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Tropical Andean glaciers provide an important flux of freshwater to communities living both in high-altitude Cordillera and population centres downstream in countries such as Peru and Bolivia. Glacier recession threatens the sustainability of these water resources, and accurate modelling of future glacier behaviour is required to manage water stress in the region. These models must capture all processes contributing significantly to overall glacier mass budgets. Here we examine supraglacial pond and ice cliff development on three clean-ice glaciers in the Cordillera Vilcanota, Peru and their overall contribution to glacier mass balance. Whilst such features are common and well-studied on debris-covered glaciers, their development on debris-free glaciers has not been examined in detail. We use high-resolution contemporary and historical satellite imagery and repeat drone surveys to examine surface structure and geometry change over three glaciers during 1977–2024. We show how cliff and pond formation is driven by aspect-dependent surface melt of crevasse walls. These features act as ice loss hotspots, which enhance glacier net mass loss by ∼10% despite accounting for <5% glacier surface area. Incorporation of such amplified ice loss processes should be a priority for glacier model advances to achieve more accurate projections of future tropical glacier recession.
Reproductive synchrony is common in populations that inhabit seasonal environments where reproductive timing is important to offspring survival. Weddell seals (Leptonychotes weddellii) reproduce in strongly seasonal Antarctic environments and are known to exhibit reproductive synchrony that varies by latitude, whereby more southerly populations give birth later. The Erebus Bay population of Weddell seals is the southernmost mammal population in the world, and birth-date synchrony has been demonstrated in this population. Various life history correlates including sex, maternal age and reproductive status have been identified as predictors of birth timing, but all prior work has been done on pups born to locally born mothers. Immigrant females originating from unknown sites in more northerly locations also produce pups in Erebus Bay and may have different birth timing due to the earlier average birth dates in their natal colonies. Using 22 years of capture-mark-recapture data for Weddell seals in Erebus Bay, we aimed to assess whether the timing of birth dates for pups born to immigrant mothers differs from that of pups born to locally born mothers. To our knowledge, this is the first time that such a comparison has been studied in a wild population. Birth dates were examined using Bayesian linear regression. We analysed birth dates from 7539 pups (4932 from locally born mothers, 2607 from immigrant mothers) born to 2210 mothers (1254 locally born, 956 immigrant) and found that there were no biologically impactful differences in the birth dates of pups born to locally born or immigrant mothers. Additionally, birth-date patterns for immigrant mothers were similar to those for locally born mothers with respect to various traits. Our results demonstrate that immigrant Weddell seal mothers can synchronize birth timing with locally born mothers. More research is required to understand the underlying mechanisms that allow for immigrant seals to achieve birth-date synchrony.
Magnets have been utilised widely for their ability to induce rapid contact – such as snapping between magnets and ferromagnetic materials. Yet, how such interactions proceed under immersion in a viscous fluid remains poorly understood. Here, we study this problem using the classical configuration of a smooth solid sphere approaching a plane in a quiescent fluid. Induced magnetic attraction, a spatially varying force analogous to short-range dispersion forces, offers a plausible route to overcome the constraint of a diverging hydrodynamic drag, which is well understood using the framework of classical lubrication theory. Instinctively, one might expect it to enable finite-time contact. However, our experiments reveal a counterintuitive result: while magnetic forces accelerate the sphere towards the surface, reducing the approach time by two orders of magnitude compared with gravity, they ultimately fail to effectuate contact in finite time, as induced magnetic interactions are unable to mitigate lubrication drag, which is singular at the thin gap limit, and transitions to an exponential descent characteristic of constant forcing. We support these findings with a simple theoretical model that accurately predicts the magnetic force law from purely kinematic observations. Finally, we outline the conditions under which spatially varying forces can enable true finite-time contact and discuss future experimental directions.
