermal plumes, enhancing vertical mixing and progressively improving heat transfer compared with a neutrally buoyant setting. Conversely, stable stratification, induced by heating from the top, suppresses the vertical motion, leading to deteriorated heat transfer that becomes invariant to the Reynolds number. The optical results provide novel insights into the complex dynamics of the directionally dependent heat transfer in the near-pseudo-critical region. These insights contribute to the reliable design of heat exchangers with highly property-variant fluids, which are critical for the decarbonisation of power and industrial heat. However, the results also highlight the need for further progress in the development of experimental techniques to generate reliable reference data for a broader range of non-ideal supercritical conditions.
The energy-harvesting performance of two oscillating hydrofoil turbines in tandem configuration is experimentally studied at a $Re$ of $20\,000$ to determine the array’s optimal kinematics. By characterising interactions between the leading foil’s wake and the trailing foil, the kinematic configuration required to maximise array power extraction is identified. This is done by prescribing leading-foil kinematics that produce specific wake regimes, identified by the maximum effective angle of attack, $\alpha _{T/4}$, parameter. The kinematics of the trailing foil are varied significantly from those of the leading foil, with heave and pitch amplitudes of $0.6c\lt h_{0,{\textit{tr}}}\lt 1.8c$ and $65^{\circ} \lt \theta _{0,{\textit{tr}}}\lt 75^{\circ}$, and inter-foil phase of $-110^{\circ} \lt \psi _{1-2}\lt 180^{\circ}$. Configurations with reduced frequencies of $0.11$ and $0.12$, and foil separations of $4c$ and $6c$ are tested within each wake regime. The power extracted by each foil over an oscillation cycle is measured through force and torque measurements. Wake–foil interactions that improve trailing foil performance are analysed with time-resolved particle image velocimetry. Constructive and destructive wake–foil interactions are compared, showing that trailing-foil performance improves by either avoiding wake vortices or interacting directly with them. By interacting with the primary wake vortex, the latter configuration sees no power loss during the cycle. System power from the two foils is found to be maximised when the leading foil operates at an intermediate $\alpha _{T/4}$ range, and when the trailing foil avoids wake vortices. This optimal array configuration sees both foils operating with different kinematics compared with the optimal kinematics of a single oscillating foil.