This work investigates the formation mechanism of the turbulent secondary vortex street (SVS) which appears in the far wake of bluff bodies, when the (primary) Kármán vortex street is absent. The turbulent wakes of four types of highly porous bluff bodies (plates/meshes) are characterised via time-resolved particle image velocimetry and large eddy simulations. The effect of ambient turbulence and initial conditions on SVS development is also examined, by installing a turbulence grid upstream of the bodies, and by varying the homogeneity of the bluff body porosity. Our results indicate that the SVS is a far-wake evolution of near-wake shear-layer vortices which, in the absence of the vortex shedding instability, continually grow and are finally arranged into alternating vortices. Free-stream turbulence and body inhomogeneity are found to significantly influence SVS development by amplifying mixing and attenuating the shear-layer instabilities of the near wake, which in turn lead to the formation of weaker and less coherent SVS structures further downstream.
We investigate the evolution of an external particle jet in a dense particle bed subjected to a radially divergent air-blast. Both random and single-mode perturbations are considered. By analysing the particle dynamics, we show that the Rayleigh–Taylor instability (RTI), the Richtmyer–Meshkov instability (RMI) and large particle inertia contribute to the formation of the external jet. The external particle jet exhibits a spike-like structure at its top and a bubble-like structure near its bottom. As the expanding particle bed lowers the internal gas pressure, particles near the bubble experience strong inward coupling forces and undergo RTI with variable acceleration. Meanwhile, particles in the spike experience weak gas–particle coupling and collision forces due to large particle inertia and low particle volume fraction, respectively. Consequently, the particles in the spike retain a nearly constant velocity, in contrast to the accelerating spikes observed in cylindrical RTI. To investigate the contributions of RMI to the particle jet growth, we track the trough-near particles in the single-mode perturbation case. It is revealed that the trough-near particles accelerate under the perturbation-induced pressure gradient, overtaking the crest-near particles and inducing phase inversion, thereby resulting in an increase in jet length. We establish a linear-growth model for the jet length increment, similar to the planar Richtmyer–Meshkov impulsive model. Combined with the jet-length-increment model, we propose an external-particle-jet-length model that is consistent with both numerical and experimental results for diverse initial gas pocket central pressures and particle bed thicknesses.
Asymptotic flow states with limiting drag modification are explored via direct numerical simulations in a moderate-curvature viscoelastic Taylor–Couette flow of the FENE-P fluid. We show that asymptotic drag modification (ADM) states are achieved at different solvent-to-total viscosity ratios ($\beta$) by gradually increasing the Weissenberg number from 10 to 150. As $\beta$ decreases from 0.99 to 0.90, for the first time, a continuous transition pathway is realised from the maximum drag reduction to the maximum drag enhancement, revealing a complete phase diagram of the ADM states. This transition originates from the competition between Reynolds stress reduction and polymer stress development, namely, a mechanistic change in angular momentum transport. Reduced $\beta$ has been found to effectively enhance elastic instability, suppressing large-scale Taylor vortices while promoting the formation of small-scale elastic Görtler vortices. The enhancement and in turn dominance of small-scale structures result in stronger incoherent transport, facilitating efficient mixing and substantial polymer stress development that ultimately drives the AMD state transition. Further analysis of the scale-decomposed transport equation of turbulent kinetic energy reveals an inverse energy cascade in the gap centre, which is attributed to the polymer-induced energy redistribution: polymers extract more energy from large scales than they can dissipate, with the excess energy redirected to smaller scales. However, the energy accumulating at smaller scales cannot be dissipated immediately and is consequently transferred back to larger scales via nonlinear interactions, thereby unravelling a novel polymer-mediated cycle for the reverse energy cascade. Overall, this study unravels the challenging puzzle of the existence of distinct dynamically connected ADM states and paves the way for coordinated experimental, simulation and theoretical studies of transition pathways to desired ADM states.
The guided-jet waves (GJWs) that may be trapped into a jet are investigated by simulating the propagation of the waves generated by an acoustic source on the axis of a jet at a Mach number of 0.95. The flow is modelled as a cylindrical shear layer to avoid reflections in the axial direction. For the source frequencies considered, GJWs belonging to the first two radial GJW axisymmetric modes are observed. They propagate in the upstream or downstream directions, and are entirely or partially contained in the flow, depending on the frequency. Their amplitudes are quantified. In the frequency–wavenumber space, they lie along the GJW dispersion curves predicted using linear-stability analysis. At specific spatial locations, they vary strongly and sharply with the frequency, exhibiting tonal-like peaks near the frequencies of the stationary points in the dispersion curves where the GJWs are standing waves with zero group velocity. Given the flow configuration, these properties can be attributed to propagation effects not requiring axial resonance between upstream- and downstream-travelling waves. Finally, it can be noted that, upstream of the source, outside the jet, the GJW amplitudes fluctuate in a reverse sawtooth manner with very intense peaks up to 30 dB higher than the levels obtained without flow at 10 jet radii from the source, similarly to the GJW footprints in the near-nozzle spectra of high-subsonic jets.
The growth of wall-mounted ice within channel flow which leads to a constriction is of significant practical relevance, especially in applications relating to aero-icing, large-scale pipe networks and mechanical systems. Whilst earlier works have treated ice constrictions as independent of the oncoming flow, few models explicitly account for the two-way coupling between the thermal and dynamical properties of the fluid and the evolving ice. To this end, the present work seeks to describe the interaction between high-Reynolds-number channel flow and constricting ice boundaries governed by Stefan conditions. Numerical simulations of the model indeed reveal that ice forming on the channel walls grows inwards towards the centreline and subsequently creates almost total constriction. In other parameter regimes, however, there is no ice formation. Using both a numerical and asymptotic approach, we identify regions of parameter space in which ice formation, and subsequently flow constriction, does or does not occur.
The clustering of inertial particles in turbulent flows is ubiquitous in many applications. This phenomenon is attributed to the influence of multiscale vortex structures in turbulent flows on particle motion. In this study, our primary goal is to further investigate the vortex effect on particle motion. We perform analytical and numerical simulations to examine the motion of particles in a counter-rotating vortex pair (CVP) with circulation ratio $\gamma \in (-1,0)$. The small, dilute, heavy inertial particles with a low particle Reynolds number are considered. In particular, the particle Stokes number and density factor satisfy $St\in (0,0.3)$ and $ R\in (0,1)$, respectively. We validate the existence of a particle-attracting ring within the CVP, which provides a simple mechanism for particle trapping. Meanwhile, there exists a critical Stokes number $St_{{cr}}$ limiting the occurrence of particle trapping. We provide a formula to predict the value of $St_{{cr}}$, which depends on both $\gamma$ and $R$. Only when $St\lt St_{{cr}}$ can the attracting ring trap the particle initially located within its basin of attraction and eventually lead to the formation of a particle clustering ring. Particles with a larger $R$ are more likely to be trapped in the CVP. While $St\gt St_{{cr}}$, the dynamics of the particles exhibits finite-time ‘leakage’. The attracting ring in the phase space coincides with the saddle point from which particles escape. Although all particles eventually escape, some may remain trapped in the vortex core region for a duration (represented by residence time). The distribution of residence time exhibits a localised exponential-like feature, indicating transient chaos.
Trophy hunting (TH) is a form of wildlife use in which individuals pay to hunt under regulated conditions and retain selected body parts as trophies. Tanzania permits TH in areas with different land-use designations, covering diverse habitats. However, human activities such as settlement, agriculture and livestock grazing can threaten habitats and wildlife populations, particularly as revenue from TH declines. Using satellite data from between 2013 and 2023, we quantified changes in functional area and land cover across 15 hunting blocks in northern Tanzania. Over this period, functional area in hunting blocks decreased by c. 22.4% in Game Controlled Areas (GCAs) and by c. 18.0% in Open Areas (OAs), attributed to human encroachment. Longido North GCA experienced the most substantial land-cover change, with a 76% increase in dense vegetation. An analysis comparing the 2013–2019 and 2019–2023 data revealed consistent shrinkages in grassland and mixed and dense vegetation within the Longido North GCA, Masai OA and Irkishbor OA hunting blocks. Although individual land-cover losses in the given periods remained below 50% (our threshold for major habitat loss), these cumulative changes reduce habitat suitability, posing risks to biodiversity and the sustainability of TH. This study reveals the extent of human pressures on hunting blocks and calls for integrated regional planning, active management and targeted conservation strategies.
Inspired by small intestine motility, we investigate the flow induced by a propagating pendular wave along the walls of a channel lined with rigid, villi-like microstructures. The villi undergo harmonic axial oscillations with a phase lag relative to their neighbours, generating travelling patterns of intervillous contraction. Using two-dimensional lattice Boltzmann simulations, we resolve the flow within the villi zone and the lumen, sampling small to moderate Womersley numbers. We uncover a mixing boundary layer (MBL) just above the villi, composed of semi-vortical structures that travel with the imposed wave. In the lumen, an axial steady flow emerges, surprisingly oriented opposite to the wave propagation direction, contrary to canonical peristaltic flows. We attribute this flow reversal to the non-reciprocal trajectories of fluid trapped between adjacent villi and derive a geometric scaling law that captures its magnitude in the Stokes regime. The MBL thickness is found to depend solely on the wave kinematics given by intervillous phase lag in the low-inertia limit. Above a critical threshold, oscillatory inertia induces dynamic confinement, limiting the radial extent of the MBL and leading to non-monotonic behaviour of the axial steady flux. We further develop an effective boundary condition at the villus tips, incorporating both steady and oscillatory components across relevant spatial scales. This framework enables coarse-grained simulations of intestinal flows without resolving individual villi. Our results shed light on the interplay among active microstructure, pendular wave and finite inertia in biological flows, and suggests new avenues for flow control in biomimetic and microfluidic systems.
Evidence for parasites in the fossil record is rare. As such, any examples present insight into parasitism in deep time. Trilobites have often been used for documenting parasites in the Paleozoic. Here we examine an Illaenus sp. pygidium from the Middle Ordovician of Estonia that displays thirteen small structures with domical to crater-like shapes. These morphologies are consistent with circular depressions on the pygidium inner surface. We propose that these structures formed while the trilobite was alive and record an infestation located within soft tissue. The trace maker seems to have influenced pygidial mineralization and caused a pathological reaction. The symbiont may have been capable of bioerosion, excavating these depressions by dissolving the trilobite’s mineral tissues; however, this scenario is less likely considering comparisons with syndromes and pathologies known in modern arthropods. The parasitic organism may have fed on the trilobite’s tissues or utilized nutrients within the trilobite’s body for growth. These observations are consistent with a parasitic organism.
The electrokinetic and unstable behaviour near strongly polarised surfaces cannot be well captured by the canonical asymptotic theory for induced-charge electro-osmosis, and the intrinsic mechanism remains unclear. Using direct numerical simulations and scaling analysis, this paper reveals that, near the strongly polarised surfaces, the strong electric double layer charging induces a strong local electric field, which drives the cations in the electrical double layer to extend to a finite region and form an extended space-charge (ESC) layer. The ESC triggers flow instability near strongly polarised surfaces, causing a transition of the velocity scaling exponent in the electric field dependence from a 2 to a 4/3 power law. The findings and mechanisms pave the way for designs of energy and biomedical systems.
A total of 12 ostracodes species belonging to 10 genera are reported for the first time from the lower Pliocene Takikawa Formation in Takikawa City, Hokkaido, northern Japan. The ostracode fauna of the formation includes both cryophilic and circumpolar species. Cryophilic species are Baffinicythere robusticostata Irizuki, 1996, Neomonoceratina tsurugasakensis (Tabuki, 1986), and Yezocythere hayashii Hanai and Ikeya, 1991, and circumpolar species are Palmenella limicola (Norman, 1867) and Semicytherura mainensis (Hazel and Valentine, 1969). The co-occurrence of cryophilic and circumpolar species suggests that the marine environment was strongly influenced by extremely cold water masses. The species diversity and equitability of the fossil ostracode assemblages indicate that the depositional topography was the inner area of a bay. One new genus and one new species, Woodeltia sorapuchiensis new genus new species, are described. This new genus occurred in a temperate environment in Japan, in the early Miocene. In the early Pliocene, Woodeltia adapted to cold environmental conditions in Hokkaido. We hypothesize that Woodeltia sorapuchiensis is an important species for Woodeltia, showing the group’s adaptation to cold environments and migration to the coast of North America.
Single-cell tornado-like vortices (TLVs) exhibit periodic wandering fluctuations around the time-averaged vortex core, a phenomenon known as vortex wandering, which constitutes the most prominent periodic behaviour in such flows. The coupling between vortex motion and wandering creates complex swirl dynamics, posing significant analytical challenges. However, the limited availability of experimental studies on vortex wandering decomposition hampers a deeper understanding of this phenomenon. To address this gap, a tornado simulator was designed to generate a controllable single-cell TLV, and high-frequency velocity data were obtained using particle image velocimetry. A sparsity-promoting dynamic mode decomposition (sp-DMD) method was developed to decouple coherent structures and analyse dynamic characteristics. Results show that as the swirl ratio increases, the vortex structure becomes more diffuse, with significant reductions in intensity. Vortex wandering is present across all swirl conditions, with its periodicity strongly modulated by the swirl ratio. Importantly, sp-DMD identified two primary modes, the time-averaged mode (first mode), representing the dominant rotational vortex motion, and the vortex-wandering-dominated modes (second and third conjugate modes), which correspond to persistent periodic velocity fluctuations and contribute the most significant pulsations. These modes exhibit a pair of oppositely rotating vortices symmetrically revolving around the central flow axis. Visualisations of the Q criterion reveal a symmetric dipole pattern. This suggests that rotational and shear effects are likely responsible for the periodic movement of the vortex core. Furthermore, as the swirl ratio increases, the energy of the vortex-wandering-dominated modes diminishes, and motion transitions from high-energy, organised dynamics to low-energy, disordered behaviour.
This paper presents an experimental and analytical investigation of the turbulent transport and flame geometric characteristics of free turbulent buoyant diffusion flames under different fuel mass fluxes and burner boundary conditions (i.e. with/without a flush floor). The stereo particle image velocimetry technique was utilised to measure the three-dimensional instantaneous velocity fields of the free methane buoyant flames with a burner diameter (d) of 0.30 m and dimensionless heat release rates ($\dot{Q}^{*}$) of 0.50–0.90. The results showed that, compared with the configuration without a floor, the time-averaged axial velocity fluctuations squared and the time-averaged radial velocity fluctuations squared decreased, and the peak values of the time-averaged radial velocity, the time-averaged radial velocity fluctuations squared and the time-averaged axial and radial fluctuation product shifted towards the burner centreline in the configuration with a flush floor. Based on the dimensional analysis and the gradient transport assumption, the mean turbulent viscosity within the mean flame height ($\nu _{t}^{=}$) was scaled. Compared with the configuration without a floor of under equal $\dot{Q}^{*}$, the turbulent viscosity decreased in the configuration with a flush floor, resulting in an increase in mean flame height and a reduction in mean flame width. Based on the concepts of turbulent mixing and equal axial convection and radial diffusion times, semi-physical models were derived for the mean flame height and the mean flame width, respectively. The two correlations agreed well with the experimental data of this work for the two burner configurations with and without a flush floor.